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<document id="643D4CA73B3BEC883921155AD40B31FA" ID-CLB-Dataset="24929" ID-DOI="10.5281/zenodo.3532762" ID-GBIF-Dataset="db23a775-d25c-4253-abda-b5a007110f6d" ID-ISSN="0374-1036" ID-Zenodo-Dep="3532762" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573206962882" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2013" docId="03A3FB753F5E1E7BFE2DFB3144E70BFB" docLanguage="en" docName="ActaEntMusNatPra.53.1.155-176.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Holotrochus agostii Irmler 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="174" masterDocId="FF9A830D3F511E68FF93FFED46660A2F" masterDocTitle="New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)" masterLastPageNumber="176" masterPageNumber="155" pageNumber="170" updateTime="1698764282444" updateUser="ExternalLinkService">
<mods:mods id="9C5A18C2866A8091F39B832C8CAA935B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C1D16E46B01FBB6D740F1F49A9E6496B">
<mods:title id="798894B171AC6080EFF3C34A928D1D73">New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)</mods:title>
</mods:titleInfo>
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<mods:roleTerm id="F05E140289F6258132C97153DFD4CD57">Author</mods:roleTerm>
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<mods:namePart id="015B302FBA40CC9C04236F9093AA6422">Irmler, Ulrich</mods:namePart>
</mods:name>
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<mods:title id="4A40E833486ADDCF6CAE390913BB0BBD">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="054102E12B1D3A19B6461892679930D4">2013</mods:date>
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<mods:number id="2B18E93D004EEE4E6526D8ACB6787CCC">2013-07-15</mods:number>
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<mods:classification id="DEC72FB106B2A457917F29A8E6178312">journal article</mods:classification>
<mods:identifier id="53DC9C95FF066C2E756CEF6A8D93927C" type="CLB-Dataset">24929</mods:identifier>
<mods:identifier id="E21024C91FD9A961303B77532E9A6722" type="DOI">10.5281/zenodo.3532762</mods:identifier>
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<treatment id="03A3FB753F5E1E7BFE2DFB3144E70BFB" ID-DOI="http://doi.org/10.5281/zenodo.4427173" ID-GBIF-Taxon="160218243" ID-Zenodo-Dep="4427173" LSID="urn:lsid:plazi:treatment:03A3FB753F5E1E7BFE2DFB3144E70BFB" httpUri="http://treatment.plazi.org/id/03A3FB753F5E1E7BFE2DFB3144E70BFB" lastPageId="19" lastPageNumber="174" pageId="15" pageNumber="170">
<subSubSection id="C31019E83F5E1E67FE2DFB3144920EDA" box="[446,756,1244,1269]" pageId="15" pageNumber="170" type="nomenclature">
<paragraph id="8BB54A633F5E1E67FE2DFB3144920EDA" blockId="15.[446,756,1244,1294]" box="[446,756,1244,1269]" pageId="15" pageNumber="170">
<heading id="D0FDFD0F3F5E1E67FE2DFB3144920EDA" bold="true" box="[446,756,1244,1269]" centered="true" fontSize="10" level="3" pageId="15" pageNumber="170" reason="4">
<emphasis id="B97E96713F5E1E67FE2DFB3144920EDA" bold="true" box="[446,756,1244,1269]" pageId="15" pageNumber="170">
<taxonomicName id="4C0A31E03F5E1E67FE2DFB3144FE0EDB" ID-CoL="3MBN9" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[446,664,1244,1268]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="agostii" status="sp. nov.">
<emphasis id="B97E96713F5E1E67FE2DFB3144FE0EDB" bold="true" box="[446,664,1244,1268]" italics="true" pageId="15" pageNumber="170">Holotrochus agostii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5E1E67FD0DFB3044920EDA" box="[670,756,1245,1269]" pageId="15" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5E1E67FD89FB1744F10F21" box="[538,663,1274,1294]" pageId="15" pageNumber="170" type="description">
<paragraph id="8BB54A633F5E1E67FD89FB1744F10F21" blockId="15.[446,756,1244,1294]" box="[538,663,1274,1294]" pageId="15" pageNumber="170">
(
<figureCitation id="133156E63F5E1E67FDB3FB1744050F21" box="[544,611,1274,1294]" captionStart="Figs 12–15. 12" captionStartId="11.[113,149,1034,1053]" captionTargetBox="[126,1068,225,1013]" captionTargetId="figure@11.[113,1088,216,1020]" captionTargetPageId="11" captionText="Figs 12–15. 12 – Holotrochus tortilis sp. nov.; 13 – H. lobatus sp. nov.; 14 – H. agostii sp.nov.; 15 – H.pseudoleticiae sp. nov. Body parts:A – aedeagus; B – antenna; C – last sternite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532770" httpUri="https://zenodo.org/record/3532770/files/figure.png" pageId="15" pageNumber="170">Figs 14</figureCitation>
,
<figureCitation id="133156E63F5E1E67FDFFFB1644F70F21" box="[620,657,1275,1294]" captionStart="Fig" captionStartId="18.[860,892,690,709]" captionTargetBox="[154,839,226,868]" captionTargetId="figure@18.[151,839,224,868]" captionTargetPageId="18" captionText="Fig. 19. Head, pronotum, and elytra of Holotrochus Erichson, 1839 showing shape, punctation, and microsculpture. A – H. agostii sp. nov.; B – H. pseudoleticiae sp. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3532780" httpUri="https://zenodo.org/record/3532780/files/figure.png" pageId="15" pageNumber="170">19A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5E1E67FFE2FAC944150F42" pageId="15" pageNumber="170" type="materials_examined">
<paragraph id="8BB54A633F5E1E67FFE2FAC947800F18" blockId="15.[113,1089,1316,1389]" box="[113,486,1316,1335]" pageId="15" pageNumber="170">
<materialsCitation id="3B62403E3F5E1E67FFE2FAC947840F18" ID-GBIF-Occurrence="2449305667" box="[113,482,1316,1335]" country="Brazil" location="Cerro Grande" pageId="15" pageNumber="170" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<emphasis id="B97E96713F5E1E67FFE2FAC9468C0F18" bold="true" box="[113,234,1316,1335]" pageId="15" pageNumber="170">
<typeStatus id="54B1F4C13F5E1E67FFE2FAC946F80F18" box="[113,158,1316,1335]" pageId="15" pageNumber="170">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D0AF33F5E1E67FF7CFAC947430F18" box="[239,293,1316,1335]" name="Brazil" pageId="15" pageNumber="170">Brazil</collectingCountry>
,
<collectingRegion id="49CE84813F5E1E67FEBDFAC947040F18" box="[302,354,1316,1335]" country="Brazil" name="Bahia" pageId="15" pageNumber="170">Bahia</collectingRegion>
,
<location id="8ED51CB83F5E1E67FEF9FAC947840F18" LSID="urn:lsid:plazi:treatment:03A3FB753F5E1E7BFE2DFB3144E70BFB:8ED51CB83F5E1E67FEF9FAC947840F18" box="[362,482,1316,1335]" country="Brazil" name="Cerro Grande" pageId="15" pageNumber="170" stateProvince="Bahia">Cerro Grande</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB54A633F5E1E67FFE2FAD244150F42" blockId="15.[113,1089,1316,1389]" pageId="15" pageNumber="170">
<emphasis id="B97E96713F5E1E67FFE2FAD246910F7D" bold="true" box="[113,247,1343,1362]" pageId="15" pageNumber="170">Type material.</emphasis>
<materialsCitation id="3B62403E3F5E1E67FF6EFAD244150F42" ID-GBIF-Occurrence="2449305661" collectingDate="1995-01-28" collectionCode="AMNH" collectorName="D. Agosti" country="Brazil" county="Urucuca" elevation="120" latitude="-14.4195" location="Atlantic forest" longLatPrecision="13" longitude="-39.057167" municipality="Cerro Grande" pageId="15" pageNumber="170" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
<typeStatus id="54B1F4C13F5E1E67FF6EFAD247360F7F" box="[253,336,1343,1362]" pageId="15" pageNumber="170" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F31D0AF33F5E1E67FEEBFAD247C80F7D" box="[376,430,1343,1362]" name="Brazil" pageId="15" pageNumber="170">Brazil</collectingCountry>
,
<collectingRegion id="49CE84813F5E1E67FE2BFAD2478A0F7D" box="[440,492,1343,1362]" country="Brazil" name="Bahia" pageId="15" pageNumber="170">Bahia</collectingRegion>
,
<collectingCounty id="62D432EF3F5E1E67FE66FAD244260F7D" box="[501,576,1343,1362]" pageId="15" pageNumber="170">Urucuca</collectingCounty>
,
<collectingMunicipality id="6BD1D0193F5E1E67FDDAFAD244D90F7D" box="[585,703,1343,1362]" pageId="15" pageNumber="170">Cerro Grande</collectingMunicipality>
(
<geoCoordinate id="EE3E2CA43F5E1E67FD5FFAD245500F7D" box="[716,822,1343,1362]" degrees="39" direction="west" minutes="03.43" orientation="longitude" pageId="15" pageNumber="170" precision="9" value="-39.057167">39°03.43’W</geoCoordinate>
,
<geoCoordinate id="EE3E2CA43F5E1E67FCACFAD245C40F7D" box="[831,930,1343,1362]" degrees="14" direction="south" minutes="25.17" orientation="latitude" pageId="15" pageNumber="170" precision="9" value="-14.4195">14°25.17’S</geoCoordinate>
), primary
<location id="8ED51CB83F5E1E67FC6FFAD246C20F42" LSID="urn:lsid:plazi:treatment:03A3FB753F5E1E7BFE2DFB3144E70BFB:8ED51CB83F5E1E67FC6FFAD246C20F42" country="Brazil" county="Urucuca" latitude="-14.4195" longLatPrecision="13" longitude="-39.057167" municipality="Cerro Grande" name="Atlantic forest" pageId="15" pageNumber="170" stateProvince="Bahia">Atlantic forest</location>
,
<quantity id="4CF2E7863F5E1E67FF3EFAB746850F42" box="[173,227,1370,1389]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="170" unit="m" value="120.0">
<elevation id="0027AD503F5E1E67FF3EFAB746850F42" box="[173,227,1370,1389]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="170" unit="m" value="120.0">120 m</elevation>
</quantity>
elevation,
<date id="FFB46CA33F5E1E67FED2FAB747FE0F42" box="[321,408,1370,1389]" pageId="15" pageNumber="170" value="1995-01-28">
<collectingDate id="EFF0954B3F5E1E67FED2FAB747FE0F42" box="[321,408,1370,1389]" pageId="15" pageNumber="170" value="1995-01-28">28.1.1995</collectingDate>
</date>
, leg.
<collectorName id="26FF2FB53F5E1E67FE56FAB744710F42" box="[453,535,1370,1389]" pageId="15" pageNumber="170">D. Agosti</collectorName>
(
<collectionCode id="ED1BD2A63F5E1E67FDB0FAB7440F0F42" box="[547,617,1370,1389]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="15" pageNumber="170">AMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5E1E67FFE2FA6844DE0C72" pageId="15" pageNumber="170" type="diagnosis">
<paragraph id="8BB54A633F5E1E67FFE2FA6844DE0C72" blockId="15.[113,1089,1413,1629]" pageId="15" pageNumber="170">
<emphasis id="B97E96713F5E1E67FFE2FA6846800FB2" bold="true" box="[113,230,1413,1437]" pageId="15" pageNumber="170">Diagnosis.</emphasis>
This species certainly belongs to the
<taxonomicName id="4C0A31E03F5E1E67FDE1FA6B456F0FB2" authorityName="Sharp" authorityYear="1876" box="[626,777,1414,1437]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="syntheticus">
<emphasis id="B97E96713F5E1E67FDE1FA6B456F0FB2" box="[626,777,1414,1437]" italics="true" pageId="15" pageNumber="170">H. syntheticus</emphasis>
</taxonomicName>
group concerning its densely pubescent abdomen. It resembles
<taxonomicName id="4C0A31E03F5E1E67FE48FA4B45730F92" authority="Blackwelder, 1943" authorityName="Blackwelder" authorityYear="1943" box="[475,789,1445,1469]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="poundi">
<emphasis id="B97E96713F5E1E67FE48FA4B44200F92" box="[475,582,1446,1469]" italics="true" pageId="15" pageNumber="170">H. poundi</emphasis>
Blackwelder, 1943
</taxonomicName>
due to the shape and sparse punctation of the pronotum. However, the posterior half of the pronotum of
<taxonomicName id="4C0A31E03F5E1E67FC16FA2B45880FF2" authorityName="Blackwelder" authorityYear="1943" box="[901,1006,1478,1501]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="poundi">
<emphasis id="B97E96713F5E1E67FC16FA2B45880FF2" box="[901,1006,1478,1501]" italics="true" pageId="15" pageNumber="170">H. poundi</emphasis>
</taxonomicName>
is more or less parallel, whereas it is slightly narrowed in
<taxonomicName id="4C0A31E03F5E1E67FD16FA0B44890FD2" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[645,751,1510,1533]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="agostii" status="sp. nov.">
<emphasis id="B97E96713F5E1E67FD16FA0B44890FD2" box="[645,751,1510,1533]" italics="true" pageId="15" pageNumber="170">H. agostii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5E1E67FD65FA0B452E0FD2" box="[758,840,1510,1533]" pageId="15" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
Nevertheless, it can be hardly differentiated from
<taxonomicName id="4C0A31E03F5E1E67FE1EF9EB479E0C32" authorityName="Blackwelder" authorityYear="1943" box="[397,504,1542,1565]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="poundi">
<emphasis id="B97E96713F5E1E67FE1EF9EB479E0C32" box="[397,504,1542,1565]" italics="true" pageId="15" pageNumber="170">H. poundi</emphasis>
</taxonomicName>
without dissection of the aedeagus: The apical lobe of the aedeagus in
<taxonomicName id="4C0A31E03F5E1E67FE8CF9CB47EC0C12" authorityName="Blackwelder" authorityYear="1943" box="[287,394,1574,1597]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="poundi">
<emphasis id="B97E96713F5E1E67FE8CF9CB47EC0C12" box="[287,394,1574,1597]" italics="true" pageId="15" pageNumber="170">H. poundi</emphasis>
</taxonomicName>
is more slender and much longer than in
<taxonomicName id="4C0A31E03F5E1E67FCDFF9CB45D00C12" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[844,950,1574,1597]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="agostii" status="sp. nov.">
<emphasis id="B97E96713F5E1E67FCDFF9CB45D00C12" box="[844,950,1574,1597]" italics="true" pageId="15" pageNumber="170">H. agostii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5E1E67FC2EF9CB42770C12" box="[957,1041,1574,1597]" pageId="15" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
The apical lobe of
<taxonomicName id="4C0A31E03F5E1E67FE9AF9AB47150C72" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[265,371,1606,1629]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="170" phylum="Arthropoda" rank="species" species="agostii" status="sp. nov.">
<emphasis id="B97E96713F5E1E67FE9AF9AB47150C72" box="[265,371,1606,1629]" italics="true" pageId="15" pageNumber="170">H. agostii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5E1E67FEEAF9AB47AA0C72" box="[377,460,1606,1629]" pageId="15" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
is thicker and shorter.
</paragraph>
</subSubSection>
<caption id="DF751AEB3F411E78FF04FC7145D909E5" ID-DOI="http://doi.org/10.5281/zenodo.3532772" ID-Zenodo-Dep="3532772" httpUri="https://zenodo.org/record/3532772/files/figure.png" pageId="16" pageNumber="171" startId="16.[151,182,924,943]" targetBox="[187,1095,223,907]" targetPageId="16">
<paragraph id="8BB54A633F411E78FF04FC7145D909E5" blockId="16.[151,1128,924,970]" pageId="16" pageNumber="171">
Fig. 16. Head, pronotum, and elytra of
<taxonomicName id="4C0A31E03F411E78FE79FC7144AD0980" authority="Coiffait, 1978" authorityName="Coiffait" authorityYear="1978" box="[490,715,924,943]" class="Insecta" family="Staphylinidae" genus="Mimogonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B97E96713F411E78FE79FC71442A0981" box="[490,588,924,942]" italics="true" pageId="16" pageNumber="171">Mimogonia</emphasis>
<bibRefCitation id="EF9B37923F411E78FDC0FC7144AD0980" author="COIFFAIT H." box="[595,715,924,943]" journalOrPublisher="Nouvelle Revue d'Entomologie" pageId="16" pageNumber="171" pagination="177 - 179" part="8" refId="ref9957" refString="COIFFAIT H. 1978: Deux genre d'Osoriinae (Col. Staphylinidae) du sud Portugal nouveaux pour la faune europeenne. Nouvelle Revue d'Entomologie 8 (2): 177 - 179." title="Deux genre d'Osoriinae (Col. Staphylinidae) du sud Portugal nouveaux pour la faune europeenne" type="journal article" year="1978">Coiffait, 1978</bibRefCitation>
</taxonomicName>
showing shape, punctation and microsculpture. A –
<taxonomicName id="4C0A31E03F411E78FF28FC5A477E09E6" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[187,280,951,969]" class="Insecta" family="Staphylinidae" genus="Mimogonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="species" species="baloghi" status="sp. nov.">
<emphasis id="B97E96713F411E78FF28FC5A477E09E6" box="[187,280,951,969]" italics="true" pageId="16" pageNumber="171">M. baloghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F411E78FE8DFC5A470509E5" box="[286,355,951,970]" pageId="16" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
; B –
<taxonomicName id="4C0A31E03F411E78FE00FC5A479909E6" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[403,511,951,969]" class="Insecta" family="Staphylinidae" genus="Mimogonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="species" species="marquesi" status="sp. nov.">
<emphasis id="B97E96713F411E78FE00FC5A479909E6" box="[403,511,951,969]" italics="true" pageId="16" pageNumber="171">M. marquesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F411E78FD97FC5A442F09E5" box="[516,585,951,970]" pageId="16" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
; C –
<taxonomicName id="4C0A31E03F411E78FDE7FC5A44AF09E6" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[628,713,951,969]" class="Insecta" family="Staphylinidae" genus="Mimogonia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="species" species="duckei" status="sp. nov.">
<emphasis id="B97E96713F411E78FDE7FC5A44AF09E6" box="[628,713,951,969]" italics="true" pageId="16" pageNumber="171">M. duckei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F411E78FD5DFC5A457709E5" box="[718,785,951,970]" pageId="16" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: 0.2 mm.
</paragraph>
</caption>
<caption id="DF751AEB3F411E78FF04F9E347270C78" ID-DOI="http://doi.org/10.5281/zenodo.3532774" ID-Zenodo-Dep="3532774" httpUri="https://zenodo.org/record/3532774/files/figure.png" pageId="16" pageNumber="171" startId="16.[151,182,1550,1569]" targetBox="[154,1121,1019,1528]" targetPageId="16">
<paragraph id="8BB54A633F411E78FF04F9E347270C78" blockId="16.[151,1128,1550,1623]" pageId="16" pageNumber="171">
Fig. 17. Head, pronotum, and elytra of
<taxonomicName id="4C0A31E03F411E78FE4CF9E344A80C0E" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" box="[479,718,1550,1569]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="genus">
<emphasis id="B97E96713F411E78FE4CF9E3442F0C0F" box="[479,585,1550,1568]" italics="true" pageId="16" pageNumber="171">Holotrochus</emphasis>
<bibRefCitation id="EF9B37923F411E78FDDDF9E344A80C0E" author="ERICHSON G. F." bookContentInfo="954 pp." box="[590,718,1550,1569]" journalOrPublisher="Morin, Berlin" pageId="16" pageNumber="171" refId="ref9991" refString="ERICHSON G. F. 1839: Genera et species Staphylinorum Insectorum Colepterorum familiae. Morin, Berlin, 954 pp." title="Genera et species Staphylinorum Insectorum Colepterorum familiae" type="book" year="1839">Erichson, 1839</bibRefCitation>
</taxonomicName>
showing shape, punctation, and microsculpture. A –
<taxonomicName id="4C0A31E03F411E78FF2EF9C447C70C14" authority="Sharp, 1876" authorityName="Sharp" authorityYear="1876" box="[189,417,1576,1595]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="B97E96713F411E78FF2EF9C447560C14" box="[189,304,1577,1595]" italics="true" pageId="16" pageNumber="171">H. pubescens</emphasis>
Sharp, 1876
</taxonomicName>
; B –
<taxonomicName id="4C0A31E03F411E78FE42F9C444A20C13" authority="Irmler, 1981" authorityName="Irmler" authorityYear="1981" box="[465,708,1576,1596]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="species" species="amazonicus">
<emphasis id="B97E96713F411E78FE42F9C444370C14" box="[465,593,1577,1595]" italics="true" pageId="16" pageNumber="171">H. amazonicus</emphasis>
<bibRefCitation id="EF9B37923F411E78FDC4F9C544A20C13" author="IRMLER U." box="[599,708,1576,1596]" journalOrPublisher="Entomologische Blatter" pageId="16" pageNumber="171" pagination="143 - 152" part="77" refId="ref10117" refString="IRMLER U. 1981 a: Neue Arten der Gattung Mimogonia Coiffait (1978) aus der Neotropis (Coleoptera, Staphylinidae). Entomologische Blatter 77: 143 - 152." title="Neue Arten der Gattung Mimogonia Coiffait (1978) aus der Neotropis (Coleoptera, Staphylinidae)" type="journal article" year="1981">Irmler, 1981</bibRefCitation>
</taxonomicName>
; C–
<taxonomicName id="4C0A31E03F411E78FD7DF9C445C10C13" authority="Irmler, 2001" authorityName="Irmler" authorityYear="2001" box="[750,935,1576,1596]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="species" species="siolii">
<emphasis id="B97E96713F411E78FD7DF9C445620C14" box="[750,772,1577,1595]" italics="true" pageId="16" pageNumber="171">H.</emphasis>
<emphasis id="B97E96713F411E78FC9AF9C445520C14" box="[777,820,1577,1595]" italics="true" pageId="16" pageNumber="171">siolii</emphasis>
<bibRefCitation id="EF9B37923F411E78FCA9F9C545C10C13" author="IRMLER U." box="[826,935,1576,1596]" journalOrPublisher="Amazoniana" pageId="16" pageNumber="171" pagination="349 - 361" part="16" refId="ref10217" refString="IRMLER U. 2001: New Neotropical species of the genera Clavilispinus, Aneucamptus, Thoracophorus, and Holotrochus (Coleoptera: Staphylinidae, Osoriinae). Amazoniana 16: 349 - 361." title="New Neotropical species of the genera Clavilispinus, Aneucamptus, Thoracophorus, and Holotrochus (Coleoptera: Staphylinidae, Osoriinae)" type="journal article" year="2001">Irmler, 2001</bibRefCitation>
</taxonomicName>
; D –
<taxonomicName id="4C0A31E03F411E78FC4BF9C4427B0C14" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[984,1053,1577,1595]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="171" phylum="Arthropoda" rank="species" species="adisi" status="sp. nov.">
<emphasis id="B97E96713F411E78FC4BF9C4427B0C14" box="[984,1053,1577,1595]" italics="true" pageId="16" pageNumber="171">H. adisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F411E78FBB7F9C4420E0C13" box="[1060,1128,1577,1596]" pageId="16" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: 0.2 mm.
</paragraph>
</caption>
<caption id="DF751AEB3F401E79FFE2F9E844490C61" ID-DOI="http://doi.org/10.5281/zenodo.3532776" ID-Zenodo-Dep="3532776" httpUri="https://zenodo.org/record/3532776/files/figure.png" pageId="17" pageNumber="172" startId="17.[113,144,1541,1560]" targetBox="[121,1067,217,1507]" targetPageId="17">
<paragraph id="8BB54A633F401E79FFE2F9E844490C61" blockId="17.[113,1089,1541,1614]" pageId="17" pageNumber="172">
Fig. 18. Head, pronotum, and elytra of
<taxonomicName id="4C0A31E03F401E79FE21F9E844FB0C37" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" box="[434,669,1541,1560]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="B97E96713F401E79FE21F9E8447D0C38" box="[434,539,1541,1559]" italics="true" pageId="17" pageNumber="172">Holotrochus</emphasis>
<bibRefCitation id="EF9B37923F401E79FD8CF9E844FB0C37" author="ERICHSON G. F." bookContentInfo="954 pp." box="[543,669,1541,1560]" journalOrPublisher="Morin, Berlin" pageId="17" pageNumber="172" refId="ref9991" refString="ERICHSON G. F. 1839: Genera et species Staphylinorum Insectorum Colepterorum familiae. Morin, Berlin, 954 pp." title="Genera et species Staphylinorum Insectorum Colepterorum familiae" type="book" year="1839">Erichson,1839</bibRefCitation>
</taxonomicName>
showing shape, punctation, and microsculpture.A –
<taxonomicName id="4C0A31E03F401E79FF17F9CD47630C1D" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[132,261,1568,1586]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="172" phylum="Arthropoda" rank="species" species="paraguensis" status="sp. nov.">
<emphasis id="B97E96713F401E79FF17F9CD47630C1D" box="[132,261,1568,1586]" italics="true" pageId="17" pageNumber="172">H. paraguensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F401E79FE9AF9CD472B0C1C" box="[265,333,1568,1587]" pageId="17" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
; B –
<taxonomicName id="4C0A31E03F401E79FEE6F9CD47A00C1D" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[373,454,1568,1586]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="172" phylum="Arthropoda" rank="species" species="tortilis" status="sp. nov.">
<emphasis id="B97E96713F401E79FEE6F9CD47A00C1D" box="[373,454,1568,1586]" italics="true" pageId="17" pageNumber="172">H.tortilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F401E79FE59F9CD44680C1C" box="[458,526,1568,1587]" pageId="17" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
; C –
<taxonomicName id="4C0A31E03F401E79FDA5F9CD44F00C1D" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[566,662,1568,1586]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="172" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="B97E96713F401E79FDA5F9CD44F00C1D" box="[566,662,1568,1586]" italics="true" pageId="17" pageNumber="172">H.hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F401E79FD08F9CD44B90C1C" box="[667,735,1568,1587]" pageId="17" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
; D –
<taxonomicName id="4C0A31E03F401E79FC98F9CD45EF0C1D" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[779,905,1568,1586]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="172" phylum="Arthropoda" rank="species" species="yanuniensis" status="sp. nov.">
<emphasis id="B97E96713F401E79FC98F9CD45EF0C1D" box="[779,905,1568,1586]" italics="true" pageId="17" pageNumber="172">H.yanuniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F401E79FC1EF9CD45B70C1C" box="[909,977,1568,1587]" pageId="17" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
; E –
<taxonomicName id="4C0A31E03F401E79FC64F9CD42270C1D" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[1015,1089,1568,1586]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="172" phylum="Arthropoda" rank="species" species="mrazi" status="sp. nov.">
<emphasis id="B97E96713F401E79FC64F9CD42270C1D" box="[1015,1089,1568,1586]" italics="true" pageId="17" pageNumber="172">H. mrazi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F401E79FFE2F9D646D10C61" box="[113,183,1595,1614]" pageId="17" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
; F –
<taxonomicName id="4C0A31E03F401E79FF4CF9D6475E0C62" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[223,312,1595,1613]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="172" phylum="Arthropoda" rank="species" species="lobatus" status="sp. nov.">
<emphasis id="B97E96713F401E79FF4CF9D6475E0C62" box="[223,312,1595,1613]" italics="true" pageId="17" pageNumber="172">H. lobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F401E79FEADF9D647E70C61" box="[318,385,1595,1614]" pageId="17" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="DF751AEB3F431E7AFCCFFD5F42600949" ID-DOI="http://doi.org/10.5281/zenodo.3532780" ID-Zenodo-Dep="3532780" httpUri="https://zenodo.org/record/3532780/files/figure.png" pageId="18" pageNumber="173" startId="18.[860,892,690,709]" targetBox="[154,839,226,868]" targetPageId="18">
<paragraph id="8BB54A633F431E7AFCCFFD5F42600949" blockId="18.[860,1128,690,870]" pageId="18" pageNumber="173">
Fig. 19. Head, pronotum, and elytra of
<taxonomicName id="4C0A31E03F431E7AFC26FD2045A708D4" authority="Erichson, 1839" authorityName="Erichson" authorityYear="1839" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="173" phylum="Arthropoda" rank="genus">
<emphasis id="B97E96713F431E7AFC26FD20424008F0" box="[949,1062,717,735]" italics="true" pageId="18" pageNumber="173">Holotrochus</emphasis>
<bibRefCitation id="EF9B37923F431E7AFBBDFD2045A708D4" author="ERICHSON G. F." bookContentInfo="954 pp." journalOrPublisher="Morin, Berlin" pageId="18" pageNumber="173" refId="ref9991" refString="ERICHSON G. F. 1839: Genera et species Staphylinorum Insectorum Colepterorum familiae. Morin, Berlin, 954 pp." title="Genera et species Staphylinorum Insectorum Colepterorum familiae" type="book" year="1839">Erichson, 1839</bibRefCitation>
</taxonomicName>
showing shape, punctation, and microsculpture. A –
<taxonomicName id="4C0A31E03F431E7AFC25FCF042770900" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[950,1041,797,815]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="173" phylum="Arthropoda" rank="species" species="agostii" status="sp. nov.">
<emphasis id="B97E96713F431E7AFC25FCF042770900" box="[950,1041,797,815]" italics="true" pageId="18" pageNumber="173">H. agostii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F431E7AFB8AFCF04202091F" box="[1049,1124,797,816]" pageId="18" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
; B –
<taxonomicName id="4C0A31E03F431E7AFC17FCD5427D0965" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[900,1051,824,842]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="173" phylum="Arthropoda" rank="species" species="pseudoleticiae" status="sp. nov.">
<emphasis id="B97E96713F431E7AFC17FCD5427D0965" box="[900,1051,824,842]" italics="true" pageId="18" pageNumber="173">H. pseudoleticiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F431E7AFBB0FCD5420E0964" box="[1059,1128,824,843]" pageId="18" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
Scale bars: 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C31019E83F431E7BFF04FC4D47EC0BBB" lastPageId="19" lastPageNumber="174" pageId="18" pageNumber="173" type="description">
<paragraph id="8BB54A633F431E7AFF04FC4D473809D7" blockId="18.[151,1127,928,1592]" pageId="18" pageNumber="173">
<emphasis id="B97E96713F431E7AFF04FC4D47440997" bold="true" box="[151,290,928,952]" pageId="18" pageNumber="173">Description.</emphasis>
Length:
<quantity id="4CF2E7863F431E7AFE11FC4D47B10998" box="[386,471,928,952]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="18" pageNumber="173" unit="mm" value="3.8">3.8 mm</quantity>
. Colouration dark brown; posterior margin of abdominal segments lighter brown; posterior margin and posterior angles of pronotum reddish; legs and antennae light red.
</paragraph>
<paragraph id="8BB54A633F431E7AFF26FBED469F0EB7" blockId="18.[151,1127,928,1592]" pageId="18" pageNumber="173">
Head
<quantity id="4CF2E7863F431E7AFF67FBED47350E38" box="[244,339,1024,1048]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="18" pageNumber="173" unit="mm" value="0.45">0.45 mm</quantity>
long,
<quantity id="4CF2E7863F431E7AFE05FBED47930E38" box="[406,501,1024,1048]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="18" pageNumber="173" unit="mm" value="0.69">0.69 mm</quantity>
wide; eyes large; distinctly longer than temples; fore-head semicircular; punctation fine and sparse; interstices nearly three times as wide as diameter of punctures; two setiferous punctures on clypeus; four setiferous punctures on vertex placed in quadrate and two supraocular setiferous punctures; without microsculpture; surface polished.
</paragraph>
<paragraph id="8BB54A633F431E7AFF26FB4D47D10F17" blockId="18.[151,1127,928,1592]" pageId="18" pageNumber="173">Antennae slightly longer than head and half of pronotum combined; second antennomere nearly globular; conical third antennomere 1.5 times longer than second; following antennomeres distinctly increasing in width; tenth antennomere twice as wide as fourth; fourth antennomere quadrate; tenth antennomere 1.5 times as wide as long; antennomeres six to eleven with dense setation.</paragraph>
<paragraph id="8BB54A633F431E7AFF26FAAD47130C17" blockId="18.[151,1127,928,1592]" pageId="18" pageNumber="173">
Pronotum
<quantity id="4CF2E7863F431E7AFEB5FAAD47E20F78" box="[294,388,1344,1368]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="18" pageNumber="173" unit="mm" value="0.69">0.69 mm</quantity>
long,
<quantity id="4CF2E7863F431E7AFE5AFAAD444E0F78" box="[457,552,1344,1368]" metricMagnitude="-4" metricUnit="m" metricValue="9.7" pageId="18" pageNumber="173" unit="mm" value="0.97">0.97 mm</quantity>
wide; widest in middle; narrowed in smooth curve to anterior and posterior angles; lateral margin thick; combined with anterior margin; posterior edge not margined; margin thickest at anterior angles; punctation sparse and fine; interstices between punctures between three to four times as wide as diameter of punctures; row of setiferous punctures at anterior margin and few setiferous punctures in lateral margin; two setiferous punctures on each side at posterior edge; without microsculpture, but partly with very weak coriaceous ground sculpture; between normal punctation with sparse micro-punctation; surface shiny.
</paragraph>
<paragraph id="8BB54A633F421E7BFF03FF3142540B1B" blockId="19.[113,1089,220,469]" pageId="19" pageNumber="174">
Elytra
<quantity id="4CF2E7863F421E7BFF45FF3047550ADB" box="[214,307,221,245]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="19" pageNumber="174" unit="mm" value="0.83">0.83 mm</quantity>
long,
<quantity id="4CF2E7863F421E7BFEE7FF3047B60ADB" box="[372,464,221,245]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" pageId="19" pageNumber="174" unit="mm" value="0.99">0.99 mm</quantity>
wide; with very weak punctation; nearly invisible in coarse coriaceous ground sculpture; lateral margin wide; numerous setiferous punctures in lateral margin; on anterior disc, two setiferous punctures on each side placed in longitudinal row.
</paragraph>
<paragraph id="8BB54A633F421E7BFF03FED147EC0BBB" blockId="19.[113,1089,220,469]" pageId="19" pageNumber="174">Aedeagus with short apical lobe; apical lobe thick and ending in hook-like apex; endophallus slightly curved in basal part; apical part with two torsions; paramere weakly sinuate and as long as apical lobe.</paragraph>
</subSubSection>
<subSubSection id="C31019E83F421E7BFFE2FE70453D0B9B" box="[113,859,412,437]" pageId="19" pageNumber="174" type="etymology">
<paragraph id="8BB54A633F421E7BFFE2FE70453D0B9B" blockId="19.[113,1089,220,469]" box="[113,859,412,437]" pageId="19" pageNumber="174">
<emphasis id="B97E96713F421E7BFFE2FE7046940B9A" bold="true" box="[113,242,413,437]" pageId="19" pageNumber="174">Etymology.</emphasis>
The specific name is derived from its collector D. Agosti.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F421E7BFFE2FE5144E70BFB" box="[113,641,444,469]" pageId="19" pageNumber="174" type="distribution">
<paragraph id="8BB54A633F421E7BFFE2FE5144E70BFB" blockId="19.[113,1089,220,469]" box="[113,641,444,469]" pageId="19" pageNumber="174">
<emphasis id="B97E96713F421E7BFFE2FE5147620BFB" bold="true" box="[113,260,444,468]" pageId="19" pageNumber="174">Distribution.</emphasis>
Known only from the
<typeStatus id="54B1F4C13F421E7BFE64FE5044430BFA" box="[503,549,445,469]" pageId="19" pageNumber="174">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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