161 lines
18 KiB
XML
161 lines
18 KiB
XML
<document id="5E686812FED7E2E913A0B203BB002602" ID-DOI="10.5281/zenodo.188725" ID-GBIF-Dataset="97a0d3d1-80c3-450b-9404-255b5a20c7f5" ID-ISSN="1175-5326" ID-Zenodo-Dep="188725" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461096479258" checkinUser="plazi" docAuthor="Shear, William A., Taylor, Steven J., Wynne, Judson & Krejca, Jean K." docDate="2009" docId="03DD87D0AB1CCC23FF2AFC24FCF2FE82" docLanguage="en" docName="zt02151p065.pdf" docOrigin="Zootaxa 2151" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nevadesmus Shear" docType="treatment" docVersion="9" lastPageNumber="59" masterDocId="FFE4FFA8AB17CC2FFFBDFFF5FFAEFFA4" masterDocTitle="Cave millipeds of the United States. VIII. New genera and species of polydesmidan millipeds from caves in the southwestern United States (Diplopoda, Polydesmida, Macrosternodesmidae)" masterLastPageNumber="65" masterPageNumber="47" pageNumber="58" updateTime="1698589473289" updateUser="plazi">
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<mods:title id="0A81EDB1BB8892FB781A4F6BB2E46CE5">Cave millipeds of the United States. VIII. New genera and species of polydesmidan millipeds from caves in the southwestern United States (Diplopoda, Polydesmida, Macrosternodesmidae)</mods:title>
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<mods:name id="1ED6D9DFA3CCA77209585BA7D38A6AF1" type="personal">
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<mods:roleTerm id="E854A1C1F3C74E58CCB1B42892141DDD">Author</mods:roleTerm>
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<mods:namePart id="DB7D406F6F390068F0A18D820148BBC1">Shear, William A.</mods:namePart>
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<mods:roleTerm id="B627FCC05BBE5CF40AA2843587C9951F">Author</mods:roleTerm>
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<mods:namePart id="7E73F3C46D1521EF8F88593E5F9D7CAE">Taylor, Steven J.</mods:namePart>
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<mods:namePart id="3CF908606BE0840DB772EB534DACAAC7">Wynne, Judson</mods:namePart>
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<mods:name id="101D784275254FC213961274511381B5" type="personal">
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<mods:namePart id="9F3EBAE14B3B4962BD7DE1F7F32F711F">Krejca, Jean K.</mods:namePart>
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<mods:typeOfResource id="B09B6989A955D63198873550017B0AFB">text</mods:typeOfResource>
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<mods:title id="D4B4D03DA56152DEF288B161D3296417">Zootaxa</mods:title>
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<mods:date id="6ECE49405CCD6E10DBDB660798221E19">2009</mods:date>
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<mods:detail id="3C320CC305A96FAA8609AE538345DE9F" type="volume">
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<mods:number id="B9BBBB20C5DE68FB197F7D503441CBAD">2151</mods:number>
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<mods:identifier id="4FFD0E8C69D1012F5C549F1A16AC488C" type="DOI">10.5281/zenodo.188725</mods:identifier>
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<mods:identifier id="BBBB42262562EB4B12C8F57871E61AC7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03DD87D0AB1CCC23FF2AFC24FCF2FE82" ID-DOI="http://doi.org/10.5281/zenodo.6213342" ID-GBIF-Taxon="119620482" ID-Zenodo-Dep="6213342" LSID="urn:lsid:plazi:treatment:03DD87D0AB1CCC23FF2AFC24FCF2FE82" httpUri="http://treatment.plazi.org/id/03DD87D0AB1CCC23FF2AFC24FCF2FE82" lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58">
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<subSubSection id="C36E654DAB1CCC24FF2AFC24FDA2FC4E" box="[151,524,976,1003]" pageId="11" pageNumber="58" type="nomenclature">
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<paragraph id="8BCB36C6AB1CCC24FF2AFC24FDA2FC4E" blockId="11.[151,524,976,1003]" box="[151,524,976,1003]" pageId="11" pageNumber="58">
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<heading id="D08381AAAB1CCC24FF2AFC24FDA2FC4E" bold="true" box="[151,524,976,1003]" fontSize="11" level="1" pageId="11" pageNumber="58" reason="1">
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<emphasis id="B900EAD4AB1CCC24FF2AFC24FDA2FC4E" bold="true" box="[151,524,976,1003]" pageId="11" pageNumber="58">
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<taxonomicName id="4C744D45AB1CCC24FF2AFC24FED3FC4E" authority="Shear" authorityName="Shear" box="[151,381,976,1003]" class="Diplopoda" family="Macrosternodesmidae" genus="Nevadesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B900EAD4AB1CCC24FF2AFC24FE81FC4F" bold="true" box="[151,303,977,1003]" italics="true" pageId="11" pageNumber="58">Nevadesmus</emphasis>
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Shear
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</taxonomicName>
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, new genus
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E654DAB1CCC24FF2AFBEFFCA2FB95" box="[151,780,1050,1074]" pageId="11" pageNumber="58" type="materials_examined">
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<paragraph id="8BCB36C6AB1CCC24FF2AFBEFFCA2FB95" blockId="11.[151,780,1050,1074]" box="[151,780,1050,1074]" pageId="11" pageNumber="58">
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<emphasis id="B900EAD4AB1CCC24FF2AFBEFFE83FB96" bold="true" box="[151,301,1050,1074]" pageId="11" pageNumber="58">
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<typeStatus id="54CF8864AB1CCC24FF2AFBEFFF61FB96" box="[151,207,1050,1074]" pageId="11" pageNumber="58">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C744D45AB1CCC24FE8EFBEFFDD5FB95" authority="Shear" authorityName="Shear" box="[307,635,1050,1073]" class="Diplopoda" family="Macrosternodesmidae" genus="Nevadesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="species" species="ophimontis" status="sp. nov.">
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<emphasis id="B900EAD4AB1CCC24FE8EFBEFFD98FB95" box="[307,566,1050,1073]" italics="true" pageId="11" pageNumber="58">Nevadesmus ophimontis</emphasis>
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Shear
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23357AFAB1CCC24FD38FBEFFCA6FB95" box="[645,776,1050,1073]" pageId="11" pageNumber="58" rank="species">new species</taxonomicNameLabel>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E654DAB1CCC24FF2AFB95FD84FB06" pageId="11" pageNumber="58" type="etymology">
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<paragraph id="8BCB36C6AB1CCC24FF2AFB95FD84FB06" blockId="11.[151,1437,1120,2036]" pageId="11" pageNumber="58">
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<emphasis id="B900EAD4AB1CCC24FF2AFB95FE82FBDE" bold="true" box="[151,300,1120,1146]" pageId="11" pageNumber="58">Etymology:</emphasis>
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Named for the state of Nevada, with the combining stem –
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<emphasis id="B900EAD4AB1CCC24FBB9FB94FBF3FBDE" box="[1028,1117,1121,1146]" italics="true" pageId="11" pageNumber="58">desmus</emphasis>
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, traditionally used for the names of polydesmidan millipeds.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E654DAB1CCC24FF7BFB5BFC7FF942" pageId="11" pageNumber="58" type="diagnosis">
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<paragraph id="8BCB36C6AB1CCC24FF7BFB5BFD27F93C" blockId="11.[151,1437,1120,2036]" pageId="11" pageNumber="58">
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<emphasis id="B900EAD4AB1CCC24FF7BFB5BFEE6FB6C" bold="true" box="[198,328,1198,1224]" pageId="11" pageNumber="58">Diagnosis:</emphasis>
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Very small (<5.0 mm length) polydesmidan millipeds with 20 trunk segments, lacking pigment; metatergites (
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<figureCitation id="134F2A43AB1CCC24FE17FB20FE5FFB4B" box="[426,497,1237,1263]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,270,1440,1464]" captionTargetBox="[151,1434,310,1416]" captionTargetId="figure@5.[151,1436,310,1417]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Segments 10, dorsal view. Fig. 3. Nevadesmus ophimontis. Fig. 2. Pratherodesmus ecclesia. Fig. 3. P. despaini. Fig. 6. P. voylesi." httpUri="https://zenodo.org/record/188727/files/figure.png" pageId="11" pageNumber="58">Fig. 3</figureCitation>
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) with three transverse rows of short, brushlike or clavate setae (
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<figureCitation id="134F2A43AB1CCC24FB4EFB20FAE5FB4B" box="[1267,1355,1237,1263]" captionStart="FIGURES 7 – 10" captionStartId="6.[151,270,1492,1516]" captionTargetBox="[153,1436,349,1459]" captionTargetId="figure@6.[153,1438,349,1461]" captionTargetPageId="6" captionText="FIGURES 7 – 10. Figs. 7 – 9. Pygidia, dorsal view. Fig. 7. Nevadesmus ophimontis. Fig. 8. Pratherodesmus despaini. Fig. 9. P. voylesi. Fig. 10. Metazonital setae anterior to ozopore of segment 10, Nevadesmus ophimontis." httpUri="https://zenodo.org/record/188728/files/figure.png" pageId="11" pageNumber="58">Fig. 10</figureCitation>
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), rows straight, setae on low pustules. Collum ovoid, narrower than head and slightly narrower than first leg-bearing segment. Paranota low, margins not strongly toothed, posteriolateral angles right-angled to acute, but not produced into processes beyond ozopores (
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<figureCitation id="134F2A43AB1CCC24FD27FABCFD41FAC7" box="[666,751,1353,1379]" captionStart="FIGURES 3 – 6" captionStartId="5.[151,270,1440,1464]" captionTargetBox="[151,1434,310,1416]" captionTargetId="figure@5.[151,1436,310,1417]" captionTargetPageId="5" captionText="FIGURES 3 – 6. Segments 10, dorsal view. Fig. 3. Nevadesmus ophimontis. Fig. 2. Pratherodesmus ecclesia. Fig. 3. P. despaini. Fig. 6. P. voylesi." httpUri="https://zenodo.org/record/188727/files/figure.png" pageId="11" pageNumber="58">Figs. 3</figureCitation>
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,
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<figureCitation id="134F2A43AB1CCC24FD40FABCFCB0FAC7" box="[765,798,1353,1379]" captionStart="FIGURES 7 – 10" captionStartId="6.[151,270,1492,1516]" captionTargetBox="[153,1436,349,1459]" captionTargetId="figure@6.[153,1438,349,1461]" captionTargetPageId="6" captionText="FIGURES 7 – 10. Figs. 7 – 9. Pygidia, dorsal view. Fig. 7. Nevadesmus ophimontis. Fig. 8. Pratherodesmus despaini. Fig. 9. P. voylesi. Fig. 10. Metazonital setae anterior to ozopore of segment 10, Nevadesmus ophimontis." httpUri="https://zenodo.org/record/188728/files/figure.png" pageId="11" pageNumber="58">10</figureCitation>
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). Pygidium blunt, nearly hemispherical when viewed dorsally, sparsely setose, with usual four spinnerets (Shear 2008) arranged in a square and set in individual depressions; pygidial process very short, continuing line of pygidium. Males with pregonopodal legs unmodified. Gonopods with coxae globular, fixed, entirely filling gonostome, tightly appressed or fused in midline; prefemora sparsely setose, strongly transverse, articulating with coxae by process fitting into coxal notch. Exomere long, sinuous, arising from distomesal margin of prefemur, endomerite large, distally with three sinuous processes, exceeding other parts of gonopod. Acropodite bulky, solenomere incurved, nearly sessile, opening of seminal canal widened, proximal subtending process widely separated from solenomere, thin, curved, acute; no obvious tibiotarsus.
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</paragraph>
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<paragraph id="8BCB36C6AB1CCC24FF7BF950FC7FF942" blockId="11.[151,1437,1120,2036]" pageId="11" pageNumber="58">
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<taxonomicName id="4C744D45AB1CCC24FF7BF950FEF7F91A" box="[198,345,1701,1726]" class="Diplopoda" family="Macrosternodesmidae" genus="Nevadesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B900EAD4AB1CCC24FF7BF950FEF7F91A" box="[198,345,1701,1726]" italics="true" pageId="11" pageNumber="58">Nevadesmus</emphasis>
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</taxonomicName>
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differs from all previously known southwestern macrosternodesmids in its small size. It may be distinguished from
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<taxonomicName id="4C744D45AB1CCC24FE20F939FDFCF941" box="[413,594,1740,1765]" class="Diplopoda" family="Macrosternodesmidae" genus="Sequoiadesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B900EAD4AB1CCC24FE20F939FDFCF941" box="[413,594,1740,1765]" italics="true" pageId="11" pageNumber="58">Sequoiadesmus</emphasis>
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</taxonomicName>
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by its much shorter solenomere.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E654DAB1CCC23FF7BF907FCF2FE82" lastPageId="12" lastPageNumber="59" pageId="11" pageNumber="58" type="distribution">
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<paragraph id="8BCB36C6AB1CCC24FF7BF907FBCBF8A8" blockId="11.[151,1437,1120,2036]" box="[198,1125,1778,1804]" pageId="11" pageNumber="58">
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<emphasis id="B900EAD4AB1CCC24FF7BF907FEC7F8A8" bold="true" box="[198,361,1778,1804]" pageId="11" pageNumber="58">Distribution:</emphasis>
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Known from caves in White Pine and Lincoln counties, Nevada.
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</paragraph>
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<paragraph id="8BCB36C6AB1CCC24FF7BF8EDFED3F850" blockId="11.[151,1437,1120,2036]" pageId="11" pageNumber="58">
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<emphasis id="B900EAD4AB1CCC24FF7BF8EDFEB8F896" bold="true" box="[198,278,1816,1842]" pageId="11" pageNumber="58">Notes:</emphasis>
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This genus is closer in its gonopod to
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<taxonomicName id="4C744D45AB1CCC24FCB3F8ECFCD7F896" box="[782,889,1817,1842]" class="Diplopoda" genus="Tidesmus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B900EAD4AB1CCC24FCB3F8ECFCD7F896" box="[782,889,1817,1842]" italics="true" pageId="11" pageNumber="58">Tidesmus</emphasis>
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</taxonomicName>
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than to
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<emphasis id="B900EAD4AB1CCC24FC51F8ECFB1CF896" box="[1004,1202,1817,1842]" italics="true" pageId="11" pageNumber="58">
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<taxonomicName id="4C744D45AB1CCC24FC51F8ECFB00F896" box="[1004,1198,1817,1842]" class="Diplopoda" family="Macrosternodesmidae" genus="Pratherodesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">Pratherodesmus</taxonomicName>
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,
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</emphasis>
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particularly in the proportions of prefemoral processes. However, the gonopods most closely resemble those of the much larger
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<taxonomicName id="4C744D45AB1CCC24FF2AF892FE5AF824" authority="Loomis 1960" authorityName="Loomis" authorityYear="1960" box="[151,500,1894,1920]" class="Diplopoda" family="Nearctodesmidae" genus="Harpogonopus" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="58" phylum="Arthropoda" rank="genus">
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<emphasis id="B900EAD4AB1CCC24FF2AF892FEE4F824" box="[151,330,1895,1920]" italics="true" pageId="11" pageNumber="58">Harpogonopus</emphasis>
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Loomis 1960
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</taxonomicName>
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, generally accepted as a nearctodesmid (
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<bibRefCitation id="EFE54B37AB1CCC24FC61F893FBD0F824" author="Shelley" box="[988,1150,1894,1920]" pageId="11" pageNumber="65" refString="Shelley, R. M. (1993) Harpagonopus (sic) confluentus Loomis, a Pacific Coast milliped of the United States and Mexico (Polydesmida: Trichopolydesmoidea). Myriapodologica, 2, 73 - 81." type="journal article" year="1993">Shelley 1993</bibRefCitation>
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,
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<bibRefCitation id="EFE54B37AB1CCC24FB36F893FB64F824" author="Shelley" box="[1163,1226,1894,1920]" pageId="11" pageNumber="65" refString="Shelley, R. M. (1994) The milliped family Nearctodesmidae in northwestern North America, with accounts of Sakophallus and S. simplex Chamberlin (Poyldesmida), Canadian Journal of Zoology, 72, 470 - 495." type="journal article" year="1994">1994</bibRefCitation>
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; Shelley & Shear 2006). Though having the general somatic appearance of a macrosternodesmid, the gonopods of species of this genus are quite nearctodesmid-like, with both endomerite and exomere well-developed, but without an obvious tibiotarsus.
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</paragraph>
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<paragraph id="8BCB36C6AB1BCC23FF7BFF6DFCF2FE82" blockId="12.[151,1436,152,294]" pageId="12" pageNumber="59">
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It now seems likely, based on size, geographical proximity, and nonsexual characters that “
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<taxonomicName id="4C744D45AB1BCC23FAA6FF6DFA3CFF16" box="[1307,1426,152,178]" class="Diplopoda" genus="Tidesmus" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="59" phylum="Arthropoda" rank="genus">Tidesmus</taxonomicName>
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”
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<taxonomicName id="4C744D45AB1BCC23FF2AFF4AFE16FF7C" authority="Chamberlin 1943" authorityName="Chamberlin" authorityYear="1943" box="[151,440,190,216]" class="Diplopoda" family="Macrosternodesmidae" genus="Nevadesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="59" phylum="Arthropoda" rank="species" species="hubbsi">
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||
<emphasis id="B900EAD4AB1BCC23FF2AFF4AFF4BFF7C" box="[151,229,191,216]" italics="true" pageId="12" pageNumber="59">hubbsi</emphasis>
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||
<bibRefCitation id="EFE54B37AB1BCC23FF51FF4BFE16FF7C" author="Chamberlin" box="[236,440,190,216]" pageId="12" pageNumber="64" refString="Chamberlin, R. V. (1943) On nine North American polydesmoid millipeds. Proceedings of the Biological Society of Washington, 56, 35 - 40." type="journal article" year="1943">Chamberlin 1943</bibRefCitation>
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</taxonomicName>
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(see
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<bibRefCitation id="EFE54B37AB1BCC23FE49FF4BFD58FF7C" author="Shear" box="[500,758,190,216]" pageId="12" pageNumber="65" refString="Shear, W. A., & Shelley, R. M. (2007) The milliped genus Tidesmus Chamberlin 1943 (Polydesmida, Macrosternodesmidae). Zootaxa, 1656, 51 - 68." type="journal article" year="2007">Shear & Shelley 2007</bibRefCitation>
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) is a member of this genus, and it is included here, though males still have not been collected. A female of
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<taxonomicName id="4C744D45AB1BCC23FD57FF10FC95FF5A" box="[746,827,229,254]" class="Diplopoda" family="Macrosternodesmidae" genus="Nevadesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="59" phylum="Arthropoda" rank="species" species="hubbsi">
|
||
<emphasis id="B900EAD4AB1BCC23FD57FF10FC95FF5A" box="[746,827,229,254]" italics="true" pageId="12" pageNumber="59">hubbsi</emphasis>
|
||
</taxonomicName>
|
||
was illustrated by
|
||
<bibRefCitation id="EFE54B37AB1BCC23FB99FF10FA98FF5B" author="Shear" box="[1060,1334,229,255]" pageId="12" pageNumber="65" refString="Shear, W. A., & Shelley, R. M. (2007) The milliped genus Tidesmus Chamberlin 1943 (Polydesmida, Macrosternodesmidae). Zootaxa, 1656, 51 - 68." type="journal article" year="2007">Shear & Shelley 2007</bibRefCitation>
|
||
, and the history and provenance of the species was discussed therein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |