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<document id="D29C47F02BA617F727C71741FAF6506E" ID-CLB-Dataset="24929" ID-DOI="10.5281/zenodo.3532762" ID-GBIF-Dataset="db23a775-d25c-4253-abda-b5a007110f6d" ID-ISSN="0374-1036" ID-Zenodo-Dep="3532762" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573206962882" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2013" docId="03A3FB753F5F1E66FE7AFF3144C10FB2" docLanguage="en" docName="ActaEntMusNatPra.53.1.155-176.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Holotrochus mrazi Irmler 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="169" masterDocId="FF9A830D3F511E68FF93FFED46660A2F" masterDocTitle="New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)" masterLastPageNumber="176" masterPageNumber="155" pageNumber="169" updateTime="1698764282444" updateUser="ExternalLinkService">
<mods:mods id="5EE6CFB943609FEEF0826359E1A53298" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="16D0E0373417D43095725B8182B87AE6">
<mods:title id="5312ECAE3707745A2287125C45D37CB3">New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)</mods:title>
</mods:titleInfo>
<mods:name id="06634A44160359DE5E8F52AB45B9B026" type="personal">
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<mods:roleTerm id="B3F84F07DB7DEB5A63271A02DCDD4093">Author</mods:roleTerm>
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<mods:namePart id="C6EC2614944F832679CDB071EA03F9CC">Irmler, Ulrich</mods:namePart>
</mods:name>
<mods:typeOfResource id="7C89C2F3D299B3357AA33F290588ED6A">text</mods:typeOfResource>
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<mods:title id="AD63D4ACFF5625E3957D2D60E840E3B7">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="CDBE18EF7857C412510EE2C3355A7721">
<mods:date id="09F4B73AD88D0E2D3D49515FFC00EBFB">2013</mods:date>
<mods:detail id="5EC46FE1C58F66654DDC1CD0AEBECC23" type="pubDate">
<mods:number id="FB8200EEBAF5872E0ABFA7C7B149C036">2013-07-15</mods:number>
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<mods:number id="8ABA5C48DAE94368DBACE6F56265030D">53</mods:number>
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<mods:number id="A8645C8D0DCF23075847FF58908476FA">1</mods:number>
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<mods:classification id="8F51E38E02F98B32901135088D4ABA2B">journal article</mods:classification>
<mods:identifier id="BC9D0304E82AA835CACF46F2580CB3A0" type="CLB-Dataset">24929</mods:identifier>
<mods:identifier id="1346529747DF81617FB159E7D8A8E64C" type="DOI">10.5281/zenodo.3532762</mods:identifier>
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<mods:identifier id="66D46E56C7A5049D661DCC42B92FD7E5" type="ISSN">0374-1036</mods:identifier>
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<treatment id="03A3FB753F5F1E66FE7AFF3144C10FB2" ID-DOI="http://doi.org/10.5281/zenodo.4427169" ID-GBIF-Taxon="160218240" ID-Zenodo-Dep="4427169" LSID="urn:lsid:plazi:treatment:03A3FB753F5F1E66FE7AFF3144C10FB2" httpUri="http://treatment.plazi.org/id/03A3FB753F5F1E66FE7AFF3144C10FB2" lastPageNumber="169" pageId="14" pageNumber="169">
<subSubSection id="C31019E83F5F1E66FE7AFF3145720ADB" box="[489,788,220,244]" pageId="14" pageNumber="169" type="nomenclature">
<paragraph id="8BB54A633F5F1E66FE7AFF3145720ADB" blockId="14.[489,788,220,269]" box="[489,788,220,244]" pageId="14" pageNumber="169">
<heading id="D0FDFD0F3F5F1E66FE7AFF3145720ADB" bold="true" box="[489,788,220,244]" centered="true" fontSize="10" level="3" pageId="14" pageNumber="169" reason="4">
<emphasis id="B97E96713F5F1E66FE7AFF3145720ADB" bold="true" box="[489,788,220,244]" pageId="14" pageNumber="169">
<taxonomicName id="4C0A31E03F5F1E66FE7AFF3144DE0ADB" ID-CoL="3MBRZ" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[489,696,220,244]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="169" phylum="Arthropoda" rank="species" species="mrazi" status="sp. nov.">
<emphasis id="B97E96713F5F1E66FE7AFF3144DE0ADB" bold="true" box="[489,696,220,244]" italics="true" pageId="14" pageNumber="169">Holotrochus mrazi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5F1E66FD2CFF3145720ADB" box="[703,788,220,244]" pageId="14" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5F1E66FDD4FF1744D00B22" box="[583,694,250,269]" pageId="14" pageNumber="169" type="description">
<paragraph id="8BB54A633F5F1E66FDD4FF1744D00B22" blockId="14.[489,788,220,269]" box="[583,694,250,269]" pageId="14" pageNumber="169">
(
<figureCitation id="133156E63F5F1E66FDDEFF1744E20B22" box="[589,644,250,269]" captionStart="Figs 8–11. 8" captionStartId="9.[113,150,1524,1543]" captionTargetBox="[150,1050,427,1501]" captionTargetId="figure@9.[143,1058,419,1513]" captionTargetPageId="9" captionText="Figs 8–11. 8– Holotrochus yasuniensis sp. nov.; 9 – H. mrazi sp. nov.; 10 – H. paraguensis sp. nov.; 11 – H. hamatus sp. nov. Body parts:A – aedeagus; B – antenna, C – tergite XII, D – last sternite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532768" httpUri="https://zenodo.org/record/3532768/files/figure.png" pageId="14" pageNumber="169">
Figs
<geoCoordinate id="EE3E2CA43F5F1E66FDEBFF1744E20B22" box="[632,644,250,269]" degrees="9, 18" direction="east" orientation="longitude" pageId="14" pageNumber="169" precision="555" value="9.18">9</geoCoordinate>
</figureCitation>
,
<figureCitation id="133156E63F5F1E66FD1EFF1744D60B22" box="[653,688,250,269]" captionStart="Fig" captionStartId="17.[113,144,1541,1560]" captionTargetBox="[121,1067,217,1507]" captionTargetId="figure@17.[113,1086,211,1515]" captionTargetPageId="17" captionText="Fig. 18. Head, pronotum, and elytra of Holotrochus Erichson,1839 showing shape, punctation, and microsculpture.A – H. paraguensis sp. nov.; B – H.tortilis sp. nov.; C – H.hamatus sp. nov.; D – H.yanuniensis sp. nov.; E – H. mrazi sp. nov.; F – H. lobatus sp. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3532776" httpUri="https://zenodo.org/record/3532776/files/figure.png" pageId="14" pageNumber="169">18E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5F1E66FF04FEC944620B43" pageId="14" pageNumber="169" type="materials_examined">
<paragraph id="8BB54A633F5F1E66FF04FEC944190B18" blockId="14.[151,1126,292,365]" box="[151,639,292,311]" pageId="14" pageNumber="169">
<materialsCitation id="3B62403E3F5F1E66FF04FEC9441C0B18" ID-GBIF-Occurrence="2449305668" box="[151,634,292,311]" country="Brazil" location="Province" pageId="14" pageNumber="169" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<emphasis id="B97E96713F5F1E66FF04FEC947760B18" bold="true" box="[151,272,292,311]" pageId="14" pageNumber="169">
<typeStatus id="54B1F4C13F5F1E66FF04FEC946A20B18" box="[151,196,292,311]" pageId="14" pageNumber="169">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D0AF33F5F1E66FE86FEC9472D0B18" box="[277,331,292,311]" name="Brazil" pageId="14" pageNumber="169">Brazil</collectingCountry>
,
<location id="8ED51CB83F5F1E66FEC7FEC947C60B18" LSID="urn:lsid:plazi:treatment:03A3FB753F5F1E66FE7AFF3144C10FB2:8ED51CB83F5F1E66FEC7FEC947C60B18" box="[340,416,292,311]" country="Brazil" name="Province" pageId="14" pageNumber="169" stateProvince="Minas Gerais">Province</location>
<collectingRegion id="49CE84813F5F1E66FE36FEC9447C0B18" box="[421,538,292,311]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="169">Minas Gerais</collectingRegion>
,
<collectingRegion id="49CE84813F5F1E66FDB1FEC9441C0B18" box="[546,634,292,311]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="169">São Paolo</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB54A633F5F1E66FF04FED244620B43" blockId="14.[151,1126,292,365]" pageId="14" pageNumber="169">
<emphasis id="B97E96713F5F1E66FF04FED2477A0B7E" bold="true" box="[151,284,318,338]" pageId="14" pageNumber="169">Type material.</emphasis>
<materialsCitation id="3B62403E3F5F1E66FEB2FED245AE0B7D" ID-GBIF-Occurrence="2449305658" box="[289,968,319,338]" collectingDate="1914-06" collectionCode="NMPC" collectorName="Mraz'" country="Brazil" location="Minas Gerais" pageId="14" pageNumber="169" specimenCount="1" stateProvince="Minas Gerais" typeStatus="holotype">
<typeStatus id="54B1F4C13F5F1E66FEB2FED247120B7F" box="[289,372,319,338]" pageId="14" pageNumber="169" type="holotype">HOLOTYPE</typeStatus>
:, ‘
<collectingCountry id="F31D0AF33F5F1E66FE0EFED247B10B7D" box="[413,471,319,338]" name="Brazil" pageId="14" pageNumber="169">Brazil</collectingCountry>
,
<collectingRegion id="49CE84813F5F1E66FE4CFED244320B7D" box="[479,596,319,338]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="169">Minas Gerais</collectingRegion>
,
<collectingRegion id="49CE84813F5F1E66FDCFFED244D20B7D" box="[604,692,319,338]" country="Brazil" name="Sao Paulo" pageId="14" pageNumber="169">Sao Pãolo</collectingRegion>
,
<date id="FFB46CA33F5F1E66FD2FFED245700B7D" box="[700,790,319,338]" pageId="14" pageNumber="169" value="1914-06">
<collectingDate id="EFF0954B3F5F1E66FD2FFED245700B7D" box="[700,790,319,338]" pageId="14" pageNumber="169" value="1914-06">June 1914</collectingDate>
</date>
, leg.
<collectorName id="26FF2FB53F5F1E66FCD1FED245130B7D" box="[834,885,319,338]" pageId="14" pageNumber="169">Mraz’</collectorName>
(
<collectionCode id="ED1BD2A63F5F1E66FC13FED245A70B7D" box="[896,961,319,338]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="14" pageNumber="169">NMPC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B62403E3F5F1E66FC5CFED244660B43" ID-GBIF-Occurrence="2449305681" collectingDate="1914-06" collectionCode="NMPC, UIC" collectorName="Mraz'" country="Brazil" location="Minas Gerais" pageId="14" pageNumber="169" specimenCount="2" specimenCount-female="2" stateProvince="Minas Gerais" typeStatus="paratype">
<typeStatus id="54B1F4C13F5F1E66FC5CFED242410B7F" box="[975,1063,319,338]" pageId="14" pageNumber="169" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D0C81EA3F5F1E66FBA1FED242070B7D" box="[1074,1121,315,338]" pageId="14" pageNumber="169" type="female">2 ♀♀</specimenCount>
, with same data as holotype (
<collectionCode id="ED1BD2A63F5F1E66FE1AFEB447AF0B43" box="[393,457,345,364]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="14" pageNumber="169">NMPC</collectionCode>
,
<collectionCode id="ED1BD2A63F5F1E66FE41FEB4479C0B43" box="[466,506,345,364]" pageId="14" pageNumber="169">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5F1E66FF04FE6845A30872" pageId="14" pageNumber="169" type="diagnosis">
<paragraph id="8BB54A633F5F1E66FF04FE6845A30872" blockId="14.[151,1127,389,1437]" pageId="14" pageNumber="169">
<emphasis id="B97E96713F5F1E66FF04FE68476A0BB2" bold="true" box="[151,268,389,413]" pageId="14" pageNumber="169">Diagnosis.</emphasis>
The species is closely related to
<taxonomicName id="4C0A31E03F5F1E66FDF6FE6844AE0BB3" authorityName="Bernhauer" authorityYear="1939" box="[613,712,389,412]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="169" phylum="Arthropoda" rank="species" species="vianai">
<emphasis id="B97E96713F5F1E66FDF6FE6844AE0BB3" box="[613,712,389,412]" italics="true" pageId="14" pageNumber="169">H. vianai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A31E03F5F1E66FD69FE6845120BB3" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[762,884,389,412]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="169" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="B97E96713F5F1E66FD69FE6845120BB3" box="[762,884,389,412]" italics="true" pageId="14" pageNumber="169">H. hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5F1E66FCE9FE6845AA0BB3" box="[890,972,389,412]" pageId="14" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
in shape, punctation, and absence of microsculpture of fore-body.All three species are characterised by the specific structure of the aedeagus with a minute tooth on the inner side of the apex and the spiralled apical part of the endophallus.
<taxonomicName id="4C0A31E03F5F1E66FDD0FE08456D0BD3" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[579,779,485,508]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="169" phylum="Arthropoda" rank="species" species="mrazi" status="sp. nov.">
<emphasis id="B97E96713F5F1E66FDD0FE08456D0BD3" box="[579,779,485,508]" italics="true" pageId="14" pageNumber="169">Holotrochus mrazi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5F1E66FC81FE0845030BD3" box="[786,869,485,508]" pageId="14" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
is distinctly longer than the two other species that are only 4.2 and
<quantity id="4CF2E7863F5F1E66FDFFFDE844D80833" box="[620,702,517,541]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="169" unit="mm" value="4.5">4.5 mm</quantity>
long. Without dissecting the aedeagus, a certain identification is not possible. In contrast to
<taxonomicName id="4C0A31E03F5F1E66FD59FDC845230813" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[714,837,549,572]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="169" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="B97E96713F5F1E66FD59FDC845230813" box="[714,837,549,572]" italics="true" pageId="14" pageNumber="169">H. hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5F1E66FCDEFDC845C60813" box="[845,928,549,572]" pageId="14" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C0A31E03F5F1E66FC47FDC8425E0813" authorityName="Bernhauer" authorityYear="1939" box="[980,1080,549,572]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="169" phylum="Arthropoda" rank="species" species="vianai">
<emphasis id="B97E96713F5F1E66FC47FDC8425E0813" box="[980,1080,549,572]" italics="true" pageId="14" pageNumber="169">H. vianai</emphasis>
</taxonomicName>
, the apical spiral part of the endophallus is larger and thicker and has more coils.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5F1E66FF04FD88459C0F13" pageId="14" pageNumber="169" type="description">
<paragraph id="8BB54A633F5F1E66FF04FD88477C08B2" blockId="14.[151,1127,389,1437]" pageId="14" pageNumber="169">
<emphasis id="B97E96713F5F1E66FF04FD8847440852" bold="true" box="[151,290,613,637]" pageId="14" pageNumber="169">Description.</emphasis>
Length:
<quantity id="4CF2E7863F5F1E66FE1AFD8847870853" box="[393,481,613,637]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="14" pageNumber="169" unit="mm" value="4.9">4.9 mm</quantity>
. Colouration black, elytra dark reddish, legs and antennae light brown.
</paragraph>
<paragraph id="8BB54A633F5F1E66FF26FD48456E0912" blockId="14.[151,1127,389,1437]" pageId="14" pageNumber="169">
Head
<quantity id="4CF2E7863F5F1E66FF60FD4847360893" box="[243,336,677,701]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="169" unit="mm" value="0.4">0.40 mm</quantity>
long,
<quantity id="4CF2E7863F5F1E66FE01FD4847890893" box="[402,495,677,701]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="14" pageNumber="169" unit="mm" value="0.84">0.84 mm</quantity>
wide; eyes slightly prominent; 1.5 times as long as temples; shape of fore-head semicircular with slight central emargination; punctation of vertex dense; on average, interstices between punctures smaller than diameter of punctures; between normal punctures with sparse micro-punctation; clypeus with remains of transverse microsculpture; vertex without microsculpture; surface polished and shiny.
</paragraph>
<paragraph id="8BB54A633F5F1E66FF26FCA847AA0992" blockId="14.[151,1127,389,1437]" pageId="14" pageNumber="169">Antennae short and thick; slightly longer than head; second antennomere globular; third antennomere conical and nearly twice as long as second; following antennomeres wider than long; antennomeres 6 to 11 wider than preceding antennomeres; penultimate antennomere nearly twice as long as wide.</paragraph>
<paragraph id="8BB54A633F5F1E66FF26FC28426F0EB2" blockId="14.[151,1127,389,1437]" pageId="14" pageNumber="169">
Pronotum
<quantity id="4CF2E7863F5F1E66FEB5FC2847E309F3" box="[294,389,965,989]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="169" unit="mm" value="0.9">0.90 mm</quantity>
long,
<quantity id="4CF2E7863F5F1E66FE58FC28444C09F3" box="[459,554,965,989]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="14" pageNumber="169" unit="mm" value="1.18">1.18 mm</quantity>
wide; widest in anterior third; anterior angles obtuse and slightly prominent; anterior edge emarginate, but with slight central prominence; sides in posterior half evenly narrowed to posterior angles; posterior angles obtusely rounded; lateral margin continuing to front edge; only central prominence not margined; punctation even, less dense, but as deep as on head; close to posterior edge denser than in anterior half; interstices between punctures in anterior half 2–3 times as wide as diameter of punctures; with indistinct impunctate midline in posterior half; at posterior angles with indistinct depression.
</paragraph>
<paragraph id="8BB54A633F5F1E66FF26FB48423D0EF2" blockId="14.[151,1127,389,1437]" pageId="14" pageNumber="169">
Elytra
<quantity id="4CF2E7863F5F1E66FF68FB4847310E93" box="[251,343,1189,1213]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="14" pageNumber="169" unit="mm" value="1.05">1.05 mm</quantity>
long,
<quantity id="4CF2E7863F5F1E66FE04FB4847950E93" box="[407,499,1189,1213]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="14" pageNumber="169" unit="mm" value="1.16">1.16 mm</quantity>
wide; with deep coriaceous ground sculpture; punctures larger than on pronotum, but hardly visible in deep ground sculpture; scutellum semicircular.
</paragraph>
<paragraph id="8BB54A633F5F1E66FF26FB0845E00ED2" blockId="14.[151,1127,389,1437]" box="[181,902,1253,1277]" pageId="14" pageNumber="169">Abdomen sparsely punctate and with weak net-like microsculpture.</paragraph>
<paragraph id="8BB54A633F5F1E66FF26FAE8459C0F13" blockId="14.[151,1127,389,1437]" pageId="14" pageNumber="169">Aedeagus hook-like at apex; apex strongly sclerotized; shortly obtuse and at inner edge of apex with short minute tooth; endophallus sack-like at base and spiralled at apex.</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5F1E66FF04FAA8468C0F52" pageId="14" pageNumber="169" type="etymology">
<paragraph id="8BB54A633F5F1E66FF04FAA8468C0F52" blockId="14.[151,1127,389,1437]" pageId="14" pageNumber="169">
<emphasis id="B97E96713F5F1E66FF04FAA8477E0F72" bold="true" box="[151,280,1349,1373]" pageId="14" pageNumber="169">Etymology.</emphasis>
The specific name refers to the collector of the species, Jaro Mráz (*1880– †1927).
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5F1E66FF04FA6844C10FB2" box="[151,679,1413,1437]" pageId="14" pageNumber="169" type="distribution">
<paragraph id="8BB54A633F5F1E66FF04FA6844C10FB2" blockId="14.[151,1127,389,1437]" box="[151,679,1413,1437]" pageId="14" pageNumber="169">
<emphasis id="B97E96713F5F1E66FF04FA68474C0FB2" bold="true" box="[151,298,1413,1437]" pageId="14" pageNumber="169">Distribution.</emphasis>
Known only from the
<typeStatus id="54B1F4C13F5F1E66FD8EFA68442D0FB2" box="[541,587,1413,1437]" pageId="14" pageNumber="169">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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