384 lines
54 KiB
XML
384 lines
54 KiB
XML
<document id="B289E846DA8995AD9A26CC11C54EC7D7" ID-CLB-Dataset="31143" ID-DOI="10.11646/zootaxa.4365.2.6" ID-GBIF-Dataset="15192bcb-c91d-4dc7-acd4-fd7caf151edc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1117560" ID-ZooBank="C785F83F-55D8-41E4-87EE-F4410B092E2A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513583977253" checkinUser="plazi" docAuthor="Mirza, Zeeshan A." docDate="2017" docId="03DD87EEFFFDF440FF053044FF58FD0F" docLanguage="en" docName="zootaxa.4365.2.6.pdf" docOrigin="Zootaxa 4365 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liocheles schalleri Mirza 2017, sp. nov." docType="treatment" docUuid="DB3072F7-8070-497F-A04E-76A1DBE4EAF8" docUuidSource="ZooBank" docVersion="8" lastPageNumber="226" masterDocId="FFE4FF96FFFEF449FF923415FFA9FFFF" masterDocTitle="Description of a new species of Liocheles Sundevall, 1833 (Hormuridae) from India" masterLastPageNumber="230" masterPageNumber="217" pageNumber="220" updateTime="1698510175314" updateUser="ExternalLinkService">
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<mods:mods id="49F7AB6E17588F673A7E2BC0B2C57D30" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="811192FFE6AAA6A86DC0A73AB16EAD13">
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<mods:title id="DDC927C3EE321B41EF77206B60B47B75">Description of a new species of Liocheles Sundevall, 1833 (Hormuridae) from India</mods:title>
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</mods:titleInfo>
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<mods:name id="C282D20C29BD240ECEBB2F995CA9D846" type="personal">
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<mods:role id="2D1CAECD99C297A9FCD8FD283268728D">
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<mods:roleTerm id="1156C7B240B6D844FF3C4C3AA56A7D36">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D03A9F8741E6D7E476FA1BAD4A42FBEE">Mirza, Zeeshan A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="197B0EA0D1C9BE666F67B142EC3B06A0">text</mods:typeOfResource>
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<mods:titleInfo id="DD69A34C78D0AF5FC212A08B0CDA96EF">
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<mods:title id="E21D1A0D83C65DF57001DE7CB977F7D6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9DA030C0DD43BAFCA72C5FF50624713F">
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<mods:date id="DA8BAEB61C6631C957BBB1FCB90527C3">2017</mods:date>
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<mods:detail id="E7A061C97ED1360960DE1651ADD0E3A1" type="pubDate">
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<mods:number id="49479D62BBC493E24D555BCC1DDDAF8C">2017-12-18</mods:number>
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</mods:detail>
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<mods:detail id="915717840579C6730857FFF20D008BA4" type="volume">
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<mods:number id="D73DE3C3F806DC613DB835C013FC457C">4365</mods:number>
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</mods:detail>
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<mods:detail id="A5FFA654224AF669E7512BE842A70E22" type="issue">
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<mods:number id="B8B47B812A83382900EB33123D24B79B">2</mods:number>
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</mods:detail>
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<mods:extent id="896F7A3D4DAA17857B4A32D408C19B4D" unit="page">
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<mods:start id="89B32AA71DB553704E0A3269365A0B3A">217</mods:start>
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<mods:end id="D6E7C3A1F03B7EF6F7719869395DD80A">230</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="606B84EF6449F9C7D06B2B09C3BEE651">journal article</mods:classification>
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<mods:identifier id="C0460D5FC34763B05A47C891F0840584" type="CLB-Dataset">31143</mods:identifier>
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<mods:identifier id="7A841EFBC63914AF12F8462ADFB93B92" type="DOI">10.11646/zootaxa.4365.2.6</mods:identifier>
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<mods:identifier id="F2BEDD5875D0E7A06BEEE12B27E8800B" type="GBIF-Dataset">15192bcb-c91d-4dc7-acd4-fd7caf151edc</mods:identifier>
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<mods:identifier id="2A6D34647100B1AC36BE356F2A76724D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="45F4C05BE32514E63A39B97FE971EE32" type="Zenodo-Dep">1117560</mods:identifier>
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<mods:identifier id="E99341BCD7DBF8E0A9AE578176BA8312" type="ZooBank">C785F83F-55D8-41E4-87EE-F4410B092E2A</mods:identifier>
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</mods:mods>
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<treatment id="03DD87EEFFFDF440FF053044FF58FD0F" ID-DOI="http://doi.org/10.5281/zenodo.6004341" ID-GBIF-Taxon="139334154" ID-Zenodo-Dep="6004341" LSID="urn:lsid:zoobank.org:act:DB3072F7-8070-497F-A04E-76A1DBE4EAF8" httpUri="http://treatment.plazi.org/id/03DD87EEFFFDF440FF053044FF58FD0F" lastPageId="9" lastPageNumber="226" pageId="3" pageNumber="220">
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<subSubSection id="C36E6573FFFDF44AFF053044FEDBFB72" pageId="3" pageNumber="220" type="nomenclature">
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<paragraph id="8BCB36F8FFFDF44AFF053044FE77FB93" blockId="3.[151,478,1105,1165]" box="[151,478,1105,1132]" pageId="3" pageNumber="220">
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<heading id="D0838194FFFDF44AFF053044FE77FB93" bold="true" box="[151,478,1105,1132]" fontSize="11" level="1" pageId="3" pageNumber="220" reason="1">
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<emphasis id="B900EAEAFFFDF44AFF053044FE77FB93" bold="true" box="[151,478,1105,1132]" pageId="3" pageNumber="220">
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<taxonomicName id="4C744D7BFFFDF44AFF053044FED3FB94" ID-CoL="3V5CP" authority="Mirza, 2017" authorityName="Mirza" authorityYear="2017" box="[151,378,1105,1131]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="schalleri" status="sp. nov.">
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<emphasis id="B900EAEAFFFDF44AFF053044FED3FB94" bold="true" box="[151,378,1105,1131]" italics="true" pageId="3" pageNumber="220">Liocheles schalleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2335791FFFDF44AFE133047FE77FB93" box="[385,478,1106,1132]" pageId="3" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCB36F8FFFDF44AFF053061FEDBFB72" blockId="3.[151,478,1105,1165]" box="[151,370,1140,1165]" pageId="3" pageNumber="220">
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(
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<figureCitation id="134F2A7DFFFDF44AFF0D3061FEA2FB72" box="[159,267,1140,1165]" captionStart-0="FIGURE 4" captionStart-1="FIGURE5" captionStart-2="FIGURE6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStartId-0="4.[151,250,654,676]" captionStartId-1="4.[151,250,1982,2004]" captionStartId-2="5.[151,250,1959,1981]" captionStartId-3="6.[151,250,1843,1865]" captionStartId-4="7.[151,250,1547,1569]" captionTargetBox-0="[163,1419,185,642]" captionTargetBox-1="[181,1403,743,1967]" captionTargetBox-2="[151,1436,793,1903]" captionTargetBox-3="[181,1405,185,1832]" captionTargetBox-4="[412,1175,597,1528]" captionTargetId-0="figure@4.[151,1436,173,652]" captionTargetId-1="figure@4.[167,1431,722,1988]" captionTargetId-2="figure@5.[151,1436,793,1903]" captionTargetId-3="figure@6.[181,1405,185,1832]" captionTargetId-4="figure@7.[402,1185,589,1537]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 4. Liocheles schalleri sp. nov. showing coloration in life (A) holotype male NCBS AG817, (B) paratype female NCBS AG818.Photo by Rajesh Sanap." captionText-1="FIGURE5. Liochelesschallerisp. nov. (A) male dorsal NCBS AG817, (B)male ventral NCBS AG817, (C)femaledorsal,(D) female ventral." captionText-2="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." captionText-3="FIGURE 7. Liocheles schalleri sp. nov. holotype male pedipalp chela showing tricobothrial pattern (A) dorsal, (B) external, (C) ventral, (D) internal." captionText-4="FIGURE 8. Liocheles schalleri sp. nov. holotype male NCBS AG817 hemispermataphore, (A) dorsal view, (B) close-up of capsule." httpUri-0="https://zenodo.org/record/1117566/files/figure.png" httpUri-1="https://zenodo.org/record/1117568/files/figure.png" httpUri-2="https://zenodo.org/record/1117570/files/figure.png" httpUri-3="https://zenodo.org/record/1117572/files/figure.png" httpUri-4="https://zenodo.org/record/1117574/files/figure.png" pageId="3" pageNumber="220">Figs. 4–8</figureCitation>
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,
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<tableCitation id="C6F60343FFFDF44AFE843061FECEFB72" box="[278,359,1140,1165]" captionStart="TABLE 1" captionStartId="9.[151,239,797,820]" captionText="TABLE 1. Morphometry of type specimens of Liocheles schalleri sp. nov." pageId="3" pageNumber="220">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E6573FFFDF44AFF0530A9FD92FA25" pageId="3" pageNumber="220" type="materials_examined">
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<paragraph id="8BCB36F8FFFDF44AFF0530A9FADFFB03" blockId="3.[151,1437,1212,2038]" pageId="3" pageNumber="220">
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<emphasis id="B900EAEAFFFDF44AFF0530A9FEA4FB2A" bold="true" box="[151,269,1212,1237]" pageId="3" pageNumber="220">
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<typeStatus id="54CF885AFFFDF44AFF0530A9FEA1FB2A" box="[151,264,1212,1237]" pageId="3" pageNumber="220" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectionCode id="ED65AE3DFFFDF44AFE8E30A8FECCFB2A" box="[284,357,1213,1237]" country="USA" httpUri="http://grbio.org/cool/x0m9-a3bs" name="Yale University" pageId="3" pageNumber="220">NCBS</collectionCode>
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AG817, male from Trishna WLS, South
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<collectingRegion id="49B0F81AFFFDF44AFCE230A9FC6AFB2A" box="[880,963,1212,1237]" country="India" name="Tripura" pageId="3" pageNumber="220">Tripura</collectingRegion>
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district,
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<collectingRegion id="49B0F81AFFFDF44AFBA530A9FB24FB2A" box="[1079,1165,1212,1237]" country="India" name="Tripura" pageId="3" pageNumber="220">Tripura</collectingRegion>
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,
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<collectingCountry id="F3637668FFFDF44AFB3230A9FB75FB2A" box="[1184,1244,1212,1237]" name="India" pageId="3" pageNumber="220">India</collectingCountry>
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,
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<geoCoordinate id="EE40503FFFFDF44AFB6230A9FA31FB2B" box="[1264,1432,1212,1237]" direction="north" orientation="latitude" pageId="3" pageNumber="220" precision="1" value="23.281126">23.281126° N</geoCoordinate>
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,
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<geoCoordinate id="EE40503FFFFDF44AFF0530F1FE98FB04" box="[151,305,1252,1276]" direction="east" orientation="longitude" pageId="3" pageNumber="220" precision="1" value="91.401">91.401004° E</geoCoordinate>
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(WGS84), elevation
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||
<quantity id="4C8C9B1DFFFDF44AFDB330F1FDFFFB03" box="[545,598,1252,1276]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="220" unit="m" value="30.0">30m</quantity>
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, collected on
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<date id="FFCA1038FFFDF44AFD6230F1FC04FB03" box="[752,941,1248,1276]" pageId="3" pageNumber="220" value="2014-10-02">2nd October 2014</date>
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by Rajesh Sanap and Zeeshan A. Mirza.
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||
</paragraph>
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||
<paragraph id="8BCB36F8FFFDF44AFF553112FCFBFA94" blockId="3.[151,1437,1212,2038]" pageId="3" pageNumber="220">
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<emphasis id="B900EAEAFFFDF44AFF553112FEE1FADF" bold="true" box="[199,328,1287,1312]" pageId="3" pageNumber="220">Paratypes:</emphasis>
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2 females NCBS AG818–AG819, 2 males NCBS AG820,
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<materialsCitation id="3B1C3CA5FFFDF44AFC4E311DFA2AFADF" ID-GBIF-Occurrence="1795578655" box="[988,1411,1287,1312]" collectionCode="NCBS" pageId="3" pageNumber="220" specimenCode="AG809" specimenCount="1">
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||
<collectionCode id="ED65AE3DFFFDF44AFC4E311DFB8CFADF" box="[988,1061,1288,1312]" country="USA" httpUri="http://grbio.org/cool/x0m9-a3bs" name="Yale University" pageId="3" pageNumber="220">NCBS</collectionCode>
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||
<specimenCode id="DBD29E83FFFDF44AFBB9311DFB29FADF" box="[1067,1152,1288,1312]" collectionCode="NCBS" country="USA" httpUri="http://grbio.org/cool/x0m9-a3bs" name="Yale University" pageId="3" pageNumber="220">AG809</specimenCode>
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, same data as holotype
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</materialsCitation>
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||
;
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<materialsCitation id="3B1C3CA5FFFDF44AFA1C311DFCFBFA94" ID-GBIF-Occurrence="1795578717" collectingDate="2014-10-09" collectionCode="NCBS" county="Sal" elevation="56" latitude="23.44166" location="Udaipur" longLatPrecision="1" longitude="91.48956" municipality="Garjee forest" pageId="3" pageNumber="220" specimenCode="AG829" specimenCount="1" specimenCount-female="1" stateProvince="Tripura">
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||
<specimenCount id="9D72FD71FFFDF44AFA1C311DFF4AFABB" pageId="3" pageNumber="220" type="female">1 female</specimenCount>
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||
<collectionCode id="ED65AE3DFFFDF44AFF7B3139FE9BFABB" box="[233,306,1324,1348]" country="USA" httpUri="http://grbio.org/cool/x0m9-a3bs" name="Yale University" pageId="3" pageNumber="220">NCBS</collectionCode>
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||
<specimenCode id="DBD29E83FFFDF44AFEAB3139FE22FABB" box="[313,395,1324,1348]" collectionCode="NCBS" country="USA" httpUri="http://grbio.org/cool/x0m9-a3bs" name="Yale University" pageId="3" pageNumber="220">AG829</specimenCode>
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from a
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<collectingCounty id="62AA4E74FFFDF44AFE73313EFDACFABB" box="[481,517,1323,1348]" pageId="3" pageNumber="220">Sal</collectingCounty>
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plantation near
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<collectingMunicipality id="6BAFAC82FFFDF44AFD2B313EFCE3FABB" box="[697,842,1323,1348]" pageId="3" pageNumber="220">Garjee forest</collectingMunicipality>
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rest house,
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<quantity id="4C8C9B1DFFFDF44AFC5C313EFC56FABB" box="[974,1023,1323,1348]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="220" unit="km" value="8.0">8km</quantity>
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south of
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<location id="8EAB6023FFFDF44AFBF5313EFB6CFABB" LSID="urn:lsid:plazi:treatment:03DD87EEFFFDF440FF053044FF58FD0F:8EAB6023FFFDF44AFBF5313EFB6CFABB" box="[1127,1221,1323,1348]" county="Sal" latitude="23.44166" longLatPrecision="1" longitude="91.48956" municipality="Garjee forest" name="Udaipur" pageId="3" pageNumber="220" stateProvince="Tripura">Udaipur</location>
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,
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<collectingRegion id="49B0F81AFFFDF44AFB5D313EFA8BFABB" box="[1231,1314,1323,1348]" country="India" name="Tripura" pageId="3" pageNumber="220">Tripura</collectingRegion>
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||
<geoCoordinate id="EE40503FFFFDF44AFABA313EFF07FA95" direction="north" orientation="latitude" pageId="3" pageNumber="220" precision="1" value="23.44166">23.44166° N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE40503FFFFDF44AFF2B3147FEECFA95" box="[185,325,1362,1387]" direction="east" orientation="longitude" pageId="3" pageNumber="220" precision="1" value="91.48956">91.48956° E</geoCoordinate>
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||
(WGS84),
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||
<elevation id="0059D1CBFFFDF44AFE553147FDCFFA94" box="[455,614,1362,1387]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="220" unit="m" value="56.0">
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||
elevation
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||
<quantity id="4C8C9B1DFFFDF44AFDA73147FDCFFA94" box="[565,614,1362,1387]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="220" unit="m" value="56.0">56m</quantity>
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||
</elevation>
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||
on
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||
<collectingDate id="EF8EE9D0FFFDF44AFD033146FCE6FA94" box="[657,847,1359,1387]" pageId="3" pageNumber="220" value="2014-10-09">
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||
<date id="FFCA1038FFFDF44AFD033146FCE6FA94" box="[657,847,1359,1387]" pageId="3" pageNumber="220" value="2014-10-09">9th October 2014</date>
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||
</collectingDate>
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||
.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BCB36F8FFFDF44AFF553163FD92FA25" blockId="3.[151,1437,1212,2038]" pageId="3" pageNumber="220">
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||
<emphasis id="B900EAEAFFFDF44AFF553163FE0EFA70" bold="true" box="[199,423,1398,1423]" pageId="3" pageNumber="220">Referred material:</emphasis>
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||
<collectionCode id="ED65AE3DFFFDF44AFE3C3162FE5EFA70" box="[430,503,1399,1423]" country="USA" httpUri="http://grbio.org/cool/x0m9-a3bs" name="Yale University" pageId="3" pageNumber="220">NCBS</collectionCode>
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AG821, female from Trishna WLS, same data as
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<typeStatus id="54CF885AFFFDF44AFBB93163FB39FA70" box="[1067,1168,1398,1423]" pageId="3" pageNumber="220" type="holotype">holotype</typeStatus>
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||
,
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||
<collectionCode id="ED65AE3DFFFDF44AFB093162FB4DFA70" box="[1179,1252,1399,1423]" country="USA" httpUri="http://grbio.org/cool/x0m9-a3bs" name="Yale University" pageId="3" pageNumber="220">NCBS</collectionCode>
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AW215, female from Bidyabil, Tulashikhar Tehsil,
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<collectingRegion id="49B0F81AFFFDF44AFDBB318FFDD5FA4C" box="[553,636,1434,1459]" country="India" name="Tripura" pageId="3" pageNumber="220">Tripura</collectingRegion>
|
||
(
|
||
<geoCoordinate id="EE40503FFFFDF44AFD1E318FFC98FA4D" box="[652,817,1434,1459]" direction="north" orientation="latitude" pageId="3" pageNumber="220" precision="1" value="24.14591">24.145910° N</geoCoordinate>
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||
, 91.699040°, E (WGS84), elevation
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||
<quantity id="4C8C9B1DFFFDF44AFB45318FFAA0FA4C" box="[1239,1289,1434,1459]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="220" unit="m" value="65.0">65m</quantity>
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collected on
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<date id="FFCA1038FFFDF44AFF0531D4FE9DFA25" box="[151,308,1470,1498]" pageId="3" pageNumber="220" value="2016-07-20">20th July 2016</date>
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||
by Akshay Khandekar.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36E6573FFFDF44AFF5531F1FCE0F852" pageId="3" pageNumber="220" type="diagnosis">
|
||
<paragraph id="8BCB36F8FFFDF44AFF5531F1FAE2F929" blockId="3.[151,1437,1212,2038]" pageId="3" pageNumber="220">
|
||
<emphasis id="B900EAEAFFFDF44AFF5531F1FEEAFA02" bold="true" box="[199,323,1508,1533]" pageId="3" pageNumber="220">Diagnosis:</emphasis>
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A fairly large species in relation to members of the genus reaching a total length of
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||
<quantity id="4C8C9B1DFFFDF44AFA9D31F1FA30FA02" box="[1295,1433,1508,1533]" metricMagnitude="-2" metricUnit="m" metricValue="3.675" metricValueMax="4.05" metricValueMin="3.3" pageId="3" pageNumber="220" unit="mm" value="36.75" valueMax="40.5" valueMin="33.0">33–40.5mm</quantity>
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||
. Overall coloration in a shade of blackish brown to light brown. Second lateral ocellus adjacent to the first ocellus, separated from the third ocellus. Patella prolateral process strongly developed forming a single median spine. Chela internomedian carinae present, composed of large sparsely placed granules. Metasoma: segment I, lacking pronounced granules on its ventral aspect; segment II, with posterior granules on ventrolateral and ventrosubmedian carinae, and dorso-submedian carinae lacking posterior spiniform granules; V, with three pairs of spiniform granules on ventrolateral carinae. Anal arch anterior carinae composed of large, subconical teeth.
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</paragraph>
|
||
<paragraph id="8BCB36F8FFFDF44AFF5532F4FCE0F852" blockId="3.[151,1437,1212,2038]" pageId="3" pageNumber="220">
|
||
<taxonomicName id="4C744D7BFFFDF44AFF5532F4FE32F906" authorityName="Mirza" authorityYear="2017" box="[199,411,1761,1785]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="schalleri">
|
||
<emphasis id="B900EAEAFFFDF44AFF5532F4FE32F906" box="[199,411,1761,1785]" italics="true" pageId="3" pageNumber="220">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C744D7BFFFDF44AFDBD32F7FD00F906" baseAuthorityName="Pocock" baseAuthorityYear="1897" box="[559,681,1762,1785]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="nigripes">
|
||
<emphasis id="B900EAEAFFFDF44AFDBD32F7FD00F906" box="[559,681,1762,1785]" italics="true" pageId="3" pageNumber="220">L. nigripes</emphasis>
|
||
</taxonomicName>
|
||
in bearing a strong spinoid prolateral process on patella (vs. absent in
|
||
<taxonomicName id="4C744D7BFFFDF44AFF263313FE87F8E2" baseAuthorityName="Pocock" baseAuthorityYear="1897" box="[180,302,1798,1821]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="nigripes">
|
||
<emphasis id="B900EAEAFFFDF44AFF263313FE87F8E2" box="[180,302,1798,1821]" italics="true" pageId="3" pageNumber="220">L. nigripes</emphasis>
|
||
</taxonomicName>
|
||
); differs from
|
||
<taxonomicName id="4C744D7BFFFDF44AFE463313FDDCF8E2" authorityName="Werner" authorityYear="1939" box="[468,629,1797,1821]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="longimanus">
|
||
<emphasis id="B900EAEAFFFDF44AFE463313FDDCF8E2" box="[468,629,1797,1821]" italics="true" pageId="3" pageNumber="220">L. longimanus</emphasis>
|
||
</taxonomicName>
|
||
in having a short and stout pedipalps (vs. long and slender pedipalps in
|
||
<taxonomicName id="4C744D7BFFFDF44AFF05333FFE9EF8BE" authorityName="Werner" authorityYear="1939" box="[151,311,1833,1857]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="longimanus">
|
||
<emphasis id="B900EAEAFFFDF44AFF05333FFE9EF8BE" box="[151,311,1833,1857]" italics="true" pageId="3" pageNumber="220">L. longimanus</emphasis>
|
||
</taxonomicName>
|
||
). The new species is most similar to
|
||
<taxonomicName id="4C744D7BFFFDF44AFD4A333FFC29F8BE" authorityName="Fabricius" authorityYear="1775" box="[728,896,1833,1857]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="B900EAEAFFFDF44AFD4A333FFC29F8BE" box="[728,896,1833,1857]" italics="true" pageId="3" pageNumber="220">L. australasiae</emphasis>
|
||
</taxonomicName>
|
||
with which it shares a similar spinoid prolateral process on patella.
|
||
<taxonomicName id="4C744D7BFFFDF44AFEEB3358FDF8F89A" authorityName="Mirza" authorityYear="2017" box="[377,593,1869,1893]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="schalleri">
|
||
<emphasis id="B900EAEAFFFDF44AFEEB3358FDF8F89A" box="[377,593,1869,1893]" italics="true" pageId="3" pageNumber="220">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C744D7BFFFDF44AFD62335BFC35F89A" authorityName="Fabricius" authorityYear="1775" box="[752,924,1869,1893]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="B900EAEAFFFDF44AFD62335BFC35F89A" box="[752,924,1869,1893]" italics="true" pageId="3" pageNumber="220">L. australasiae</emphasis>
|
||
</taxonomicName>
|
||
by the medial granules on the ventrolateral carinae metasomal segment II with medial granules on ventrolateral carinae which are less pronounced than the preceding segment in
|
||
<taxonomicName id="4C744D7BFFFDF44AFE1F3383FDA4F852" authorityName="Mirza" authorityYear="2017" box="[397,525,1941,1965]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="schalleri">
|
||
<emphasis id="B900EAEAFFFDF44AFE1F3383FDA4F852" box="[397,525,1941,1965]" italics="true" pageId="3" pageNumber="220">L. schalleri</emphasis>
|
||
</taxonomicName>
|
||
but large in
|
||
<taxonomicName id="4C744D7BFFFDF44AFD083383FCEBF852" authorityName="Fabricius" authorityYear="1775" box="[666,834,1941,1965]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="B900EAEAFFFDF44AFD083383FCEBF852" box="[666,834,1941,1965]" italics="true" pageId="3" pageNumber="220">L. australasiae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6573FFFDF44AFF5533ACFB09F809" pageId="3" pageNumber="220" type="etymology">
|
||
<paragraph id="8BCB36F8FFFDF44AFF5533ACFB09F809" blockId="3.[151,1437,1212,2038]" pageId="3" pageNumber="220">
|
||
<emphasis id="B900EAEAFFFDF44AFF5533ACFEF8F82D" bold="true" box="[199,337,1977,2002]" pageId="3" pageNumber="220">Etymology:</emphasis>
|
||
The specific epithet is a patronym honoring Wildlife Biologist Dr. George Beals Schaller of the Wildlife Conservation Society and Panthera, for his contribution to conservation of wildlife.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0B6670FFFAF44DFF05369BFD81FD3D" httpUri="https://zenodo.org/record/1117566/files/figure.png" pageId="4" pageNumber="221" targetBox="[163,1419,185,642]" targetPageId="4">
|
||
<paragraph id="8BCB36F8FFFAF44DFF05369BFD81FD3D" blockId="4.[151,1436,654,706]" pageId="4" pageNumber="221">
|
||
<emphasis id="B900EAEAFFFAF44DFF05369BFEBCFD5C" bold="true" box="[151,277,654,676]" pageId="4" pageNumber="221">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C744D7BFFFAF44DFE8C369BFE48FD5B" authority="Mirza, 2017" authorityName="Mirza" authorityYear="2017" box="[286,481,654,676]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="221" phylum="Arthropoda" rank="species" species="schalleri" status="sp. nov.">
|
||
<emphasis id="B900EAEAFFFAF44DFE8C369BFE48FD5B" box="[286,481,654,676]" italics="true" pageId="4" pageNumber="221">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2335791FFFAF44DFE79369BFD97FD5C" box="[491,574,654,675]" pageId="4" pageNumber="221" rank="species">
|
||
<emphasis id="B900EAEAFFFAF44DFE79369BFD97FD5C" bold="true" box="[491,574,654,675]" pageId="4" pageNumber="221">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
showing coloration in life (A) holotype male NCBS AG817, (B) paratype female NCBS AG818. Photo by Rajesh Sanap.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0B6670FFFAF44DFF0533ABFE99F80C" httpUri="https://zenodo.org/record/1117568/files/figure.png" pageId="4" pageNumber="221" targetBox="[181,1403,743,1967]" targetPageId="4">
|
||
<paragraph id="8BCB36F8FFFAF44DFF0533ABFE99F80C" blockId="4.[151,1436,1982,2035]" pageId="4" pageNumber="221">
|
||
<emphasis id="B900EAEAFFFAF44DFF0533ABFEBBF82B" bold="true" box="[151,274,1982,2004]" pageId="4" pageNumber="221">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C744D7BFFFAF44DFE8A33ABFE7EF82B" authority="Mirza, 2017" authorityName="Mirza" authorityYear="2017" box="[280,471,1982,2004]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="221" phylum="Arthropoda" rank="species" species="schalleri" status="sp. nov.">
|
||
<emphasis id="B900EAEAFFFAF44DFE8A33ABFE7EF82B" box="[280,471,1982,2004]" italics="true" pageId="4" pageNumber="221">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2335791FFFAF44DFE4E33AAFD85F82B" box="[476,556,1983,2004]" pageId="4" pageNumber="221" rank="species">
|
||
<emphasis id="B900EAEAFFFAF44DFE4E33AAFD85F82B" bold="true" box="[476,556,1983,2004]" pageId="4" pageNumber="221">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(A) male dorsal NCBS AG817, (B) male ventral NCBS AG817, (C) female dorsal, (D) female ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36E6573FFFBF440FF553482FF58FD0F" lastPageId="9" lastPageNumber="226" pageId="5" pageNumber="222" type="description">
|
||
<paragraph id="8BCB36F8FFFBF44CFF553482FCA8FE77" blockId="5.[151,1437,151,717]" pageId="5" pageNumber="222">
|
||
<emphasis id="B900EAEAFFFBF44CFF553482FD74FF4F" bold="true" box="[199,733,151,176]" pageId="5" pageNumber="222">
|
||
Description of
|
||
<typeStatus id="54CF885AFFFBF44CFEE83482FE48FF4F" box="[378,481,151,176]" pageId="5" pageNumber="222" type="holotype">holotype</typeStatus>
|
||
male NCBS-AG817:
|
||
</emphasis>
|
||
Coloration: ‘In life’ (
|
||
<figureCitation id="134F2A7DFFFBF44CFC453482FBEFFF4F" box="[983,1094,151,176]" captionStart-0="FIGURE 4" captionStart-1="FIGURE5" captionStartId-0="4.[151,250,654,676]" captionStartId-1="4.[151,250,1982,2004]" captionTargetBox-0="[163,1419,185,642]" captionTargetBox-1="[181,1403,743,1967]" captionTargetId-0="figure@4.[151,1436,173,652]" captionTargetId-1="figure@4.[167,1431,722,1988]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="FIGURE 4. Liocheles schalleri sp. nov. showing coloration in life (A) holotype male NCBS AG817, (B) paratype female NCBS AG818.Photo by Rajesh Sanap." captionText-1="FIGURE5. Liochelesschallerisp. nov. (A) male dorsal NCBS AG817, (B)male ventral NCBS AG817, (C)femaledorsal,(D) female ventral." httpUri-0="https://zenodo.org/record/1117566/files/figure.png" httpUri-1="https://zenodo.org/record/1117568/files/figure.png" pageId="5" pageNumber="222">Figs. 4–5</figureCitation>
|
||
): Overall in a shade of glossy black, extremities of legs with a brownish tinge. Color faded after preservation, the black coloration on the carapace, mesosoma and metasoma is retained; however, on the legs and pedipalps the color has faded to dark brown. Chelicerae yellowish brown with dark brown extremities of fingers and reticulate markings on cheliceral tibia. Telson pale yellow. Sternum anteriorly brownish grey gradually becoming tan in its posterior half. Pectines and pectinal plate pale yellow. Sternites brownish grey for more than half their length anteriorly excluding the stigmata and posterior half which are yellowish brown.
|
||
</paragraph>
|
||
<paragraph id="8BCB36F8FFFBF44CFF553581FAEEFD32" blockId="5.[151,1437,151,717]" pageId="5" pageNumber="222">
|
||
Carapace (
|
||
<figureCitation id="134F2A7DFFFBF44CFED13581FE0AFE52" box="[323,419,404,429]" captionStart="FIGURE6" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[151,1436,793,1903]" captionTargetId="figure@5.[151,1436,793,1903]" captionTargetPageId="5" captionText="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="5" pageNumber="222">Fig. 6A</figureCitation>
|
||
): Overall brown, borders and area surrounding the eye black. Surface of the carapace punctuate throughout except for two smooth, glossy patches between lateral and median oculi. Anterior margin with a subtle indentation scarcely forming a shallow cavity, with nearly a horizontal orientation. Posterior median furrow anterior to the posterior border of carapace deep, posterior lateral furrow on each side of the median furrow shallow (
|
||
<figureCitation id="134F2A7DFFFBF44CFF6C3631FEF1FDC2" box="[254,344,548,573]" captionStart="FIGURE6" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[151,1436,793,1903]" captionTargetId="figure@5.[151,1436,793,1903]" captionTargetPageId="5" captionText="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="5" pageNumber="222">Fig. 6A</figureCitation>
|
||
). Median ocelli lacking a raised ocular tubercle. Median ocelli situated anteriorly in the ration of 1:2.2. Second lateral ocellus adjacent to the first ocellus, separated from the third ocellus. Chelicerae immovable finger with fused basal and middle tooth into a bicusp, basal tooth much smaller than median tooth. Movable finger with four teeth, basal and sub-basal touching their bases; distal and sub-distal adjacent separated from the basal group by a very short diastema. Ratio of cheliceral movable finger length to chelicerae tibia length is 1:1.5.
|
||
</paragraph>
|
||
<caption id="DF0B6670FFFBF44CFF0533B2FB29F824" httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="5" pageNumber="222" targetBox="[151,1436,793,1903]" targetPageId="5">
|
||
<paragraph id="8BCB36F8FFFBF44CFF0533B2FB29F824" blockId="5.[151,1436,1959,2011]" pageId="5" pageNumber="222">
|
||
<emphasis id="B900EAEAFFFBF44CFF0533B2FEBBF843" bold="true" box="[151,274,1959,1981]" pageId="5" pageNumber="222">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C744D7BFFFBF44CFE8B33B2FE70F842" authority="Mirza, 2017" authorityName="Mirza" authorityYear="2017" box="[281,473,1959,1981]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="222" phylum="Arthropoda" rank="species" species="schalleri" status="sp. nov.">
|
||
<emphasis id="B900EAEAFFFBF44CFE8B33B2FE70F842" box="[281,473,1959,1981]" italics="true" pageId="5" pageNumber="222">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2335791FFFBF44CFE4D33B2FD86F843" box="[479,559,1959,1980]" pageId="5" pageNumber="222" rank="species">
|
||
<emphasis id="B900EAEAFFFBF44CFE4D33B2FD86F843" bold="true" box="[479,559,1959,1980]" pageId="5" pageNumber="222">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patella external, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomal segment II.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0B6670FFF8F44FFF053326FE39F897" httpUri="https://zenodo.org/record/1117572/files/figure.png" pageId="6" pageNumber="223" targetBox="[181,1405,185,1832]" targetPageId="6">
|
||
<paragraph id="8BCB36F8FFF8F44FFF053326FE39F897" blockId="6.[151,1436,1843,1896]" pageId="6" pageNumber="223">
|
||
<emphasis id="B900EAEAFFF8F44FFF053326FEBDF8B6" bold="true" box="[151,276,1843,1865]" pageId="6" pageNumber="223">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C744D7BFFF8F44FFE8E3326FE74F8B6" authority="Mirza, 2017" authorityName="Mirza" authorityYear="2017" box="[284,477,1843,1865]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="223" phylum="Arthropoda" rank="species" species="schalleri" status="sp. nov.">
|
||
<emphasis id="B900EAEAFFF8F44FFE8E3326FE74F8B6" box="[284,477,1843,1865]" italics="true" pageId="6" pageNumber="223">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2335791FFF8F44FFE763321FD9EF8B6" box="[484,567,1844,1865]" pageId="6" pageNumber="223" rank="species">
|
||
<emphasis id="B900EAEAFFF8F44FFE763321FD9EF8B6" bold="true" box="[484,567,1844,1865]" pageId="6" pageNumber="223">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
holotype male pedipalp chela showing tricobothrial pattern (A) dorsal, (B) external, (C) ventral, (D) internal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36F8FFF8F44FFF553380FA34F80A" blockId="6.[151,1437,1940,2037]" pageId="6" pageNumber="223">Mesosoma: All mesosomal tergites in a shade of blackish brown. Sternites tan yellow with darker posterior and lateral borders. Sternite VII dark brown to grey in color with slight tan yellow coloration in the anterior median portion of the segment. Sternites smooth glossy throughout, except for sternite VII which is granular throughout.</paragraph>
|
||
<paragraph id="8BCB36F8FFF9F44EFF053482FEF4FEE2" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="224">Tergites I–VII: with coarse granulation throughout; median carinae smooth and well developed. Sternum pentagonal, as wide as long. Genital operculum oval shaped, divided into two halves and with two genital papillae protruding from its posterior border. Pectine teeth well developed, seven each side. Pectinal plate with a median furrow anteriorly.</paragraph>
|
||
<paragraph id="8BCB36F8FFF9F44EFF55353DFC2AFDE7" blockId="7.[151,1437,151,536]" pageId="7" pageNumber="224">
|
||
Metasoma (
|
||
<figureCitation id="134F2A7DFFF9F44EFED93532FE4DFEBF" box="[331,484,295,320]" captionStart="FIGURE6" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[151,1436,793,1903]" captionTargetId="figure@5.[151,1436,793,1903]" captionTargetPageId="5" captionText="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="7" pageNumber="224">Fig. 6E & 6F</figureCitation>
|
||
): Metasomal segment I dark brown, segment II–V blackish brown. Telson tan yellow in color. Dorsal median furrow almost indistinct. Granular throughout excluding posterior half of segment V. Ventromedian and ventro-submedian carinae poorly developed, moderately developed on segment II–IV with scattered granules. Segment I lacking pronounced granules on its ventral surface (
|
||
<figureCitation id="134F2A7DFFF9F44EFBF53581FB6DFE52" box="[1127,1220,404,429]" captionStart="FIGURE6" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[151,1436,793,1903]" captionTargetId="figure@5.[151,1436,793,1903]" captionTargetPageId="5" captionText="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="7" pageNumber="224">Fig. 6E</figureCitation>
|
||
). Segment II with posterior granules on ventrolateral and ventrosubmedian carinae (
|
||
<figureCitation id="134F2A7DFFF9F44EFC1435A2FC74FE2F" box="[902,989,439,464]" captionStart="FIGURE6" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[151,1436,793,1903]" captionTargetId="figure@5.[151,1436,793,1903]" captionTargetPageId="5" captionText="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="7" pageNumber="224">Fig. 6F</figureCitation>
|
||
). Segment II, dorso-submedian carinae lacking posterior spiniform granules. Segment V with three pairs of spiniform granules on ventrolateral carinae. Anal arch anterior carinae composed of large, subconical granules.
|
||
</paragraph>
|
||
<caption id="DF0B6670FFF9F44EFF05321EFF43F9BF" httpUri="https://zenodo.org/record/1117574/files/figure.png" pageId="7" pageNumber="224" targetBox="[412,1175,597,1528]" targetPageId="7">
|
||
<paragraph id="8BCB36F8FFF9F44EFF05321EFF43F9BF" blockId="7.[151,1436,1547,1600]" pageId="7" pageNumber="224">
|
||
<emphasis id="B900EAEAFFF9F44EFF05321EFEBDF9DF" bold="true" box="[151,276,1547,1569]" pageId="7" pageNumber="224">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C744D7BFFF9F44EFE8E321EFE77F9DE" authority="Mirza, 2017" authorityName="Mirza" authorityYear="2017" box="[284,478,1547,1569]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="schalleri" status="sp. nov.">
|
||
<emphasis id="B900EAEAFFF9F44EFE8E321EFE77F9DE" box="[284,478,1547,1569]" italics="true" pageId="7" pageNumber="224">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EAEAFFF9F44EFE75321EFD90F9DF" bold="true" box="[487,569,1547,1568]" pageId="7" pageNumber="224">
|
||
<taxonomicNameLabel id="A2335791FFF9F44EFE75321EFD90F9DF" box="[487,569,1547,1568]" pageId="7" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype male NCBS AG817 hemispermataphore, (A) dorsal view, (B) close-up of capsule.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36F8FFF9F44EFF553272FDB6F881" blockId="7.[151,1436,1639,2027]" pageId="7" pageNumber="224">
|
||
Pedipalp (
|
||
<figureCitation id="134F2A7DFFF9F44EFEAB3272FE52F97F" box="[313,507,1639,1664]" captionStart="FIGURE6" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[151,1436,793,1903]" captionTargetId="figure@5.[151,1436,793,1903]" captionTargetPageId="5" captionText="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="7" pageNumber="224">Figs. 6B, 6C, 6D</figureCitation>
|
||
,
|
||
<figureCitation id="134F2A7DFFF9F44EFD953272FDB1F980" box="[519,536,1639,1663]" captionStart="FIGURE 7" captionStartId="6.[151,250,1843,1865]" captionTargetBox="[181,1405,185,1832]" captionTargetId="figure@6.[181,1405,185,1832]" captionTargetPageId="6" captionText="FIGURE 7. Liocheles schalleri sp. nov. holotype male pedipalp chela showing tricobothrial pattern (A) dorsal, (B) external, (C) ventral, (D) internal." httpUri="https://zenodo.org/record/1117572/files/figure.png" pageId="7" pageNumber="224">7</figureCitation>
|
||
): Pedipalp femur, patella and chela reddish brown dorsally, slightly paler on the venter. Fingers of pedipalp chela darker with the tip of the fingers with a reddish tinge. Coloration darker on carinae Chela width to length ration 1:2.9. Pedipalp chela with 26 trichobothria;
|
||
<emphasis id="B900EAEAFFF9F44EFBBD32BAFBE2F938" box="[1071,1099,1711,1735]" italics="true" pageId="7" pageNumber="224">db</emphasis>
|
||
,
|
||
<emphasis id="B900EAEAFFF9F44EFBC832BAFBDCF938" box="[1114,1141,1711,1735]" italics="true" pageId="7" pageNumber="224">eb</emphasis>
|
||
3,
|
||
<emphasis id="B900EAEAFFF9F44EFB1E32BAFB18F938" box="[1164,1201,1711,1735]" italics="true" pageId="7" pageNumber="224">esb</emphasis>
|
||
aligned in a straight vertical line (
|
||
<figureCitation id="134F2A7DFFF9F44EFEBC32C0FE2EF911" box="[302,391,1749,1774]" captionStart="FIGURE 7" captionStartId="6.[151,250,1843,1865]" captionTargetBox="[181,1405,185,1832]" captionTargetId="figure@6.[181,1405,185,1832]" captionTargetPageId="6" captionText="FIGURE 7. Liocheles schalleri sp. nov. holotype male pedipalp chela showing tricobothrial pattern (A) dorsal, (B) external, (C) ventral, (D) internal." httpUri="https://zenodo.org/record/1117572/files/figure.png" pageId="7" pageNumber="224">Fig. 7B</figureCitation>
|
||
);
|
||
<emphasis id="B900EAEAFFF9F44EFE0F32C3FE1AF911" box="[413,435,1750,1774]" italics="true" pageId="7" pageNumber="224">dt</emphasis>
|
||
on internal aspect of fixed finger with
|
||
<emphasis id="B900EAEAFFF9F44EFCE332C2FC14F911" box="[881,957,1750,1774]" italics="true" pageId="7" pageNumber="224">it & ib</emphasis>
|
||
(
|
||
<figureCitation id="134F2A7DFFF9F44EFC5F32C0FB81F911" box="[973,1064,1749,1774]" captionStart="FIGURE 7" captionStartId="6.[151,250,1843,1865]" captionTargetBox="[181,1405,185,1832]" captionTargetId="figure@6.[181,1405,185,1832]" captionTargetPageId="6" captionText="FIGURE 7. Liocheles schalleri sp. nov. holotype male pedipalp chela showing tricobothrial pattern (A) dorsal, (B) external, (C) ventral, (D) internal." httpUri="https://zenodo.org/record/1117572/files/figure.png" pageId="7" pageNumber="224">Fig. 7D</figureCitation>
|
||
);
|
||
<emphasis id="B900EAEAFFF9F44EFBAC32C3FBC1F911" box="[1086,1128,1750,1774]" italics="true" pageId="7" pageNumber="224">Esb</emphasis>
|
||
,
|
||
<emphasis id="B900EAEAFFF9F44EFBE532C3FB69F911" box="[1143,1216,1750,1774]" italics="true" pageId="7" pageNumber="224">
|
||
Eb
|
||
<quantity id="4C8C9B1DFFF9F44EFB0532C3FB69F911" box="[1175,1216,1750,1774]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="7" pageNumber="224" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">1–3</quantity>
|
||
</emphasis>
|
||
in a group in basal region. Pedipalp chela manus granular, immovable and movable finger smooth. Dentate margin of pedipalp chela finger with a distinct sculpture (
|
||
<figureCitation id="134F2A7DFFF9F44EFD983308FDCCF8C9" box="[522,613,1821,1846]" captionStart="FIGURE 7" captionStartId="6.[151,250,1843,1865]" captionTargetBox="[181,1405,185,1832]" captionTargetId="figure@6.[181,1405,185,1832]" captionTargetPageId="6" captionText="FIGURE 7. Liocheles schalleri sp. nov. holotype male pedipalp chela showing tricobothrial pattern (A) dorsal, (B) external, (C) ventral, (D) internal." httpUri="https://zenodo.org/record/1117572/files/figure.png" pageId="7" pageNumber="224">Fig. 7B</figureCitation>
|
||
). Patella prolateral view with a median spine like protrusion (
|
||
<figureCitation id="134F2A7DFFF9F44EFAA03308FA26F8C9" box="[1330,1423,1821,1846]" captionStart="FIGURE6" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[151,1436,793,1903]" captionTargetId="figure@5.[151,1436,793,1903]" captionTargetPageId="5" captionText="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="7" pageNumber="224">Fig. 6D</figureCitation>
|
||
). Patella retro-lateral basally with a shallow concavity. Chela internomedian carinae present, composed of large sparsely placed granules (
|
||
<figureCitation id="134F2A7DFFF9F44EFE2A3370FDBBF881" box="[440,530,1893,1918]" captionStart="FIGURE 7" captionStartId="6.[151,250,1843,1865]" captionTargetBox="[181,1405,185,1832]" captionTargetId="figure@6.[181,1405,185,1832]" captionTargetPageId="6" captionText="FIGURE 7. Liocheles schalleri sp. nov. holotype male pedipalp chela showing tricobothrial pattern (A) dorsal, (B) external, (C) ventral, (D) internal." httpUri="https://zenodo.org/record/1117572/files/figure.png" pageId="7" pageNumber="224">Fig. 7D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCB36F8FFF9F44EFF55339EFD10F814" blockId="7.[151,1436,1639,2027]" pageId="7" pageNumber="224">Legs: Dorsally in a shade of dark brown and lighter ventrally. Dorsal surfaces of trochanter, femur & patella granular, basitarsus and tarsus smooth and glossy. Tarsi lacking median row of ventromedian spinules. Two rows of ventral macrosetae. Tarsal claws of equal length.</paragraph>
|
||
<caption id="DF0B6670FFF6F441FF053736FED4FC89" httpUri="https://zenodo.org/record/1117576/files/figure.png" pageId="8" pageNumber="225" targetBox="[151,1436,193,781]" targetPageId="8">
|
||
<paragraph id="8BCB36F8FFF6F441FF053736FED4FC89" blockId="8.[151,1436,803,886]" pageId="8" pageNumber="225">
|
||
<emphasis id="B900EAEAFFF6F441FF053736FEBCFCC7" bold="true" box="[151,277,803,825]" pageId="8" pageNumber="225">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C744D7BFFF6F441FE8F3736FE49FCC6" authority="Mirza, 2017" authorityName="Mirza" authorityYear="2017" box="[285,480,803,825]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="225" phylum="Arthropoda" rank="species" species="schalleri" status="sp. nov.">
|
||
<emphasis id="B900EAEAFFF6F441FE8F3736FE49FCC6" box="[285,480,803,825]" italics="true" pageId="8" pageNumber="225">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2335791FFF6F441FE7B3736FD92FCC7" box="[489,571,803,824]" pageId="8" pageNumber="225" rank="species">
|
||
<emphasis id="B900EAEAFFF6F441FE7B3736FD92FCC7" bold="true" box="[489,571,803,824]" pageId="8" pageNumber="225">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
female NCBS AG818, (A) femur dorsal, (B) patella prolateral process, (C) manus internal view, (D) ventral view of metasomal segment I, (E) ventral view of metasomal segment II, (F) dorsal view of metasomal segment II.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0B6670FFF6F441FF053339FB6CF8BE" box="[151,1221,1836,1858]" httpUri="https://zenodo.org/record/1117578/files/figure.png" pageId="8" pageNumber="225" targetBox="[191,1331,944,1813]" targetPageId="8">
|
||
<paragraph id="8BCB36F8FFF6F441FF053339FB6CF8BE" blockId="8.[151,1221,1836,1858]" box="[151,1221,1836,1858]" pageId="8" pageNumber="225">
|
||
<emphasis id="B900EAEAFFF6F441FF053339FEB6F8BE" bold="true" box="[151,287,1836,1858]" pageId="8" pageNumber="225">FIGURE 10.</emphasis>
|
||
Map of eastern northeastern India showing known localities for
|
||
<taxonomicName id="4C744D7BFFF6F441FC3C3339FBC6F8BD" authority="Mirza, 2017" authorityName="Mirza" authorityYear="2017" box="[942,1135,1836,1858]" class="Arachnida" family="Hormuridae" genus="Liocheles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="225" phylum="Arthropoda" rank="species" species="schalleri" status="sp. nov.">
|
||
<emphasis id="B900EAEAFFF6F441FC3C3339FBC6F8BD" box="[942,1135,1836,1858]" italics="true" pageId="8" pageNumber="225">Liocheles schalleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2335791FFF6F441FBE63339FB6CF8BE" box="[1140,1221,1836,1857]" pageId="8" pageNumber="225" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB36F8FFF6F441FF55337BFACCF80B" blockId="8.[151,1436,1902,2036]" pageId="8" pageNumber="225">
|
||
Hemispermatophore (
|
||
<figureCitation id="134F2A7DFFF6F441FE2E337BFDABF878" box="[444,514,1902,1927]" captionStart="FIGURE 8" captionStartId="7.[151,250,1547,1569]" captionTargetBox="[412,1175,597,1528]" captionTargetId="figure@7.[402,1185,589,1537]" captionTargetPageId="7" captionText="FIGURE 8. Liocheles schalleri sp. nov. holotype male NCBS AG817 hemispermataphore, (A) dorsal view, (B) close-up of capsule." httpUri="https://zenodo.org/record/1117574/files/figure.png" pageId="8" pageNumber="225">Fig. 8</figureCitation>
|
||
): Lamelliform with a complex capsule; distal lamellum stout, with a rounded tip and a slight protrusion at the apex (
|
||
<figureCitation id="134F2A7DFFF6F441FDA73386FD3BF854" box="[565,658,1939,1964]" captionStart="FIGURE 8" captionStartId="7.[151,250,1547,1569]" captionTargetBox="[412,1175,597,1528]" captionTargetId="figure@7.[402,1185,589,1537]" captionTargetPageId="7" captionText="FIGURE 8. Liocheles schalleri sp. nov. holotype male NCBS AG817 hemispermataphore, (A) dorsal view, (B) close-up of capsule." httpUri="https://zenodo.org/record/1117574/files/figure.png" pageId="8" pageNumber="225">Fig. 8A</figureCitation>
|
||
). Flagellum length equal to that of the capsule region and trunk. A single lamellar hook arising from the transverse ridge, hook short, curved and pointing upwards. Lamellae arising from the posterior lobe is clearly apart from the lamellar hook, vertical in orientation with a slight angular tilt.
|
||
</paragraph>
|
||
<paragraph id="8BCB36F8FFF7F440FF553482FF68FE53" blockId="9.[151,1437,151,752]" pageId="9" pageNumber="226">
|
||
<emphasis id="B900EAEAFFF7F440FF553482FEE5FF4F" bold="true" box="[199,332,151,176]" pageId="9" pageNumber="226">Variations:</emphasis>
|
||
Morphometric variation is presented in
|
||
<tableCitation id="C6F60343FFF7F440FCA33482FC23FF4F" box="[817,906,151,176]" captionStart="TABLE 1" captionStartId="9.[151,239,797,820]" captionText="TABLE 1. Morphometry of type specimens of Liocheles schalleri sp. nov." pageId="9" pageNumber="226">Table 1</tableCitation>
|
||
, which highlight notable sexual dimorphism especially with regards to the robustness of the pedipalp chela in the males and can also be seen in the graphic output from PCA (
|
||
<figureCitation id="134F2A7DFFF7F440FEE734CAFE62FF07" box="[373,459,223,248]" captionStart="FIGURE 11" captionStartId="10.[151,250,1021,1043]" captionTargetBox="[190,1356,208,999]" captionTargetId="figure@10.[175,1419,194,1010]" captionTargetPageId="10" captionText="FIGURE 11. Multivariate PCA plot. Blue- L. nigripes, black- Liocheles schalleri sp. nov., male indicated by solid square and females by solid circles." httpUri="https://zenodo.org/record/1117580/files/figure.png" pageId="9" pageNumber="226">Fig. 11</figureCitation>
|
||
). The males of this species are paler in coloration in comparison with the females. Ventral spiniform granules on metasomal segment I & II more pronounced in males, in females these are more depressed and less demarcated (
|
||
<figureCitation id="134F2A7DFFF7F440FD943532FD2AFEBF" box="[518,643,295,320]" captionStart="FIGURE 9" captionStartId="8.[151,250,803,825]" captionTargetBox="[151,1436,193,781]" captionTargetId="figure@8.[151,1436,193,781]" captionTargetPageId="8" captionText="FIGURE 9. Liocheles schalleri sp. nov. female NCBS AG818, (A) femur dorsal, (B) patella prolateral process, (C) manus internal view, (D) ventral view of metasomal segment I, (E) ventral view of metasomal segment II, (F) dorsal view of metasomal segment II." httpUri="https://zenodo.org/record/1117576/files/figure.png" pageId="9" pageNumber="226">Fig. 9D–E</figureCitation>
|
||
). Dentate margin of pedipalp chela finger with a distinct sculpture in males, absent in females. Carinae on patella of males composed of large tubercles in basal half and smooth in the distal half (
|
||
<figureCitation id="134F2A7DFFF7F440FE87357AFEC4FE77" box="[277,365,367,392]" captionStart="FIGURE6" captionStartId="5.[151,250,1959,1981]" captionTargetBox="[151,1436,793,1903]" captionTargetId="figure@5.[151,1436,793,1903]" captionTargetPageId="5" captionText="FIGURE6. Liocheles schalleri sp. nov. holotype male NCBS AG817 (A) carapace, (B) femur dorsal, (C) patellaexternal, (D) patella dorsal, (E) ventral view of metasomal segment I, (F) ventral view of metasomalsegment II." httpUri="https://zenodo.org/record/1117570/files/figure.png" pageId="9" pageNumber="226">Fig. 6D</figureCitation>
|
||
) whereas the carinae in females is smooth in its basal half and tuberculate in its distal half (
|
||
<figureCitation id="134F2A7DFFF7F440FAFD357AFF11FE53" captionStart="FIGURE 9" captionStartId="8.[151,250,803,825]" captionTargetBox="[151,1436,193,781]" captionTargetId="figure@8.[151,1436,193,781]" captionTargetPageId="8" captionText="FIGURE 9. Liocheles schalleri sp. nov. female NCBS AG818, (A) femur dorsal, (B) patella prolateral process, (C) manus internal view, (D) ventral view of metasomal segment I, (E) ventral view of metasomal segment II, (F) dorsal view of metasomal segment II." httpUri="https://zenodo.org/record/1117576/files/figure.png" pageId="9" pageNumber="226">Fig. 9B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BCB36F8FFF7F440FF5535A2FF58FD0F" blockId="9.[151,1437,151,752]" pageId="9" pageNumber="226">
|
||
<emphasis id="B900EAEAFFF7F440FF5535A2FE23FE2F" bold="true" box="[199,394,439,464]" pageId="9" pageNumber="226">Natural history:</emphasis>
|
||
Specimens of this species were found in slit like burrows ~
|
||
<quantity id="4C8C9B1DFFF7F440FBBD35ADFBC6FE2F" box="[1071,1135,440,464]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="226" unit="cm" value="30.0">30cm</quantity>
|
||
deep on mud escarpments near ponds, streams and in crevices on tree trunks, among boulders. Some burrows were vertical in their orientation and others horizontal depending on presence of obstacles like roots or boulders. Most burrows were observed in mud and a few in the root system of a tree. This is a low elevation species and is found bellow 50 MSL. Burrows of this species are concentrated at a single site and completely absent from areas where suitable mud escarpments are absent. The species is recorded from Trishna Wildlife Sanctuary, Bison National Park, Garjee reserve forest and Bidyabil (
|
||
<figureCitation id="134F2A7DFFF7F440FE9B369AFEF7FD57" box="[265,350,655,680]" captionStart="FIGURE 10" captionStartId="8.[151,250,1836,1858]" captionTargetBox="[191,1331,944,1813]" captionTargetId="figure@8.[151,1436,924,1833]" captionTargetPageId="8" captionText="FIGURE 10. Map of easternnortheastern India showing known localities for Liocheles schallerisp. nov." httpUri="https://zenodo.org/record/1117578/files/figure.png" pageId="9" pageNumber="226">Fig. 10</figureCitation>
|
||
), however; is likely widespread across parts of
|
||
<collectingRegion id="49B0F81AFFF7F440FC16369AFC7EFD57" box="[900,983,655,680]" country="India" name="Tripura" pageId="9" pageNumber="226">Tripura</collectingRegion>
|
||
and likely adjoining
|
||
<collectingCountry id="F3637668FFF7F440FB5F369AFAF9FD57" box="[1229,1360,655,680]" name="Bangladesh" pageId="9" pageNumber="226">Bangladesh</collectingCountry>
|
||
as the neighboring areas of
|
||
<collectingCountry id="F3637668FFF7F440FE1D36A1FDBBFD32" box="[399,530,692,717]" name="Bangladesh" pageId="9" pageNumber="226">Bangladesh</collectingCountry>
|
||
share similar habitat and the country boarded is less than
|
||
<quantity id="4C8C9B1DFFF7F440FB5036A1FB5AFD32" box="[1218,1267,692,717]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="226" unit="km" value="3.0">3km</quantity>
|
||
from the
|
||
<typeStatus id="54CF885AFFF7F440FAFE36A0FA35FD32" box="[1388,1436,693,717]" pageId="9" pageNumber="226">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |