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<document id="87D2344A47660F1BB06120491EF71168" ID-CLB-Dataset="2946" ID-DOI="10.11646/zootaxa.5082.1.3" ID-GBIF-Dataset="c601c282-b593-47ff-bd7f-0b901cbf7b6c" ID-ISSN="1175-5326" ID-Zenodo-Dep="5783154" ID-ZooBank="06372651-6702-4741-AA55-2B302F34D21F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639564268802" checkinUser="plazi" docAuthor="Hutchinson, Paul M. &amp; Allsopp, Peter G." docDate="2021" docId="03EF120EFFBABB53098DBBB5FBE106FF" docLanguage="en" docName="zootaxa.5082.1.3.pdf" docOrigin="Zootaxa 5082 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudholophylla castaneipennis" docType="treatment" docVersion="8" lastPageNumber="34" masterDocId="FFD66A76FFBEBB56091ABB22FF980629" masterDocTitle="Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae): a sixth species of Pseudholophylla Blackburn, 1911, and notes on the other known species" masterLastPageNumber="40" masterPageNumber="30" pageNumber="34" updateTime="1699472537074" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C7C4A1226DB3F3EC31018BA14A89EF40">Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae): a sixth species of Pseudholophylla Blackburn, 1911, and notes on the other known species</mods:title>
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<mods:namePart id="F4E1295FDC78D05E07835B76BC35FFE8">Hutchinson, Paul M.</mods:namePart>
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<mods:namePart id="BF3457789687F01E4FB6F467E25E43C1">Allsopp, Peter G.</mods:namePart>
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<treatment id="03EF120EFFBABB53098DBBB5FBE106FF" ID-DOI="http://doi.org/10.5281/zenodo.5783812" ID-GBIF-Taxon="191647344" ID-Zenodo-Dep="5783812" LSID="urn:lsid:plazi:treatment:03EF120EFFBABB53098DBBB5FBE106FF" httpUri="http://treatment.plazi.org/id/03EF120EFFBABB53098DBBB5FBE106FF" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="34">
<subSubSection id="C35CF093FFBABB52098DBBB5FD700698" box="[151,744,151,178]" pageId="4" pageNumber="34" type="nomenclature">
<paragraph id="8BF9A318FFBABB52098DBBB5FD700698" blockId="4.[151,744,151,213]" box="[151,744,151,178]" pageId="4" pageNumber="34">
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<taxonomicName id="4C46D89BFFBABB52098DBBB5FD700698" authority="(Macleay, 1888)" baseAuthorityName="Macleay" baseAuthorityYear="1888" box="[151,744,151,178]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="B9327F0AFFBABB52098DBBB5FD700698" bold="true" box="[151,744,151,178]" pageId="4" pageNumber="34">
<emphasis id="B9327F0AFFBABB52098DBBB5FD840698" bold="true" box="[151,540,151,178]" italics="true" pageId="4" pageNumber="34">Pseudholophylla castaneipennis</emphasis>
(
<bibRefCitation id="EFD7DEE9FFBABB520B31BBB5FD780698" author="Macleay, W." box="[555,736,151,177]" pageId="4" pageNumber="34" pagination="897 - 924" refId="ref6071" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. Part II. The lamellicornes. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 897 - 924. https: // doi. org / 10.5962 / bhl. part. 29226" type="journal article" year="1888">Macleay, 1888</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFBABB52098DBB99FEB806FC" box="[151,288,187,213]" pageId="4" pageNumber="34" type="description">
<paragraph id="8BF9A318FFBABB52098DBB99FEB806FC" blockId="4.[151,744,151,213]" box="[151,288,187,213]" pageId="4" pageNumber="34">
(
<figureCitation id="137DBF9DFFBABB520985BB99FF7606FC" box="[159,238,187,213]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="4" pageNumber="34">Figs. 8</figureCitation>
,
<figureCitation id="137DBF9DFFBABB5209E1BB99FE8006FC" box="[251,280,187,213]" captionStart="FIGURE 17" captionStartId="8.[151,250,1498,1522]" captionTargetBox="[151,1436,345,1474]" captionTargetId="figure-84@8.[151,1436,345,1474]" captionTargetPageId="8" captionText="FIGURE 17. Known distributions of Pseudholophylla species across northern Australia. Upper map shows distribution relative to topography; lower map shows Köppen-Geiger classifications for comparison with climates. Pseudholophylla furfuracea (Burmeister, 1855) is recorded only from “Australia”." figureDoi="http://doi.org/10.5281/zenodo.5783166" httpUri="https://zenodo.org/record/5783166/files/figure.png" pageId="4" pageNumber="34">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFBABB52098DBA22FBAE071F" pageId="4" pageNumber="34" type="reference_group">
<paragraph id="8BF9A318FFBABB52098DBA22FDFC0731" blockId="4.[151,1078,256,310]" box="[151,612,256,280]" pageId="4" pageNumber="34">
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<treatmentCitation id="0AE78509FFBABB52098DBA22FDF80731" author="Macleay, W." box="[151,608,256,280]" page="919" pageId="4" pageNumber="34" year="1888">
<taxonomicName id="4C46D89BFFBABB52098DBA22FDF80731" authority="Macleay, 1888: 919" authorityName="Macleay" authorityPageNumber="919" authorityYear="1888" box="[151,608,256,280]" class="Insecta" family="Melolonthidae" genus="Rhopaea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="B9327F0AFFBABB52098DBA22FE17073E" box="[151,399,256,280]" italics="true" pageId="4" pageNumber="34">Rhopaea castaneipennis</emphasis>
<bibRefCitation id="EFD7DEE9FFBABB52088EBA22FDF80731" author="Macleay, W." box="[404,608,256,280]" pageId="4" pageNumber="34" pagination="897 - 924" refId="ref6071" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. Part II. The lamellicornes. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 897 - 924. https: // doi. org / 10.5962 / bhl. part. 29226" type="journal article" year="1888">Macleay, 1888: 919</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8BF9A318FFBABB52098DBA3CFBAE071F" blockId="4.[151,1078,256,310]" box="[151,1078,286,310]" pageId="4" pageNumber="34">
<treatmentCitationGroup id="AB568436FFBABB52098DBA3CFBAE071F" box="[151,1078,286,310]" pageId="4" pageNumber="34">
<treatmentCitation id="0AE78509FFBABB52098DBA3CFD0E071F" author="Britton, E. B." box="[151,662,286,310]" page="34" pageId="4" pageNumber="34" year="1978">
<taxonomicName id="4C46D89BFFBABB52098DBA3CFD0E071F" authority=": Britton 1978: 34" authorityName="Britton" authorityPageNumber="34" authorityYear="1978" box="[151,662,286,310]" class="Insecta" family="Melolonthidae" genus="Pseudholophylla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="34" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="B9327F0AFFBABB52098DBA3CFE47071F" box="[151,479,286,310]" italics="true" pageId="4" pageNumber="34">Pseudholophylla castaneipennis</emphasis>
:
<bibRefCitation id="EFD7DEE9FFBABB5208F6BA3CFD0E071F" author="Britton, E. B." box="[492,662,286,310]" pageId="4" pageNumber="34" pagination="1 - 150" refId="ref5766" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton 1978: 34</bibRefCitation>
</taxonomicName>
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, Figs. 97
<geoCoordinate id="EE72C5DFFFBABB520A1ABA3CFCC5071F" box="[768,861,286,310]" degrees="99, 274" direction="east" orientation="longitude" pageId="4" pageNumber="34" precision="55" value="99.274">99, 274E</geoCoordinate>
;
<treatmentCitation id="0AE78509FFBABB520A7CBA3CFBAA071F" ID-DOI="http://doi.org/10.5281/zenodo.4340912" ID-Zenodo-Dep="4340912" author="Allsopp, P. G." box="[870,1074,286,310]" httpUri="http://treatment.plazi.org/id/C264702F452335072BE79A08FBDCF9EA" page="476" pageId="4" pageNumber="34" year="2020">
<bibRefCitation id="EFD7DEE9FFBABB520A7CBA3CFBAA071F" author="Allsopp, P. G." box="[870,1074,286,310]" pageId="4" pageNumber="34" pagination="451 - 486" refId="ref5561" refString="Allsopp, P. G. (2020 b) Clarification of the status of the types of Australian Melolonthini (Coleoptera: Scarabaeidae: Melolonthinae) described before 1950. Zootaxa, 4885 (4), 451 - 486. https: // doi. org / 10.11646 / zootaxa. 4885.4.1" type="journal article" year="2020">Allsopp 2020b: 476</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFBABB52098DBA47FB5B07EE" pageId="4" pageNumber="34" type="materials_examined">
<paragraph id="8BF9A318FFBABB52098DBA47FE48078A" blockId="4.[151,1437,356,671]" pageId="4" pageNumber="34">
<emphasis id="B9327F0AFFBABB52098DBA47FEBB0757" bold="true" box="[151,291,356,383]" pageId="4" pageNumber="34">Type series.</emphasis>
<materialsCitation id="3B2EA945FFBABB520830BA47FE48078A" ID-GBIF-Occurrence="3418631315" collectionCode="ANIC" collectorName="Range N. W. Aust." location="WESTERN AUSTRALIA" pageId="4" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="lectotype">
<typeStatus id="54FD1DBAFFBABB520830BA47FE020756" box="[298,410,357,383]" pageId="4" pageNumber="34" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D406891FFBABB5208BBBA47FE200756" box="[417,440,357,383]" pageId="4" pageNumber="34" type="male"></specimenCount>
, designated by
<bibRefCitation id="EFD7DEE9FFBABB520B7CBA46FC930756" author="Britton, E. B." box="[614,779,356,383]" pageId="4" pageNumber="34" pagination="1 - 150" refId="ref5766" refString="Britton, E. B. (1978) A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae). Vol. 2. Tribe Melolonthini. Australian Journal of Zoology, Supplementary Series, 60, 1 - 150. https: // doi. org / 10.1071 / AJZS 060" type="journal article" year="1978">Britton (1978)</bibRefCitation>
.
<collectingRegion id="49826DFAFFBABB520A0FBA47FBD90756" box="[789,1089,357,383]" country="Australia" name="Western Australia" pageId="4" pageNumber="34">WESTERN AUSTRALIA</collectingRegion>
. Barrior [Barrier]
<collectorName id="26B3C6CEFFBABB520C17BA47FF4F078A" pageId="4" pageNumber="34">Range N. W. Aust.</collectorName>
; in
<collectionCode id="ED573BDDFFBABB5209E7BAABFED9078A" box="[253,321,393,419]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="4" pageNumber="34">ANIC</collectionCode>
[examined].
</materialsCitation>
</paragraph>
<paragraph id="8BF9A318FFBABB5209DDBA8EFB5B07EE" blockId="4.[151,1437,356,671]" box="[199,1219,428,455]" pageId="4" pageNumber="34">
<materialsCitation id="3B2EA945FFBABB5209DDBA8EFB2707EE" ID-GBIF-Occurrence="3418631313" box="[199,1215,428,455]" collectionCode="ANIC" collectorName="N. W. Austra" location="WESTERN AUSTRALIA" pageId="4" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia" typeStatus="paralectotype">
<typeStatus id="54FD1DBAFFBABB5209DDBA8EFEC707EF" box="[199,351,428,454]" pageId="4" pageNumber="34" type="paralectotype">Paralectotype</typeStatus>
<specimenCount id="9D406891FFBABB52087DBA8FFEE707EE" box="[359,383,429,455]" pageId="4" pageNumber="34" type="male"></specimenCount>
.
<collectingRegion id="49826DFAFFBABB520893BA8FFD2D07EE" box="[393,693,429,455]" country="Australia" name="Western Australia" pageId="4" pageNumber="34">WESTERN AUSTRALIA</collectingRegion>
.
<collectorName id="26B3C6CEFFBABB520BA5BA8FFCD207EE" box="[703,842,429,455]" pageId="4" pageNumber="34">N.W. Austra</collectorName>
[North-west Australia]; in
<collectionCode id="ED573BDDFFBABB520D6DBA8FFB2707EE" box="[1143,1215,429,455]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="4" pageNumber="34">ANIC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFBABB5209DDBAF2FC6404B7" pageId="4" pageNumber="34" type="diagnosis">
<paragraph id="8BF9A318FFBABB5209DDBAF2FC6404B7" blockId="4.[151,1437,356,671]" pageId="4" pageNumber="34">
<emphasis id="B9327F0AFFBABB5209DDBAF2FE3707C2" bold="true" box="[199,431,464,491]" pageId="4" pageNumber="34">Diagnosis of males.</emphasis>
Body approximately
<quantity id="4CBE0EFDFFBABB520BBDBAF2FD6007C2" box="[679,760,464,491]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="34" unit="mm" value="16.0">16 mm</quantity>
long. Colour reddish brown to dark brown. Lamella of antennomere 4 approximately one-third length of lamella of antennomere 5. Maxillary palp with apical palpomere slender, with an elongate, flat, dull area on upper surface. Pronotum with greatest width 1.6x mid length; anterior margin defined by groove towards anterior angles, without defined posterior margin; pygidium with evenly distributed sparse setae. Parameres (
<figureCitation id="137DBF9DFFBABB5208FCB942FDB00452" box="[486,552,608,635]" captionStart="FIGURES 813" captionStartId="4.[151,264,1785,1809]" captionTargetBox="[168,1426,702,1754]" captionTargetId="figure-170@4.[161,1426,694,1761]" captionTargetPageId="4" captionText="FIGURES 813. Pseudholophylla aedeagi. Each set with lateral view left, dorsal view top right, ventral view bottom right. 8, Pseudholophylla castaneipennis (Macleay, 1888); 9, P. cavifrons (Lea, 1919); 10, P. furfuracea (Burmeister, 1855); 11, P. lepidoptera (Blackburn, 1912); 12, P. soror Britton, 1978; 13, P. hurai Hutchinson &amp; Allsopp, new species, holotype male. Figs. 812 reproduced from Britton (1978) (CSIRO, used with permission)." figureDoi="http://doi.org/10.5281/zenodo.5783160" httpUri="https://zenodo.org/record/5783160/files/figure.png" pageId="4" pageNumber="34">Fig. 8</figureCitation>
) without a transverse apical process, dorsal and ventral margins evenly curved and tapering to rounded apex in lateral view, apex folded back in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="C35CF093FFBABB5309DDBC89FBE106FF" lastPageId="5" lastPageNumber="35" pageId="4" pageNumber="34" type="distribution">
<paragraph id="8BF9A318FFBABB5309DDBC89FBE106FF" blockId="4.[151,1437,1963,2026]" lastBlockId="5.[151,1437,150,214]" lastPageId="5" lastPageNumber="35" pageId="4" pageNumber="34">
<emphasis id="B9327F0AFFBABB5209DDBC89FEC101EC" bold="true" box="[199,345,1963,1989]" pageId="4" pageNumber="34">Distribution</emphasis>
(
<figureCitation id="137DBF9DFFBABB520872BC89FE2001EC" box="[360,440,1963,1989]" captionStart="FIGURE 17" captionStartId="8.[151,250,1498,1522]" captionTargetBox="[151,1436,345,1474]" captionTargetId="figure-84@8.[151,1436,345,1474]" captionTargetPageId="8" captionText="FIGURE 17. Known distributions of Pseudholophylla species across northern Australia. Upper map shows distribution relative to topography; lower map shows Köppen-Geiger classifications for comparison with climates. Pseudholophylla furfuracea (Burmeister, 1855) is recorded only from “Australia”." figureDoi="http://doi.org/10.5281/zenodo.5783166" httpUri="https://zenodo.org/record/5783166/files/figure.png" pageId="4" pageNumber="34">Fig. 17</figureCitation>
). The precise type locality was not given by
<bibRefCitation id="EFD7DEE9FFBABB520ABCBC89FBCF01EC" author="Macleay, W." box="[934,1111,1963,1989]" pageId="4" pageNumber="34" pagination="897 - 924" refId="ref6071" refString="Macleay, W. (1888) The insects of King's Sound and its vicinity. Part II. The lamellicornes. Proceedings of the Linnean Society of New South Wales, Series 2, 3, 897 - 924. https: // doi. org / 10.5962 / bhl. part. 29226" type="journal article" year="1888">Macleay (1888)</bibRefCitation>
but the label on the
<typeStatus id="54FD1DBAFFBABB520C2CBC89FA0501EC" box="[1334,1437,1963,1989]" pageId="4" pageNumber="34" type="lectotype">lectotype</typeStatus>
suggests that it was Barrier Range, now Napier Range. This was where the collector, Walter Froggatt, stayed near Windjana Gorge [
<geoCoordinate id="EE72C5DFFFBBBB530847BBB5FE230698" box="[349,443,151,177]" degrees="17.43" direction="south" orientation="latitude" pageId="5" pageNumber="35" precision="555" value="-17.43">17.43°S</geoCoordinate>
,
<geoCoordinate id="EE72C5DFFFBBBB5308DDBBB4FDAB0698" box="[455,563,150,177]" degrees="124.98" direction="east" orientation="longitude" pageId="5" pageNumber="35" precision="555" value="124.98">124.98°E</geoCoordinate>
] during JulyAugust and December 1887 and early 1888 (
<bibRefCitation id="EFD7DEE9FFBBBB530DDBBBB5FAFD0698" author="Froggatt, W. W." box="[1217,1381,151,177]" pageId="5" pageNumber="35" pagination="69 - 82" refId="ref5856" refString="Froggatt, W. W. (1934) A naturalist in Kimberley in 1887. The Australian Naturalist, 9, 69 - 82." type="journal article" year="1934">Froggatt 1934</bibRefCitation>
). We have used that locality in
<figureCitation id="137DBF9DFFBBBB5308ACBB99FD9106FC" box="[438,521,187,213]" captionStart="FIGURE 17" captionStartId="8.[151,250,1498,1522]" captionTargetBox="[151,1436,345,1474]" captionTargetId="figure-84@8.[151,1436,345,1474]" captionTargetPageId="8" captionText="FIGURE 17. Known distributions of Pseudholophylla species across northern Australia. Upper map shows distribution relative to topography; lower map shows Köppen-Geiger classifications for comparison with climates. Pseudholophylla furfuracea (Burmeister, 1855) is recorded only from “Australia”." figureDoi="http://doi.org/10.5281/zenodo.5783166" httpUri="https://zenodo.org/record/5783166/files/figure.png" pageId="5" pageNumber="35">Fig. 17</figureCitation>
; it has a Köppen-Geiger climate of Bsh (hot semi-arid).
</paragraph>
</subSubSection>
</treatment>
</document>