303 lines
36 KiB
XML
303 lines
36 KiB
XML
<document id="5B5CD6E8D6205E7F333FA8E07A3A1AC3" ID-CLB-Dataset="24929" ID-DOI="10.5281/zenodo.3532762" ID-GBIF-Dataset="db23a775-d25c-4253-abda-b5a007110f6d" ID-ISSN="0374-1036" ID-Zenodo-Dep="3532762" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573206962882" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2013" docId="03A3FB753F591E62FE55FF3142720B5B" docLanguage="en" docName="ActaEntMusNatPra.53.1.155-176.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Holotrochus paraguensis Irmler 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="165" masterDocId="FF9A830D3F511E68FF93FFED46660A2F" masterDocTitle="New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)" masterLastPageNumber="176" masterPageNumber="155" pageNumber="163" updateTime="1698764282444" updateUser="ExternalLinkService">
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<mods:mods id="F30DFD80437794C6596D6CC02605D2CA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CAFB6A603B6FE8BF932C697330F5A5B7">
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<mods:title id="E5621F8385489EF39CE0412C6E3BC6E4">New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="9DC34116AE21BADE0E6CEED03BD31424" type="personal">
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<mods:roleTerm id="51FFF00839526C67E63F732D460DEEE8">Author</mods:roleTerm>
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<mods:namePart id="4B9706AE57D0A75475D1A72ED86F561F">Irmler, Ulrich</mods:namePart>
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<mods:typeOfResource id="7AB76D43619C3588683722A550AFA607">text</mods:typeOfResource>
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<mods:title id="C3F4783BC91408BB95FE1CE75F88B66D">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="463A266EFE5455BB71B6E803CE856862">
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<mods:date id="7EA2B80C865F422EDBEFB252339589BC">2013</mods:date>
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<mods:number id="A97DDD6AA05CC5190FDED094EF6E5C87">2013-07-15</mods:number>
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<mods:number id="5C2197538BAE93EA7BEBC3ED2F2295C2">53</mods:number>
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<mods:classification id="6CCA9CDE58EC1B51F28FB2E34A96999B">journal article</mods:classification>
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<mods:identifier id="A7022AF4F542E5AC3FE60F70AFA04593" type="CLB-Dataset">24929</mods:identifier>
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<mods:identifier id="DFFA02DB2B6E34ABD524EA2650DB3F4B" type="DOI">10.5281/zenodo.3532762</mods:identifier>
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<mods:identifier id="4A43F9CC2258C8C0A035FA7A6BA2B9E4" type="ISSN">0374-1036</mods:identifier>
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<treatment id="03A3FB753F591E62FE55FF3142720B5B" ID-DOI="http://doi.org/10.5281/zenodo.4427161" ID-GBIF-Taxon="160218246" ID-Zenodo-Dep="4427161" LSID="urn:lsid:plazi:treatment:03A3FB753F591E62FE55FF3142720B5B" httpUri="http://treatment.plazi.org/id/03A3FB753F591E62FE55FF3142720B5B" lastPageId="10" lastPageNumber="165" pageId="8" pageNumber="163">
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<subSubSection id="C31019E83F591E60FE55FF31455E0ADB" box="[454,824,220,244]" pageId="8" pageNumber="163" type="nomenclature">
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<paragraph id="8BB54A633F591E60FE55FF31455E0ADB" blockId="8.[454,824,220,269]" box="[454,824,220,244]" pageId="8" pageNumber="163">
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<heading id="D0FDFD0F3F591E60FE55FF31455E0ADB" bold="true" box="[454,824,220,244]" centered="true" fontSize="10" level="3" pageId="8" pageNumber="163" reason="4">
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<emphasis id="B97E96713F591E60FE55FF31455E0ADB" bold="true" box="[454,824,220,244]" pageId="8" pageNumber="163">
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<taxonomicName id="4C0A31E03F591E60FE55FF3144BD0ADB" ID-CoL="3MBSN" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[454,731,220,244]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="paraguensis" status="sp. nov.">
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<emphasis id="B97E96713F591E60FE55FF3144BD0ADB" bold="true" box="[454,731,220,244]" italics="true" pageId="8" pageNumber="163">Holotrochus paraguensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24D2B0A3F591E60FD71FF31455E0ADB" box="[738,824,220,244]" pageId="8" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31019E83F591E60FDD3FF1744010B18" pageId="8" pageNumber="163" type="description">
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<paragraph id="8BB54A633F591E60FDD3FF1744DB0B22" blockId="8.[454,824,220,269]" box="[576,701,250,269]" pageId="8" pageNumber="163">
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(
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<figureCitation id="133156E63F591E60FDD5FF1744EE0B22" box="[582,648,250,269]" captionStart="Figs 8–11. 8" captionStartId="9.[113,150,1524,1543]" captionTargetBox="[150,1050,427,1501]" captionTargetId="figure@9.[143,1058,419,1513]" captionTargetPageId="9" captionText="Figs 8–11. 8– Holotrochus yasuniensis sp. nov.; 9 – H. mrazi sp. nov.; 10 – H. paraguensis sp. nov.; 11 – H. hamatus sp. nov. Body parts:A – aedeagus; B – antenna, C – tergite XII, D – last sternite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532768" httpUri="https://zenodo.org/record/3532768/files/figure.png" pageId="8" pageNumber="163">Figs 10</figureCitation>
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,
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<figureCitation id="133156E63F591E60FD02FF1744D10B22" box="[657,695,250,269]" captionStart="Fig" captionStartId="17.[113,144,1541,1560]" captionTargetBox="[121,1067,217,1507]" captionTargetId="figure@17.[113,1086,211,1515]" captionTargetPageId="17" captionText="Fig. 18. Head, pronotum, and elytra of Holotrochus Erichson,1839 showing shape, punctation, and microsculpture.A – H. paraguensis sp. nov.; B – H.tortilis sp. nov.; C – H.hamatus sp. nov.; D – H.yanuniensis sp. nov.; E – H. mrazi sp. nov.; F – H. lobatus sp. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3532776" httpUri="https://zenodo.org/record/3532776/files/figure.png" pageId="8" pageNumber="163">18A</figureCitation>
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)
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</paragraph>
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||
<paragraph id="8BB54A633F591E60FF04FEC944010B18" blockId="8.[151,1128,292,445]" box="[151,615,292,311]" pageId="8" pageNumber="163">
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<materialsCitation id="3B62403E3F591E60FF04FEC944020B18" ID-GBIF-Occurrence="2449305649" box="[151,612,292,311]" country="Paraguay" location="Puerto Presidente Stroessner" pageId="8" pageNumber="163" specimenCount="1" typeStatus="holotype">
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<emphasis id="B97E96713F591E60FF04FEC947760B18" bold="true" box="[151,272,292,311]" pageId="8" pageNumber="163">
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<typeStatus id="54B1F4C13F591E60FF04FEC946A20B18" box="[151,196,292,311]" pageId="8" pageNumber="163">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F31D0AF33F591E60FE86FEC947030B18" box="[277,357,292,311]" name="Paraguay" pageId="8" pageNumber="163">Paraguay</collectingCountry>
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||
,
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<location id="8ED51CB83F591E60FEFDFEC944020B18" LSID="urn:lsid:plazi:treatment:03A3FB753F591E62FE55FF3142720B5B:8ED51CB83F591E60FEFDFEC944020B18" box="[366,612,292,311]" country="Paraguay" name="Puerto Presidente Stroessner" pageId="8" pageNumber="163">Puerto Presidente Stroessner</location>
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31019E83F591E60FF04FED245B60B93" pageId="8" pageNumber="163" type="materials_examined">
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<paragraph id="8BB54A633F591E60FF04FED245B60B93" blockId="8.[151,1128,292,445]" pageId="8" pageNumber="163">
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||
<emphasis id="B97E96713F591E60FF04FED2477A0B7E" bold="true" box="[151,284,318,338]" pageId="8" pageNumber="163">Type material.</emphasis>
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||
<materialsCitation id="3B62403E3F591E60FEB2FED244F30B43" ID-GBIF-Occurrence="2449305673" collectingDate="1966-02-01" collectionCode="HNHM" collectorName="I. Loksa'" country="Paraguay" latitude="-25.516666" location="Puerto Presidente Stroessner" longLatPrecision="1246" longitude="-54.6" municipality="Hungarian" pageId="8" pageNumber="163" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="54B1F4C13F591E60FEB2FED247120B7F" box="[289,372,319,338]" pageId="8" pageNumber="163" type="holotype">HOLOTYPE</typeStatus>
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||
:,
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<emphasis id="B97E96713F591E60FE09FED347C70B7E" bold="true" box="[410,417,318,337]" pageId="8" pageNumber="163">‘</emphasis>
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<collectingCountry id="F31D0AF33F591E60FE32FED247970B7D" box="[417,497,319,338]" name="Paraguay" pageId="8" pageNumber="163">Paraguay</collectingCountry>
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,
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||
<location id="8ED51CB83F591E60FE69FED244890B7D" LSID="urn:lsid:plazi:treatment:03A3FB753F591E62FE55FF3142720B5B:8ED51CB83F591E60FE69FED244890B7D" box="[506,751,319,338]" country="Paraguay" latitude="-25.516666" longLatPrecision="1246" longitude="-54.6" municipality="Hungarian" name="Puerto Presidente Stroessner" pageId="8" pageNumber="163">Puerto Presidente Stroessner</location>
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,
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||
<collectingMunicipality id="6BD1D0193F591E60FD6BFED245340B7D" box="[760,850,319,338]" pageId="8" pageNumber="163">Hungarian</collectingMunicipality>
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(
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<geoCoordinate id="EE3E2CA43F591E60FCCDFED345D60B7D" box="[862,944,318,338]" degrees="54" direction="west" minutes="36" orientation="longitude" pageId="8" pageNumber="163" precision="925" value="-54.6">54°36’ W</geoCoordinate>
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,
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||
<geoCoordinate id="EE3E2CA43F591E60FC2AFED342650B7D" box="[953,1027,318,338]" degrees="25" direction="south" minutes="31" orientation="latitude" pageId="8" pageNumber="163" precision="925" value="-25.516666">25°31’ S</geoCoordinate>
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||
) Soil-Zool. Exp., Acaray waterfall,
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||
<date id="FFB46CA33F591E60FEF1FEB447A30B43" box="[354,453,345,364]" pageId="8" pageNumber="163" value="1966-02-01">
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||
<collectingDate id="EFF0954B3F591E60FEF1FEB447A30B43" box="[354,453,345,364]" pageId="8" pageNumber="163" value="1966-02-01">02.01.1966</collectingDate>
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||
</date>
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||
, leg.
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||
<collectorName id="26FF2FB53F591E60FE61FEB744580B43" box="[498,574,345,365]" pageId="8" pageNumber="163">I. Loksa’</collectorName>
|
||
(
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||
<collectionCode id="ED1BD2A63F591E60FDDAFEB444E90B43" box="[585,655,345,364]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="163">HNHM</collectionCode>
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||
)
|
||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B62403E3F591E60FD0EFEB7477E0BA8" ID-GBIF-Occurrence="2449305656" collectingDate="1966-02-01" collectionCode="HNHM, UIC" collectorName="I. Loksa'" country="Paraguay" latitude="-25.516666" location="Puerto Presidente Stroessner" longLatPrecision="1246" longitude="-54.6" municipality="Hungarian" pageId="8" pageNumber="163" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<typeStatus id="54B1F4C13F591E60FD0EFEB744930B45" box="[669,757,346,365]" pageId="8" pageNumber="163" type="paratype">PARATYPES</typeStatus>
|
||
: 1
|
||
<specimenCount id="9D0C81EA3F591E60FCB6FEB445320B43" box="[805,852,341,364]" pageId="8" pageNumber="163" type="female">2 ♀♀</specimenCount>
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||
, with the same data as holotype (
|
||
<collectionCode id="ED1BD2A63F591E60FF0EFE9946840BA8" box="[157,226,372,391]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="163">HNHM</collectionCode>
|
||
,
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||
<collectionCode id="ED1BD2A63F591E60FF79FE9947740BA8" box="[234,274,372,391]" pageId="8" pageNumber="163">UIC</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="3B62403E3F591E60FEB2FE9942320BA8" ID-GBIF-Occurrence="2449305646" box="[289,1108,368,391]" collectingDate="1965-12-26" collectionCode="HNHM" collectorName="I. Loksa" country="Paraguay" latitude="-25.516666" location="Acaray" longLatPrecision="1246" longitude="-54.6" municipality="Hungarian" pageId="8" pageNumber="163" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<specimenCount id="9D0C81EA3F591E60FEB2FE9947290BA8" box="[289,335,368,391]" pageId="8" pageNumber="163" type="female">2 ♀♀</specimenCount>
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||
from the same location, but not from
|
||
<location id="8ED51CB83F591E60FD03FE9944A80BA8" LSID="urn:lsid:plazi:treatment:03A3FB753F591E62FE55FF3142720B5B:8ED51CB83F591E60FD03FE9944A80BA8" box="[656,718,372,391]" name="Acaray" pageId="8" pageNumber="163">Acaray</location>
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||
waterfall,
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||
<date id="FFB46CA33F591E60FCBBFE9945EC0BA8" box="[808,906,372,391]" pageId="8" pageNumber="163" value="1965-12-26">
|
||
<collectingDate id="EFF0954B3F591E60FCBBFE9945EC0BA8" box="[808,906,372,391]" pageId="8" pageNumber="163" value="1965-12-26">26.12.1965</collectingDate>
|
||
</date>
|
||
, leg,
|
||
<collectorName id="26FF2FB53F591E60FC24FE99459A0BA8" box="[951,1020,372,391]" pageId="8" pageNumber="163">I. Loksa</collectorName>
|
||
(
|
||
<collectionCode id="ED1BD2A63F591E60FB94FE99422B0BA8" box="[1031,1101,372,391]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="163">HNHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B62403E3F591E60FBCFFE9945670B8D" ID-GBIF-Occurrence="2449305670" collectingDate="1965-12-30" collectionCode="HNHM" collectorName="L. S. Mahunka" country="Paraguay" latitude="-25.516666" location="Acaray" longLatPrecision="1246" longitude="-54.6" municipality="Hungarian" pageId="8" pageNumber="163" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D0C81EA3F591E60FBCFFE9946C10B8D" pageId="8" pageNumber="163" type="female">1 ♀</specimenCount>
|
||
; from the same location, but
|
||
<date id="FFB46CA33F591E60FE32FE6344640B8E" box="[417,514,398,417]" pageId="8" pageNumber="163" value="1965-12-30">
|
||
<collectingDate id="EFF0954B3F591E60FE32FE6344640B8E" box="[417,514,398,417]" pageId="8" pageNumber="163" value="1965-12-30">30.12.1965</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26FF2FB53F591E60FDA3FE6244CC0B8D" box="[560,682,399,418]" pageId="8" pageNumber="163">L.S. Mahunka</collectorName>
|
||
(
|
||
<collectionCode id="ED1BD2A63F591E60FD26FE62449D0B8D" box="[693,763,399,418]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="163">HNHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B62403E3F591E60FC9AFE6247300B93" ID-GBIF-Occurrence="2449305672" collectionCode="HNHM" collectorName="B. J. Gyorgy" country="Brazil" location="Sao Paulo" pageId="8" pageNumber="163" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="paratype">
|
||
<specimenCount id="9D0C81EA3F591E60FC9AFE62454F0B8D" box="[777,809,395,418]" pageId="8" pageNumber="163" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingCountry id="F31D0AF33F591E60FCA0FE62450F0B8D" box="[819,873,399,418]" name="Brazil" pageId="8" pageNumber="163">Brazil</collectingCountry>
|
||
,
|
||
<collectingRegion id="49CE84813F591E60FCE1FE6245AC0B8D" box="[882,970,399,418]" country="Brazil" name="Sao Paulo" pageId="8" pageNumber="163">São Paulo</collectingRegion>
|
||
, without date, leg.
|
||
<collectorName id="26FF2FB53F591E60FF04FE4746990B93" box="[151,255,425,445]" pageId="8" pageNumber="163">B.J. György</collectorName>
|
||
(
|
||
<collectionCode id="ED1BD2A63F591E60FE99FE4447360B93" box="[266,336,425,444]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="163">HNHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B62403E3F591E60FECDFE4445AA0B93" ID-GBIF-Occurrence="2449305676" box="[350,972,421,445]" collectingDate="1966-05-01" collectionCode="HNHM" collectorName="I. Loksa'" country="Brazil" location="National Park" municipality="Iguacu" pageId="8" pageNumber="163" specimenCount="1" specimenCount-female="1" stateProvince="Parana" typeStatus="paratype">
|
||
<specimenCount id="9D0C81EA3F591E60FECDFE4447190B93" box="[350,383,421,444]" pageId="8" pageNumber="163" type="female">1 ♀</specimenCount>
|
||
, ‘
|
||
<collectingRegion id="49CE84813F591E60FE1FFE4447AA0B93" box="[396,460,425,444]" country="Brazil" name="Parana" pageId="8" pageNumber="163">Paraná</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BD1D0193F591E60FE46FE4444770B93" box="[469,529,425,444]" pageId="8" pageNumber="163">Iguacú</collectingMunicipality>
|
||
,
|
||
<location id="8ED51CB83F591E60FD8AFE4444F70B93" LSID="urn:lsid:plazi:treatment:03A3FB753F591E62FE55FF3142720B5B:8ED51CB83F591E60FD8AFE4444F70B93" box="[537,657,425,444]" country="Brazil" municipality="Iguacu" name="National Park" pageId="8" pageNumber="163" stateProvince="Parana">National Park</location>
|
||
,
|
||
<date id="FFB46CA33F591E60FD09FE44449A0B93" box="[666,764,425,444]" pageId="8" pageNumber="163" value="1966-05-01">
|
||
<collectingDate id="EFF0954B3F591E60FD09FE44449A0B93" box="[666,764,425,444]" pageId="8" pageNumber="163" value="1966-05-01">05.01.1966</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26FF2FB53F591E60FCBAFE4745130B93" box="[809,885,425,445]" pageId="8" pageNumber="163">I. Loksa’</collectorName>
|
||
(
|
||
<collectionCode id="ED1BD2A63F591E60FC13FE4445A00B93" box="[896,966,425,444]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="8" pageNumber="163">HNHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31019E83F591E60FF04FE3845EE0963" pageId="8" pageNumber="163" type="diagnosis">
|
||
<paragraph id="8BB54A633F591E60FF04FE3845EE0963" blockId="8.[151,1127,469,1549]" pageId="8" pageNumber="163">
|
||
<emphasis id="B97E96713F591E60FF04FE38476A0BC2" bold="true" box="[151,268,469,493]" pageId="8" pageNumber="163">Diagnosis.</emphasis>
|
||
This species belongs to the
|
||
<taxonomicName id="4C0A31E03F591E60FDD1FE3845690BC3" authorityName="Fauvel" authorityYear="1863" box="[578,783,469,492]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B97E96713F591E60FDD1FE3845690BC3" box="[578,783,469,492]" italics="true" pageId="8" pageNumber="163">Holotrochus minor</emphasis>
|
||
</taxonomicName>
|
||
group due to its small size, the absence of pubescence on the abdomen and the partly margined front edge of the pronotum. Regarding the shape of the pronotum with its more or less parallel sides, it mostly resembles
|
||
<taxonomicName id="4C0A31E03F591E60FF04FDD8471E0862" authority="Irmler, 1981" authorityName="Irmler" authorityYear="1981" box="[151,376,565,589]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="inpai">
|
||
<emphasis id="B97E96713F591E60FF04FDD846880863" box="[151,238,565,588]" italics="true" pageId="8" pageNumber="163">H. inpai</emphasis>
|
||
<bibRefCitation id="EF9B37923F591E60FF67FDD8471E0862" author="IRMLER U." box="[244,376,565,589]" journalOrPublisher="Entomologische Blatter" pageId="8" pageNumber="163" pagination="143 - 152" part="77" refId="ref10117" refString="IRMLER U. 1981 a: Neue Arten der Gattung Mimogonia Coiffait (1978) aus der Neotropis (Coleoptera, Staphylinidae). Entomologische Blatter 77: 143 - 152." title="Neue Arten der Gattung Mimogonia Coiffait (1978) aus der Neotropis (Coleoptera, Staphylinidae)" type="journal article" year="1981">Irmler, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A31E03F591E60FE38FDD844F60862" authority="Irmler, 2005" authorityName="Irmler" authorityYear="2005" box="[427,656,565,589]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="ashei">
|
||
<emphasis id="B97E96713F591E60FE38FDD844650863" box="[427,515,565,588]" italics="true" pageId="8" pageNumber="163">H. ashei</emphasis>
|
||
<bibRefCitation id="EF9B37923F591E60FD9AFDD844F60862" author="IRMLER U." box="[521,656,565,589]" journalOrPublisher="Studies on Neotropical Fauna and Environment" pageId="8" pageNumber="163" pagination="49 - 54" part="40" refId="ref10251" refString="IRMLER U. 2005 a: New species of the genus Mimogonia (Coleoptera: Staphylinidae, Osoriinae) from the Neotropical region. Studies on Neotropical Fauna and Environment 40: 49 - 54." title="New species of the genus Mimogonia (Coleoptera: Staphylinidae, Osoriinae) from the Neotropical region" type="journal article" year="2005">Irmler, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
. However, the species in the
|
||
<taxonomicName id="4C0A31E03F591E60FC51FDD842440863" authorityName="Fauvel" authorityYear="1863" box="[962,1058,565,588]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="minor">
|
||
<emphasis id="B97E96713F591E60FC51FDD842440863" box="[962,1058,565,588]" italics="true" pageId="8" pageNumber="163">H. minor</emphasis>
|
||
</taxonomicName>
|
||
group are mainly separated by the structure of the endophallus. Most other species in the group have a straight basal part and a twisted apical part of the endophallus. A throughout twisted endophallus is also developed in
|
||
<taxonomicName id="4C0A31E03F591E60FE65FD78456E0882" authority="Irmler, 1981" authorityName="Irmler" authorityYear="1981" box="[502,776,661,685]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="schubarti">
|
||
<emphasis id="B97E96713F591E60FE65FD78441D0883" box="[502,635,661,684]" italics="true" pageId="8" pageNumber="163">H. schubarti</emphasis>
|
||
<bibRefCitation id="EF9B37923F591E60FD12FD78456E0882" author="IRMLER U." box="[641,776,661,685]" journalOrPublisher="Entomologische Blatter" pageId="8" pageNumber="163" pagination="143 - 152" part="77" refId="ref10117" refString="IRMLER U. 1981 a: Neue Arten der Gattung Mimogonia Coiffait (1978) aus der Neotropis (Coleoptera, Staphylinidae). Entomologische Blatter 77: 143 - 152." title="Neue Arten der Gattung Mimogonia Coiffait (1978) aus der Neotropis (Coleoptera, Staphylinidae)" type="journal article" year="1981">Irmler, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
, but in this species the endophallus developed 5 coils with a distinctly larger apical coil. Thus, the species mostly resembles
|
||
<taxonomicName id="4C0A31E03F591E60FF04FD38476608C3" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[151,256,725,748]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="tortilis" status="sp. nov.">
|
||
<emphasis id="B97E96713F591E60FF04FD38476608C3" box="[151,256,725,748]" italics="true" pageId="8" pageNumber="163">H. tortilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D2B0A3F591E60FE9AFD38473B08C3" box="[265,349,725,748]" pageId="8" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
that also has a twisted endophallus, but in
|
||
<taxonomicName id="4C0A31E03F591E60FCA6FD3845F808C3" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[821,926,725,748]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="tortilis" status="sp. nov.">
|
||
<emphasis id="B97E96713F591E60FCA6FD3845F808C3" box="[821,926,725,748]" italics="true" pageId="8" pageNumber="163">H. tortilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D2B0A3F591E60FC35FD38459D08C3" box="[934,1019,725,748]" pageId="8" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
two wide torsions are developed, while in
|
||
<taxonomicName id="4C0A31E03F591E60FE65FD1844FF0923" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[502,665,757,780]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="paraguensis" status="sp. nov.">
|
||
<emphasis id="B97E96713F591E60FE65FD1844FF0923" box="[502,665,757,780]" italics="true" pageId="8" pageNumber="163">H. paraguensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D2B0A3F591E60FD32FD1844920923" box="[673,756,757,780]" pageId="8" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
three short coils are existing. Furthermore, the species can be easily separated by the elytra that have rectangular shoulders in
|
||
<taxonomicName id="4C0A31E03F591E60FF04FCD846990963" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[151,255,821,844]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="tortilis" status="sp. nov.">
|
||
<emphasis id="B97E96713F591E60FF04FCD846990963" box="[151,255,821,844]" italics="true" pageId="8" pageNumber="163">H. tortilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D2B0A3F591E60FE95FCD8473D0963" box="[262,347,821,844]" pageId="8" pageNumber="163" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, but smoothly rounded shoulders in
|
||
<taxonomicName id="4C0A31E03F591E60FD4DFCD845E70963" authorityName="Irmler" authorityYear="2013" box="[734,897,821,844]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="163" phylum="Arthropoda" rank="species" species="paraguensis">
|
||
<emphasis id="B97E96713F591E60FD4DFCD845E70963" box="[734,897,821,844]" italics="true" pageId="8" pageNumber="163">H. paraguensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31019E83F591E62FF04FCB844F30B3B" lastPageId="10" lastPageNumber="165" pageId="8" pageNumber="163" type="description">
|
||
<paragraph id="8BB54A633F591E60FF04FCB8458809A2" blockId="8.[151,1127,469,1549]" pageId="8" pageNumber="163">
|
||
<emphasis id="B97E96713F591E60FF04FCB847440942" bold="true" box="[151,290,853,877]" pageId="8" pageNumber="163">Description.</emphasis>
|
||
Length: 3.0 mm. Colouration dark brown; posterior margin of pronotum and elytra, and parallel to suture of elytra, lighter reddish; legs red; antennae yellow.
|
||
</paragraph>
|
||
<paragraph id="8BB54A633F591E60FF26FC7844610E82" blockId="8.[151,1127,469,1549]" pageId="8" pageNumber="163">
|
||
Head
|
||
<quantity id="4CF2E7863F591E60FF65FC7847310983" box="[246,343,917,941]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="163" unit="mm" value="0.34">0.34 mm</quantity>
|
||
long,
|
||
<quantity id="4CF2E7863F591E60FE0EFC7847990983" box="[413,511,917,941]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="163" unit="mm" value="0.5">0.50 mm</quantity>
|
||
wide; eyes slightly prominent, as long as temples; forehead smoothly rounded, with supraocular margin continuing to base of antennae; within margin four setiferous punctures; on a line between posterior edges of eyes four setiferous punctures; interstices between inner setiferous punctures nearly four times as wide as interstices between inner puncture and outer puncture; along front edge of clypeus with row of short yellow setae; irregular punctation on vertex moderately dense and deep; on average, interstices between punctures slightly wider than diameter of punctures; sparse micro-punctation on disc; at neck, punctation slightly sparser than on fore-head; surface without microsculpture, polished.
|
||
</paragraph>
|
||
<paragraph id="8BB54A633F591E60FF26FB5847AA0F02" blockId="8.[151,1127,469,1549]" pageId="8" pageNumber="163">Antennae slightly longer than head; second antennomere oval, distinctly ticker than conical third; fourth antennomere as thick as third, but only half as long as third; following antennomeres increasing in width; fourth antennomere nearly quadrate; tenth antennomere nearly twice as wide as long.</paragraph>
|
||
<paragraph id="8BB54A633F591E60FF26FAD844D80C22" blockId="8.[151,1127,469,1549]" pageId="8" pageNumber="163">
|
||
Pronotum
|
||
<quantity id="4CF2E7863F591E60FEB5FAD847E00F63" box="[294,390,1333,1357]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="163" unit="mm" value="0.54">0.54 mm</quantity>
|
||
long,
|
||
<quantity id="4CF2E7863F591E60FE5FFAD8444D0F63" box="[460,555,1333,1357]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="8" pageNumber="163" unit="mm" value="0.67">0.67 mm</quantity>
|
||
wide; widest in the posterior third; nearly parallel in middle and shortly rounded at posterior and anterior angles; sides with narrow margin from posterior angles to anterior angles and shortly continuing to anterior edge; shortly in front of anterior angles, margin hidden in dorsal aspect, but again distinctly visible at anterior angles; punctation moderately deep and dense; on average, interstices between punctures slightly wider than diameter of punctures; distinct micro-punctation between normal punctation; surface without microsculpture, polished and shiny.
|
||
</paragraph>
|
||
<paragraph id="8BB54A633F581E61FF03FF3146870B1B" blockId="9.[113,1089,220,404]" pageId="9" pageNumber="164">
|
||
Elytra
|
||
<quantity id="4CF2E7863F581E61FF4BFF3147510ADB" box="[216,311,220,244]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="9" pageNumber="164" unit="mm" value="0.64">0.64 mm</quantity>
|
||
long,
|
||
<quantity id="4CF2E7863F581E61FEE8FF3147BC0ADB" box="[379,474,220,244]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="9" pageNumber="164" unit="mm" value="0.69">0.69 mm</quantity>
|
||
wide; shoulders smoothly rounded; with finer punctation than pronotum, but with deep coriaceous ground-sculpture; surface less shiny than head and pronotum.
|
||
</paragraph>
|
||
<paragraph id="8BB54A633F581E61FF03FED147B80BBB" blockId="9.[113,1089,220,404]" pageId="9" pageNumber="164">Abdomen with fine punctation and weak net-like microsculpture; laterally, with few setiferous punctures; surface still less shiny than elytra. Shape of last sternite triangular and with punctures and ridges at apex.</paragraph>
|
||
<caption id="DF751AEB3F581E61FFE2FA1945A10C0D" ID-DOI="http://doi.org/10.5281/zenodo.3532768" ID-Zenodo-Dep="3532768" httpUri="https://zenodo.org/record/3532768/files/figure.png" pageId="9" pageNumber="164" startId="9.[113,150,1524,1543]" targetBox="[150,1050,427,1501]" targetPageId="9">
|
||
<paragraph id="8BB54A633F581E61FFE2FA1945A10C0D" blockId="9.[113,1088,1524,1570]" pageId="9" pageNumber="164">
|
||
Figs 8–11. 8–
|
||
<taxonomicName id="4C0A31E03F581E61FF74FA1947D10C29" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[231,439,1524,1542]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="164" phylum="Arthropoda" rank="species" species="yasuniensis" status="sp. nov.">
|
||
<emphasis id="B97E96713F581E61FF74FA1947D10C29" box="[231,439,1524,1542]" italics="true" pageId="9" pageNumber="164">Holotrochus yasuniensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D2B0A3F581E61FE2FFA1844660C27" box="[444,512,1525,1544]" pageId="9" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 9 –
|
||
<taxonomicName id="4C0A31E03F581E61FDB7FA1944080C29" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[548,622,1524,1542]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="164" phylum="Arthropoda" rank="species" species="mrazi" status="sp. nov.">
|
||
<emphasis id="B97E96713F581E61FDB7FA1944080C29" box="[548,622,1524,1542]" italics="true" pageId="9" pageNumber="164">H. mrazi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D2B0A3F581E61FDE2FA1844D30C27" box="[625,693,1525,1544]" pageId="9" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 10 –
|
||
<taxonomicName id="4C0A31E03F581E61FD7BFA19450F0C29" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[744,873,1524,1542]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="164" phylum="Arthropoda" rank="species" species="paraguensis" status="sp. nov.">
|
||
<emphasis id="B97E96713F581E61FD7BFA19450F0C29" box="[744,873,1524,1542]" italics="true" pageId="9" pageNumber="164">H. paraguensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D2B0A3F581E61FCFEFA1845D70C27" box="[877,945,1525,1544]" pageId="9" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 11 –
|
||
<taxonomicName id="4C0A31E03F581E61FC4CFA1942260C29" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[991,1088,1524,1542]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="164" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
|
||
<emphasis id="B97E96713F581E61FC4CFA1942260C29" box="[991,1088,1524,1542]" italics="true" pageId="9" pageNumber="164">H. hamatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24D2B0A3F581E61FFE2F9E246D20C0D" box="[113,180,1551,1570]" pageId="9" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Body parts:A – aedeagus; B – antenna, C – tergite XII, D – last sternite. Scale bar: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB54A633F5B1E62FF26FF3144F30B3B" blockId="10.[151,1126,220,373]" pageId="10" pageNumber="165">Aedeagus with straight apex and more or less straight parameres; endophallus with three short torsions and additional half apical torsion.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31019E83F5B1E62FF04FEF0476B0B7B" pageId="10" pageNumber="165" type="etymology">
|
||
<paragraph id="8BB54A633F5B1E62FF04FEF0476B0B7B" blockId="10.[151,1126,220,373]" pageId="10" pageNumber="165">
|
||
<emphasis id="B97E96713F5B1E62FF04FEF0477E0B1A" bold="true" box="[151,280,285,309]" pageId="10" pageNumber="165">Etymology.</emphasis>
|
||
The specific name is derived from the name of the country where the species was found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31019E83F5B1E62FF04FEB142720B5B" box="[151,1044,348,373]" pageId="10" pageNumber="165" type="distribution">
|
||
<paragraph id="8BB54A633F5B1E62FF04FEB142720B5B" blockId="10.[151,1126,220,373]" box="[151,1044,348,373]" pageId="10" pageNumber="165">
|
||
<emphasis id="B97E96713F5B1E62FF04FEB1474C0B5B" bold="true" box="[151,298,348,372]" pageId="10" pageNumber="165">Distribution.</emphasis>
|
||
Known only from
|
||
<collectingCountry id="F31D0AF33F5B1E62FE65FEB0443F0B5A" box="[502,601,349,373]" name="Paraguay" pageId="10" pageNumber="165">Paraguay</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="F31D0AF33F5B1E62FD7CFEB145560B5B" box="[751,816,348,372]" name="Brazil" pageId="10" pageNumber="165">Brazil</collectingCountry>
|
||
(State of
|
||
<collectingRegion id="49CE84813F5B1E62FC0AFEB0426E0B5B" box="[921,1032,348,373]" country="Brazil" name="Sao Paulo" pageId="10" pageNumber="165">São Paulo</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |