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<document id="9C2DDA320F5D56C94FC65AB6E1D7EF82" ID-CLB-Dataset="46508" ID-DOI="10.11646/zootaxa.3760.4.6" ID-GBIF-Dataset="2b5d31f4-2ab4-4398-a158-a9dc11d678ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="230710" ID-ZooBank="4218FEB8-1E58-4653-924A-E7CC25733AEC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460379911691" checkinUser="plazi" docAuthor="Souto, Paula Malaquias, Da-Silva, Elidiomar Ribeiro &amp; Nessimian, Jorge Luiz" docDate="2014" docId="03DD87C4B84F7523D7F074A9C7E3FB4A" docLanguage="en" docName="zt03760p578.pdf" docOrigin="Zootaxa 3760 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thraulodes pinga Souto, Da-Silva &amp; Nessimian, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="577" masterDocId="FFE4FFBCB84B7525D7677639C27FFF8E" masterDocTitle="Two new species of Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Southeast Brazil" masterLastPageNumber="578" masterPageNumber="571" pageNumber="575" updateTime="1698296676006" updateUser="ExternalLinkService">
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<mods:title id="2D8D0F5B6A7669605326D708B312C429">Two new species of Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Southeast Brazil</mods:title>
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<mods:namePart id="4C99577CE971707FB4BA8D17334B64E6">Souto, Paula Malaquias</mods:namePart>
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<mods:namePart id="F70681B7C873AD21580E8B01A958B13F">Da-Silva, Elidiomar Ribeiro</mods:namePart>
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<mods:namePart id="FC5BE9E6F9438F5F0A4C8F4DEAC921BA">Nessimian, Jorge Luiz</mods:namePart>
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<mods:date id="D00DB0B68270788985A83A4B4C98A895">2014</mods:date>
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<treatment id="03DD87C4B84F7523D7F074A9C7E3FB4A" ID-DOI="http://doi.org/10.5281/zenodo.6495683" ID-GBIF-Taxon="119414033" ID-Zenodo-Dep="6495683" LSID="urn:lsid:plazi:treatment:03DD87C4B84F7523D7F074A9C7E3FB4A" httpUri="http://treatment.plazi.org/id/03DD87C4B84F7523D7F074A9C7E3FB4A" lastPageId="6" lastPageNumber="577" pageId="4" pageNumber="575">
<subSubSection id="C36E6559B84F7521D7F074A9C13CFD24" box="[151,835,656,682]" pageId="4" pageNumber="575" type="nomenclature">
<paragraph id="8BCB36D2B84F7521D7F074A9C13CFD24" blockId="4.[151,835,656,682]" box="[151,835,656,682]" pageId="4" pageNumber="575">
<heading id="D08381BEB84F7521D7F074A9C13CFD24" bold="true" box="[151,835,656,682]" fontSize="11" level="1" pageId="4" pageNumber="575" reason="1">
<emphasis id="B900EAC0B84F7521D7F074A9C13CFD24" bold="true" box="[151,835,656,682]" pageId="4" pageNumber="575">
<taxonomicName id="4C744D51B84F7521D7F074A9C312FD24" box="[151,365,656,682]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="575" phylum="Arthropoda" rank="species" species="pinga" status="sp. nov.">
<emphasis id="B900EAC0B84F7521D7F074A9C312FD24" bold="true" box="[151,365,656,682]" italics="true" pageId="4" pageNumber="575">Thraulodes pinga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357BBB84F7521D61374A9C3ADFD24" box="[372,466,656,682]" pageId="4" pageNumber="575" rank="species">sp. nov.</taxonomicNameLabel>
Souto, Da-Silva &amp; Nessimian
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36E6559B84F7521D7F074EFC617FCF1" pageId="4" pageNumber="575" type="diagnosis">
<paragraph id="8BCB36D2B84F7521D7F074EFC617FCF1" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7F074EFC375FD61" bold="true" box="[151,266,726,751]" pageId="4" pageNumber="575">Diagnosis</emphasis>
.
<emphasis id="B900EAC0B84F7521D67F74EEC320FD60" box="[280,351,727,750]" italics="true" pageId="4" pageNumber="575">Imago</emphasis>
: 1) Forewings with 1 weakly marked cross vein basal to bullae (
<figureCitation id="134F2A57B84F7521D35374EFC6F7FD61" box="[1076,1160,726,751]" captionStart="FIGURES 10 14" captionStartId="5.[151,264,1640,1663]" captionTargetBox="[170,1418,201,1618]" captionTargetId="figure@5.[162,1425,193,1619]" captionTargetPageId="5" captionText="FIGURES 10 14. Thraulodes pinga sp. nov., male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm)." httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="4" pageNumber="575">Fig. 10</figureCitation>
); 2) costal and subcostal areas of forewing translucent; 3) femora of forelegs with one band and one submedian macula (
<figureCitation id="134F2A57B84F7521D27574C3C715FC9D" box="[1298,1386,762,787]" captionStart="FIGURES 10 14" captionStartId="5.[151,264,1640,1663]" captionTargetBox="[170,1418,201,1618]" captionTargetId="figure@5.[162,1425,193,1619]" captionTargetPageId="5" captionText="FIGURES 10 14. Thraulodes pinga sp. nov., male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm)." httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="4" pageNumber="575">Fig. 12</figureCitation>
); 4) abdomen color pattern as in
<figureCitation id="134F2A57B84F7521D6B07527C012FCB9" box="[471,621,798,823]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1984,2007]" captionTargetBox="[359,1222,193,1936]" captionTargetId="figure@2.[359,1224,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 4. Thraulodes spp. nov., male imagos: 1) T. luisae (lateral view); 2) T. luisae (dorsal view); 3) T. pinga (lateral view); 4) T. pinga (dorsal view). Scale: 1.0 mm." httpUri="https://zenodo.org/record/230711/files/figure.png" pageId="4" pageNumber="575">Figs. 3 and 4</figureCitation>
; 5) styliger plate possessing median projection short with wide base and truncated apex (
<figureCitation id="134F2A57B84F7521D632757BC3D2FCD5" box="[341,429,834,859]" captionStart="FIGURES 10 14" captionStartId="5.[151,264,1640,1663]" captionTargetBox="[170,1418,201,1618]" captionTargetId="figure@5.[162,1425,193,1619]" captionTargetPageId="5" captionText="FIGURES 10 14. Thraulodes pinga sp. nov., male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm)." httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="4" pageNumber="575">Fig. 13</figureCitation>
); 6) penes short and wide, with lateral pouch and apicolateral rounded with ear-like projection, penes spines long and narrow, projected internally and posteriorly (
<figureCitation id="134F2A57B84F7521D360755FC625FCF1" box="[1031,1114,870,895]" captionStart="FIGURES 10 14" captionStartId="5.[151,264,1640,1663]" captionTargetBox="[170,1418,201,1618]" captionTargetId="figure@5.[162,1425,193,1619]" captionTargetPageId="5" captionText="FIGURES 10 14. Thraulodes pinga sp. nov., male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm)." httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="4" pageNumber="575">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36E6559B84F7520D7A075B3C3DEF8E3" lastPageId="5" lastPageNumber="576" pageId="4" pageNumber="575" type="description">
<paragraph id="8BCB36D2B84F7521D7A075B3C1B1FC49" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A075B3C39EFC2D" bold="true" box="[199,481,906,931]" italics="true" pageId="4" pageNumber="575">
Male imago (
<typeStatus id="54CF8870B84F7521D60075B3C3AEFC2D" box="[359,465,906,931]" pageId="4" pageNumber="575" type="holotype">Holotype</typeStatus>
):
</emphasis>
Length of body: 6.0 mm; forewings:
<quantity id="4C8C9B37B84F7521D4FB75B3C187FC2C" box="[924,1016,906,931]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="4" pageNumber="575" unit="mm" value="6.6">6.6 mm</quantity>
; hind wings:
<quantity id="4C8C9B37B84F7521D3F175B3C68DFC2C" box="[1174,1266,906,931]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="575" unit="mm" value="1.5">1.5 mm</quantity>
. General color brown with some translucent segments. Wings hyaline with veins yellow.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A075EBC6E8FB81" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A075EBC37BFC65" box="[199,260,978,1003]" italics="true" pageId="4" pageNumber="575">Head</emphasis>
(
<figureCitation id="134F2A57B84F7521D67275EBC3FAFC65" box="[277,389,978,1003]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1984,2007]" captionTargetBox="[359,1222,193,1936]" captionTargetId="figure@2.[359,1224,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 4. Thraulodes spp. nov., male imagos: 1) T. luisae (lateral view); 2) T. luisae (dorsal view); 3) T. pinga (lateral view); 4) T. pinga (dorsal view). Scale: 1.0 mm." httpUri="https://zenodo.org/record/230711/files/figure.png" pageId="4" pageNumber="575">Figs. 3, 4</figureCitation>
): Brown, shaded with dark brown. Scapes light brown; pedicels dark brown; flagella light brown. Eyes black, turbinate portion orange; ocelli white, surrounded by black ring at base.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A07223C019FBD9" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A07223C367FBBD" box="[199,280,1050,1075]" italics="true" pageId="4" pageNumber="575">Thorax</emphasis>
(
<figureCitation id="134F2A57B84F7521D6417223C3F0FBBD" box="[294,399,1050,1075]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1984,2007]" captionTargetBox="[359,1222,193,1936]" captionTargetId="figure@2.[359,1224,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 4. Thraulodes spp. nov., male imagos: 1) T. luisae (lateral view); 2) T. luisae (dorsal view); 3) T. pinga (lateral view); 4) T. pinga (dorsal view). Scale: 1.0 mm." httpUri="https://zenodo.org/record/230711/files/figure.png" pageId="4" pageNumber="575">Figs. 3, 4</figureCitation>
): Nota brown with light brown suture. Prosternum white; mesosternum medially light brown and laterally brown; metasternum brown.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A0725DC0BCFB4D" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A0725DC283FBF5" box="[199,252,1124,1147]" italics="true" pageId="4" pageNumber="575">Legs</emphasis>
(
<figureCitation id="134F2A57B84F7521D66C725BC322FBF5" box="[267,349,1122,1147]" captionStart="FIGURES 10 14" captionStartId="5.[151,264,1640,1663]" captionTargetBox="[170,1418,201,1618]" captionTargetId="figure@5.[162,1425,193,1619]" captionTargetPageId="5" captionText="FIGURES 10 14. Thraulodes pinga sp. nov., male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm)." httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="4" pageNumber="575">Fig. 12</figureCitation>
): Legs I and II with coxae light brown; leg II with coxae dark brown. Femora of all legs with an apical transversal band heavily marked and one submedian maculae. Tibia of all legs yellowish; tibia of foreleg with apical dark mark. Tarsi and claws yellowish.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A072F6C33CFAA1" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A072F6C375FB68" box="[199,266,1231,1254]" italics="true" pageId="4" pageNumber="575">Wings</emphasis>
(
<figureCitation id="134F2A57B84F7521D67972F7C3D6FB69" box="[286,425,1230,1255]" captionStart="FIGURES 10 14" captionStartId="5.[151,264,1640,1663]" captionTargetBox="[170,1418,201,1618]" captionTargetId="figure@5.[162,1425,193,1619]" captionTargetPageId="5" captionText="FIGURES 10 14. Thraulodes pinga sp. nov., male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm)." httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="4" pageNumber="575">Figs. 10, 11</figureCitation>
): Membrane of wings hyaline with dark brown stain at base; veins yellow. Membrane between C and Sc of forewings white on apical third. Forewings with 1 cross vein weakly marked basal and 13 distal to bullae.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A07303C6CDFA15" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A07303C34EFADD" box="[199,305,1338,1363]" italics="true" pageId="4" pageNumber="575">Abdomen</emphasis>
(
<figureCitation id="134F2A57B84F7521D6277303C3D4FADD" box="[320,427,1338,1363]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1984,2007]" captionTargetBox="[359,1222,193,1936]" captionTargetId="figure@2.[359,1224,193,1938]" captionTargetPageId="2" captionText="FIGURES 1 4. Thraulodes spp. nov., male imagos: 1) T. luisae (lateral view); 2) T. luisae (dorsal view); 3) T. pinga (lateral view); 4) T. pinga (dorsal view). Scale: 1.0 mm." httpUri="https://zenodo.org/record/230711/files/figure.png" pageId="4" pageNumber="575">Figs. 3, 4</figureCitation>
): Terga IVI translucent; terga VIIX brown; terga IIVI with a pair of submedian spots, a pair of midway spots and a pair of stigmatic dots; terga VIIX with a dark oblique lateral streak; tergum VII with small dorsal dots. Sterna translucent. Caudal filaments yellowish white with dark annulations.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A0739FC0D1F9C1" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A0739FC34EFA31" box="[199,305,1446,1471]" italics="true" pageId="4" pageNumber="575">Genitalia</emphasis>
(
<figureCitation id="134F2A57B84F7521D658739FC3BAFA31" box="[319,453,1446,1471]" captionStart="FIGURES 10 14" captionStartId="5.[151,264,1640,1663]" captionTargetBox="[170,1418,201,1618]" captionTargetId="figure@5.[162,1425,193,1619]" captionTargetPageId="5" captionText="FIGURES 10 14. Thraulodes pinga sp. nov., male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm)." httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="4" pageNumber="575">Figs. 13, 14</figureCitation>
): Styliger plate wide and light brown, with posterior median projection short, with wide base and truncated apex. Forceps white; segment I with dark stain at apex and with strong constriction near the middle; inner margin of segment I with several short setae basal to constriction. Penes short and wide, with lateral pouch and an apicolateral rounded with ear-like projection; penes spines long and narrow, with 1.1x the width of penes base, projected internally and posteriorly.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A07063C0AEF919" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A07063C3FDF9FD" bold="true" box="[199,386,1626,1651]" italics="true" pageId="4" pageNumber="575">Male subimago:</emphasis>
Similar to imago, however the caudal filaments have two black annulations, one weaker than the other and the general color is paler than imago.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A0709BC38BF951" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A0709BC307F935" bold="true" box="[199,376,1698,1723]" italics="true" pageId="4" pageNumber="575">Female imago:</emphasis>
Length of body:
<quantity id="4C8C9B37B84F7521D521709BC0F2F934" box="[582,653,1698,1723]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="575" unit="mm" value="6.0">6 mm</quantity>
; forewings:
<quantity id="4C8C9B37B84F7521D47A709AC11BF934" box="[797,868,1699,1722]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="575" unit="mm" value="7.0">7 mm</quantity>
; hind wings:
<quantity id="4C8C9B37B84F7521D365709AC636F934" box="[1026,1097,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="575" unit="mm" value="1.0">1 mm</quantity>
. General color brown. Wings hyaline with light brown veins.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A070D3C0BCF8A9" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A070D3C372F88D" box="[199,269,1770,1795]" italics="true" pageId="4" pageNumber="575">Head:</emphasis>
Brown, shaded with dark brown. Eyes black. Ocelli white, surrounding by black at base. Scapes light brown; pedicels dark brown; flagella light brown.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A0710BC071F8E1" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A0710BC35EF8C5" box="[199,289,1842,1867]" italics="true" pageId="4" pageNumber="575">Thorax:</emphasis>
Nota brown, with suture light brown. Prosternum white; median region light brown and lateral regions brown; metasternum light brown.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A07145C66AF81D" blockId="4.[151,1436,726,2011]" box="[199,1045,1914,1939]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A07145C379F81D" box="[199,262,1916,1939]" italics="true" pageId="4" pageNumber="575">Legs:</emphasis>
Similar to male, except forelegs smaller. Tarsi and claws light brown.
</paragraph>
<paragraph id="8BCB36D2B84F7521D7A071A6C173F855" blockId="4.[151,1436,726,2011]" pageId="4" pageNumber="575">
<emphasis id="B900EAC0B84F7521D7A071A6C36CF838" box="[199,275,1951,1974]" italics="true" pageId="4" pageNumber="575">Wings:</emphasis>
Membrane of wings hyaline with dark brown stain at base; veins brown. Presence of 2 cross veins weakly marked basal and 15 cross veins distal to bullae.
</paragraph>
<caption id="DF0B665AB84E7520D7F07051C077F935" httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="5" pageNumber="576" targetBox="[170,1418,201,1618]" targetPageId="5">
<paragraph id="8BCB36D2B84E7520D7F07051C077F935" blockId="5.[151,1436,1640,1723]" pageId="5" pageNumber="576">
<emphasis id="B900EAC0B84E7520D7F07051C32CF9F3" bold="true" box="[151,339,1640,1663]" pageId="5" pageNumber="576">FIGURES 1014.</emphasis>
<taxonomicName id="4C744D51B84E7520D63D7051C071F9F0" box="[346,526,1640,1662]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="pinga" status="sp. nov.">
<emphasis id="B900EAC0B84E7520D63D7051C071F9F0" box="[346,526,1640,1662]" italics="true" pageId="5" pageNumber="576">Thraulodes pinga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357BBB84E7520D5737050C01AF9F0" box="[532,613,1641,1662]" pageId="5" pageNumber="576" rank="species">
<emphasis id="B900EAC0B84E7520D5737050C01AF9F0" bold="true" box="[532,613,1641,1662]" pageId="5" pageNumber="576">sp. nov.</emphasis>
</taxonomicNameLabel>
, male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm).
</paragraph>
</caption>
<paragraph id="8BCB36D2B84E7520D7A070D1C66BF8AB" blockId="5.[151,1436,1768,2009]" pageId="5" pageNumber="576">
<emphasis id="B900EAC0B84E7520D7A070D1C345F88F" box="[199,314,1768,1793]" italics="true" pageId="5" pageNumber="576">Abdomen:</emphasis>
Terga brown with oblique lateral streak. Terga IVI darker than others, with dorsal black mark. Sterna light brown. Sternum X apex with a V-shape cleft. Caudal filaments lost.
</paragraph>
<paragraph id="8BCB36D2B84E7520D7A07109C7EBF8C7" blockId="5.[151,1436,1768,2009]" box="[199,1428,1840,1865]" pageId="5" pageNumber="576">
<emphasis id="B900EAC0B84E7520D7A07109C3E1F8C7" bold="true" box="[199,414,1840,1865]" italics="true" pageId="5" pageNumber="576">Female subimago:</emphasis>
Similar to female imago, but with paler color. Caudal filaments similar to male subimago.
</paragraph>
<paragraph id="8BCB36D2B84E7520D7A0716DC3DEF8E3" blockId="5.[151,1436,1768,2009]" box="[199,417,1876,1901]" pageId="5" pageNumber="576">
<emphasis id="B900EAC0B84E7520D7A0716DC35BF8E3" bold="true" box="[199,292,1876,1901]" italics="true" pageId="5" pageNumber="576">Nymph:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36E6559B84E7520D7A07141C347F857" pageId="5" pageNumber="576" type="biology_ecology">
<paragraph id="8BCB36D2B84E7520D7A07141C347F857" blockId="5.[151,1436,1768,2009]" pageId="5" pageNumber="576">
<emphasis id="B900EAC0B84E7520D7A07141C358F81F" bold="true" box="[199,295,1912,1937]" pageId="5" pageNumber="576">Biology.</emphasis>
Specimens of the new species live in sympatry with specimens of
<taxonomicName id="4C744D51B84E7520D3467141C71AF81F" authority="Goncalves" authorityName="Goncalves" box="[1057,1381,1912,1937]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="jones">
<emphasis id="B900EAC0B84E7520D3467141C69BF81E" box="[1057,1252,1912,1937]" italics="true" pageId="5" pageNumber="576">Thraulodes jones</emphasis>
Gonçalves
</taxonomicName>
, Da- Silva &amp;
<bibRefCitation id="EFE54B23B84E7520D79471A5C3CCF83B" author="Nessimian" box="[243,435,1948,1973]" pageId="5" pageNumber="576" refString="Goncalves, I. C., Da-Silva, E. R. &amp; Nessimian, J. L. (2010) A new species of Thraulodes Ulmer (Ephemeroptera: Leptophlebiidae) from Southeastern Brazil. Zootaxa, 2438, 61 - 68." type="journal article" year="2010">Nessimian, 2010</bibRefCitation>
.
<taxonomicName id="4C744D51B84E7520D6DA71A5C0FCF83A" box="[445,643,1948,1973]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="pinga" status="sp. nov.">
<emphasis id="B900EAC0B84E7520D6DA71A5C0FCF83A" box="[445,643,1948,1973]" italics="true" pageId="5" pageNumber="576">Thraulodes pinga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357BBB84E7520D5EE71A5C09CF83B" box="[649,739,1948,1973]" pageId="5" pageNumber="576" rank="species">
<emphasis id="B900EAC0B84E7520D5EE71A5C09CF83B" bold="true" box="[649,739,1948,1973]" pageId="5" pageNumber="576">sp. nov.</emphasis>
</taxonomicNameLabel>
occurs only at low altitudes, ranging between
<quantity id="4C8C9B37B84E7520D38E71A5C731F83A" box="[1257,1358,1948,1973]" metricMagnitude="1" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="576" unit="m" value="70.0" valueMax="80.0" valueMin="60.0">6080 m</quantity>
, flying after nightfall.
</paragraph>
</subSubSection>
<subSubSection id="C36E6559B84D7523D7A076AEC65BFF5B" pageId="6" pageNumber="577" type="etymology">
<paragraph id="8BCB36D2B84D7523D7A076AEC65BFF5B" blockId="6.[151,1436,151,1005]" pageId="6" pageNumber="577">
<emphasis id="B900EAC0B84D7523D7A076AEC331FF3E" bold="true" box="[199,334,151,176]" pageId="6" pageNumber="577">Etymology.</emphasis>
<taxonomicName id="4C744D51B84D7523D63876AEC3DCFF3E" box="[351,419,151,176]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pinga" status="sp. nov.">Pinga</taxonomicName>
” is an informal name for cachaça, a Brazilian alcoholic beverage derived from sugarcane. Paraty, the type-locality of the
<typeStatus id="54CF8870B84D7523D6977685C02AFF5B" box="[496,597,188,213]" pageId="6" pageNumber="577" type="holotype">holotype</typeStatus>
, is a municipality famous for its cachaça.
</paragraph>
</subSubSection>
<subSubSection id="C36E6559B84D7523D7A076E6C670FE5E" pageId="6" pageNumber="577" type="materials_examined">
<paragraph id="8BCB36D2B84D7523D7A076E6C670FE5E" blockId="6.[151,1436,151,1005]" pageId="6" pageNumber="577">
<emphasis id="B900EAC0B84D7523D7A076E6C307FF76" bold="true" box="[199,376,223,248]" pageId="6" pageNumber="577">
<typeStatus id="54CF8870B84D7523D7A076E6C37DFF76" box="[199,258,223,248]" pageId="6" pageNumber="577">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="F3637642B84D7523D6E576E6C3B6FF76" box="[386,457,223,248]" name="Brazil" pageId="6" pageNumber="577">Brazil</collectingCountry>
, Rio de Janeiro State:
<typeStatus id="54CF8870B84D7523D5B476E6C144FF76" box="[723,827,223,248]" pageId="6" pageNumber="577" type="holotype">
<emphasis id="B900EAC0B84D7523D5B476E6C144FF76" bold="true" box="[723,827,223,248]" italics="true" pageId="6" pageNumber="577">Holotype</emphasis>
</typeStatus>
: Paraty, Sertão do Taquari, Afluente do Rio Taquari (PA2),
<geoCoordinate id="EE405015B84D7523D78D773DC3EDFE93" box="[234,402,260,285]" direction="south" orientation="latitude" pageId="6" pageNumber="577" precision="1" value="-23.041777">23°0230,40S</geoCoordinate>
/
<geoCoordinate id="EE405015B84D7523D6FB773DC02BFE93" box="[412,596,260,285]" direction="west" orientation="longitude" pageId="6" pageNumber="577" precision="1" value="-44.69589">44°4145,20W</geoCoordinate>
,
<date id="FFCA1012B84D7523D504773DC0AEFE92" box="[611,721,260,284]" pageId="6" pageNumber="577" value="2013-01-01">01.I.2013</date>
, Souto, P.M., Hoffmann, A. &amp; Silveira, L.F.
<emphasis id="B900EAC0B84D7523D386773DC774FE93" box="[1249,1291,260,285]" italics="true" pageId="6" pageNumber="577">leg.</emphasis>
male imago (
<collectionCode id="ED65AE17B84D7523D7F97711C29CFECE" box="[158,227,296,320]" pageId="6" pageNumber="577">DZRJ</collectionCode>
2368).
<typeStatus id="54CF8870B84D7523D65E7711C3D5FEB1" box="[313,426,296,319]" pageId="6" pageNumber="577" type="paratype">
<emphasis id="B900EAC0B84D7523D65E7711C3D5FEB1" bold="true" box="[313,426,296,319]" italics="true" pageId="6" pageNumber="577">Paratypes</emphasis>
</typeStatus>
: Same data,
<specimenCount id="9D72FD5BB84D7523D55D7711C0FAFECE" box="[570,645,295,320]" pageId="6" pageNumber="577" type="male">1 male</specimenCount>
imago, pair of wings on slide (
<collectionCode id="ED65AE17B84D7523D4877711C65AFECE" box="[992,1061,296,320]" pageId="6" pageNumber="577">DZRJ</collectionCode>
2369); Same data,
<specimenCount id="9D72FD5BB84D7523D39A7711C737FECE" box="[1277,1352,295,320]" pageId="6" pageNumber="577" type="male">1 male</specimenCount>
imago, genitalia on slide (
<collectionCode id="ED65AE17B84D7523D60E7775C3D1FEEA" box="[361,430,332,356]" pageId="6" pageNumber="577">DZRJ</collectionCode>
2370); Same data,
<specimenCount id="9D72FD5BB84D7523D5EB7775C091FEEB" box="[652,750,332,357]" pageId="6" pageNumber="577" type="female">1 female</specimenCount>
imago (
<collectionCode id="ED65AE17B84D7523D42A7775C1EDFEEA" box="[845,914,332,356]" pageId="6" pageNumber="577">DZRJ</collectionCode>
2371); Same data,
<specimenCount id="9D72FD5BB84D7523D3177775C6ADFEEB" box="[1136,1234,332,357]" pageId="6" pageNumber="577" type="female">1 female</specimenCount>
subimago (
<collectionCode id="ED65AE17B84D7523D2307775C7E3FEEA" box="[1367,1436,332,356]" pageId="6" pageNumber="577">DZRJ</collectionCode>
2372); Same locality,
<date id="FFCA1012B84D7523D6F37749C057FE06" box="[404,552,368,392]" pageId="6" pageNumber="577" value="2012-08-18">18.VIII.2012</date>
, Souto, P.M. &amp; Gonçalves, R.S.
<emphasis id="B900EAC0B84D7523D4C37756C1ACFE06" box="[932,979,367,392]" italics="true" pageId="6" pageNumber="577">leg.,</emphasis>
<specimenCount id="9D72FD5BB84D7523D4BA7749C654FE06" box="[989,1067,367,392]" pageId="6" pageNumber="577" type="male">3 male</specimenCount>
subimagos (
<collectionCode id="ED65AE17B84D7523D3DA7749C77DFE06" box="[1213,1282,368,392]" pageId="6" pageNumber="577">DZRJ</collectionCode>
2373); Same locality,
<date id="FFCA1012B84D7523D79A77ADC309FE22" box="[253,374,404,428]" pageId="6" pageNumber="577" value="2012-10-12">12.X.2012</date>
, Souto, P.M. &amp; Gonçalves, R.S.
<emphasis id="B900EAC0B84D7523D46C77ADC152FE23" box="[779,813,404,429]" italics="true" pageId="6" pageNumber="577">leg</emphasis>
.
<specimenCount id="9D72FD5BB84D7523D42677ADC1CBFE23" box="[833,948,404,429]" pageId="6" pageNumber="577" type="female">2 females</specimenCount>
subimagos (
<collectionCode id="ED65AE17B84D7523D32977ADC6ECFE22" box="[1102,1171,404,428]" pageId="6" pageNumber="577">DZRJ</collectionCode>
2374); Same locality,
<date id="FFCA1012B84D7523D7F07781C370FE5E" box="[151,271,440,464]" pageId="6" pageNumber="577" value="2012-10-12">12.X.2012</date>
, Souto, P.M. &amp; Gonçalves, R.S.
<emphasis id="B900EAC0B84D7523D51A778EC0E0FE5E" box="[637,671,439,464]" italics="true" pageId="6" pageNumber="577">leg</emphasis>
.
<specimenCount id="9D72FD5BB84D7523D5CA7781C086FE5E" box="[685,761,439,464]" pageId="6" pageNumber="577" type="male">1 male</specimenCount>
subimago (
<collectionCode id="ED65AE17B84D7523D41C7781C1BFFE5E" box="[891,960,440,464]" pageId="6" pageNumber="577">DZRJ</collectionCode>
2375).
</paragraph>
</subSubSection>
<subSubSection id="C36E6559B84D7523D7A077E5C7E3FB4A" pageId="6" pageNumber="577" type="discussion">
<paragraph id="8BCB36D2B84D7523D7A077E5C3A3FD43" blockId="6.[151,1436,151,1005]" pageId="6" pageNumber="577">
<emphasis id="B900EAC0B84D7523D7A077E5C333FE7B" bold="true" box="[199,332,476,501]" pageId="6" pageNumber="577">Discussion.</emphasis>
<taxonomicName id="4C744D51B84D7523D63077E5C05CFE7A" box="[343,547,476,501]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pinga" status="sp. nov.">
<emphasis id="B900EAC0B84D7523D63077E5C05CFE7A" box="[343,547,476,501]" italics="true" pageId="6" pageNumber="577">Thraulodes pinga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357BBB84D7523D54977E5C0F3FE7B" box="[558,652,476,501]" pageId="6" pageNumber="577" rank="species">
<emphasis id="B900EAC0B84D7523D54977E5C0F3FE7B" bold="true" box="[558,652,476,501]" pageId="6" pageNumber="577">sp. nov.</emphasis>
</taxonomicNameLabel>
resembles
<taxonomicName id="4C744D51B84D7523D47477E5C675FE7B" box="[787,1034,476,501]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pelicanus">
<emphasis id="B900EAC0B84D7523D47477E5C675FE7B" box="[787,1034,476,501]" italics="true" pageId="6" pageNumber="577">Thraulodes pelicanus</emphasis>
</taxonomicName>
, specially by the abdominal color pattern, short penes with a lateral pouch and apicolateral rounded with ear-like projection. However, the new species can be distinguished from
<taxonomicName id="4C744D51B84D7523D57A741CC0DAFDB3" box="[541,677,548,573]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pelicanus">
<emphasis id="B900EAC0B84D7523D57A741CC0DAFDB3" box="[541,677,548,573]" italics="true" pageId="6" pageNumber="577">T. pelicanus</emphasis>
</taxonomicName>
by the styliger plate format with posterior median projection short with truncate apex and the presence of one cross vein weakly marked basal to bullae.
<taxonomicName id="4C744D51B84D7523D300747EC726FDEE" box="[1127,1369,583,608]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pelicanus">
<emphasis id="B900EAC0B84D7523D300747EC726FDEE" box="[1127,1369,583,608]" italics="true" pageId="6" pageNumber="577">Thraulodes pelicanus</emphasis>
</taxonomicName>
has a styliger plate with posterior median projection long, almost the size of the penes, and cross veins basal to bullae are absent in forewings. Moreover,
<taxonomicName id="4C744D51B84D7523D56D74B6C0ABFD26" box="[522,724,655,680]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pinga" status="sp. nov.">
<emphasis id="B900EAC0B84D7523D56D74B6C0ABFD26" box="[522,724,655,680]" italics="true" pageId="6" pageNumber="577">Thraulodes pinga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357BBB84D7523D5B974B6C144FD26" box="[734,827,655,680]" pageId="6" pageNumber="577" rank="species">
<emphasis id="B900EAC0B84D7523D5B974B6C144FD26" bold="true" box="[734,827,655,680]" pageId="6" pageNumber="577">sp. nov.</emphasis>
</taxonomicNameLabel>
has one band and one maculae on femora, while
<taxonomicName id="4C744D51B84D7523D2E074A8C37CFD43" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pelicanus">
<emphasis id="B900EAC0B84D7523D2E074A8C37CFD43" italics="true" pageId="6" pageNumber="577">T. pelicanus</emphasis>
</taxonomicName>
has only one band.
</paragraph>
<paragraph id="8BCB36D2B84D7523D7A074EEC193FC2A" blockId="6.[151,1436,151,1005]" pageId="6" pageNumber="577">
The two new species described here can be distinguished by the styliger plate and penes format, the number of cross veins basal to bullae on forewings, the general color of the body and the abdominal color pattern. Also, the two new species will key to
<taxonomicName id="4C744D51B84D7523D6807526C108FCB6" authority="Edmunds, 1950" authorityName="Edmunds" authorityYear="1950" box="[487,887,799,824]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B900EAC0B84D7523D6807526C0C5FCB6" box="[487,698,799,824]" italics="true" pageId="6" pageNumber="577">Thraulodes ulmeri</emphasis>
Edmunds,
<quantity id="4C8C9B37B84D7523D4587526C108FCB6" box="[831,887,799,824]" metricMagnitude="1" metricUnit="m" metricValue="4.953" pageId="6" pageNumber="577" unit="in" value="1950.0">1950</quantity>
</taxonomicName>
in
<bibRefCitation id="EFE54B23B84D7523D4C67526C606FCB6" box="[929,1145,799,824]" pageId="6" pageNumber="577" refString="Lima, L. R. C., Mariano, R. &amp; Pinheiro, U. (2013) New species for Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and the first key to adults from Brazil. Zootaxa, 3709 (3), 230 - 242. http: // dx. doi. org / 10.11646 / zootaxa. 3709.3.2" type="journal article">
Lima
<emphasis id="B900EAC0B84D7523D4817518C65DFCB6" box="[998,1058,799,824]" italics="true" pageId="6" pageNumber="577">et al.</emphasis>
(2013)
</bibRefCitation>
. The new species can be distinguished from
<taxonomicName id="4C744D51B84D7523D616757CC3AAFCD3" box="[369,469,836,861]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B900EAC0B84D7523D616757CC3AAFCD3" box="[369,469,836,861]" italics="true" pageId="6" pageNumber="577">T. ulmeri</emphasis>
</taxonomicName>
by the following characters: femora with one band and maculae, being one band in
<taxonomicName id="4C744D51B84D7523D2E0757CC29FFC0E" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B900EAC0B84D7523D2E0757CC29FFC0E" italics="true" pageId="6" pageNumber="577">T. ulmeri</emphasis>
</taxonomicName>
; coastal membrane basal do bullae with 23 cross veins in
<taxonomicName id="4C744D51B84D7523D41E7550C1A8FC0E" box="[889,983,871,896]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="luisae" status="sp. nov.">
<emphasis id="B900EAC0B84D7523D41E7550C1A8FC0E" box="[889,983,871,896]" italics="true" pageId="6" pageNumber="577">T. luisae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357BBB84D7523D4B9755EC647FC0E" box="[990,1080,871,896]" pageId="6" pageNumber="577" rank="species">
<emphasis id="B900EAC0B84D7523D4B9755EC647FC0E" bold="true" box="[990,1080,871,896]" pageId="6" pageNumber="577">sp. nov.</emphasis>
</taxonomicNameLabel>
and 1 cross vein in
<taxonomicName id="4C744D51B84D7523D27F7550C70BFC0E" box="[1304,1396,873,896]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pinga" status="sp. nov.">
<emphasis id="B900EAC0B84D7523D27F7550C70BFC0E" box="[1304,1396,873,896]" italics="true" pageId="6" pageNumber="577">T. pinga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357BBB84D7523D21D755EC2B7FC2B" pageId="6" pageNumber="577" rank="species">
<emphasis id="B900EAC0B84D7523D21D755EC2B7FC2B" bold="true" pageId="6" pageNumber="577">sp. nov.</emphasis>
</taxonomicNameLabel>
, being 4 cross veins in
<taxonomicName id="4C744D51B84D7523D6A875B4C04BFC2B" box="[463,564,908,933]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B900EAC0B84D7523D6A875B4C04BFC2B" box="[463,564,908,933]" italics="true" pageId="6" pageNumber="577">T. ulmeri</emphasis>
</taxonomicName>
; penes with shorter spines in
<taxonomicName id="4C744D51B84D7523D4E675B4C19AFC2B" box="[897,997,908,933]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B900EAC0B84D7523D4E675B4C19AFC2B" box="[897,997,908,933]" italics="true" pageId="6" pageNumber="577">T. ulmeri</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCB36D2B84D7523D7A07596C62FFC63" blockId="6.[151,1436,151,1005]" pageId="6" pageNumber="577">
The key for male imagos of
<taxonomicName id="4C744D51B84D7523D5667596C0FEFC46" box="[513,641,943,968]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="genus">
<emphasis id="B900EAC0B84D7523D5667596C0FEFC46" box="[513,641,943,968]" italics="true" pageId="6" pageNumber="577">Thraulodes</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFE54B23B84D7523D5C37596C111FC46" box="[676,878,943,968]" pageId="6" pageNumber="577" refString="Lima, L. R. C., Mariano, R. &amp; Pinheiro, U. (2013) New species for Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) and the first key to adults from Brazil. Zootaxa, 3709 (3), 230 - 242. http: // dx. doi. org / 10.11646 / zootaxa. 3709.3.2" type="journal article">
Lima
<emphasis id="B900EAC0B84D7523D5827588C162FC46" box="[741,797,943,968]" italics="true" pageId="6" pageNumber="577">et al.</emphasis>
(2013)
</bibRefCitation>
may be emended by modifying the fourth couplet to separate the new species from
<taxonomicName id="4C744D51B84D7523D56D75ECC013FC63" box="[522,620,980,1005]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B900EAC0B84D7523D56D75ECC013FC63" box="[522,620,980,1005]" italics="true" pageId="6" pageNumber="577">T. ulmeri</emphasis>
</taxonomicName>
and adding two more couplets, as follows:
</paragraph>
<paragraph id="8BCB36D2B84D7523D7F07222C7E3FB4A" blockId="6.[151,1436,1051,1220]" pageId="6" pageNumber="577">
4(3). Penes long and slender, without lateral pouch............................................................ 5 4. Penes short to moderate length and width, with lateral pouch................................................ 15 15(4). Femora with one band............................................................
<taxonomicName id="4C744D51B84D7523D3C3726FC7E4FBE5" authority="Edmunds, 1950" authorityName="Edmunds" authorityYear="1950" box="[1188,1435,1109,1131]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="ulmeri">
<emphasis id="B900EAC0B84D7523D3C3726FC680FBE5" box="[1188,1279,1110,1131]" italics="true" pageId="6" pageNumber="577">T. ulmeri</emphasis>
Edmunds, 1950
</taxonomicName>
15. Femora with one band and maculae 6 16 (15). Costal membrane basal to bullae of forewings with 1 cross vein (
<figureCitation id="134F2A57B84D7523D40972A9C1CBFB28" box="[878,948,1168,1190]" captionStart="FIGURES 10 14" captionStartId="5.[151,264,1640,1663]" captionTargetBox="[170,1418,201,1618]" captionTargetId="figure@5.[162,1425,193,1619]" captionTargetPageId="5" captionText="FIGURES 10 14. Thraulodes pinga sp. nov., male imago: 10) Forewing (scale: 1.0 mm); 11) Hind wing (scale: 1.0 mm); 12) Foreleg (scale: 1.0 mm); 13) General aspect of genitalia, ventral view (scale: 0.5 mm); 14) Detail of penes (left: ventral view, right: dorsal view). (Scale: 0.2 mm)." httpUri="https://zenodo.org/record/230713/files/figure.png" pageId="6" pageNumber="577">Fig. 10</figureCitation>
)...................
<taxonomicName id="4C744D51B84D7523D3F872A8C737FB28" box="[1183,1352,1169,1190]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="pinga" status="sp. nov.">
<emphasis id="B900EAC0B84D7523D3F872A8C737FB28" box="[1183,1352,1169,1190]" italics="true" pageId="6" pageNumber="577">Thraulodes pinga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23357BBB84D7523D22872A9C7E3FB28" box="[1359,1436,1168,1190]" pageId="6" pageNumber="577" rank="species">
<emphasis id="B900EAC0B84D7523D22872A9C7E3FB28" bold="true" box="[1359,1436,1168,1190]" pageId="6" pageNumber="577">
sp.
<date id="FFCA1012B84D7523D21572A9C7E3FB28" box="[1394,1436,1168,1190]" pageId="6" pageNumber="577" value="1916-11">nov.</date>
</emphasis>
</taxonomicNameLabel>
16. Costal membrane basal to bullae of forewings with 2 or 3 cross vein (
<figureCitation id="134F2A57B84D7523D4FD7294C1ADFB4D" box="[922,978,1197,1219]" captionStart="FIGURES 5 9" captionStartId="3.[151,264,1477,1500]" captionTargetBox="[163,1424,199,1452]" captionTargetId="figure@3.[154,1432,193,1456]" captionTargetPageId="3" captionText="FIGURES 5 9. Thraulodes luisae sp. nov., male imago: 5) Forewing (scale: 1.0 mm); 6) Hind wing (scale: 1.0 mm); 7) Foreleg (scale: 1.0 mm); 8) General aspect of genitalia, ventral view (scale: 0.5 mm); 9) Detail of penes (left: dorsal view, right: ventral view). (Scale: 0.1 mm)." httpUri="https://zenodo.org/record/230712/files/figure.png" pageId="6" pageNumber="577">Fig. 5</figureCitation>
)...............
<taxonomicName id="4C744D51B84D7523D3FB7297C736FB4D" box="[1180,1353,1198,1219]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="luisae" status="sp. nov.">
<emphasis id="B900EAC0B84D7523D3FB7297C736FB4D" box="[1180,1353,1198,1219]" italics="true" pageId="6" pageNumber="577">Thraulodes luisae</emphasis>
</taxonomicName>
<emphasis id="B900EAC0B84D7523D2287297C7E3FB4A" bold="true" box="[1359,1436,1198,1220]" pageId="6" pageNumber="577">
<taxonomicNameLabel id="A23357BBB84D7523D2287297C7E3FB4A" box="[1359,1436,1198,1220]" pageId="6" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>