164 lines
21 KiB
XML
164 lines
21 KiB
XML
<document id="97C5AD20B1B05B3CDC7756F11608F01E" ID-CLB-Dataset="46226" ID-DOI="10.11646/zootaxa.3779.2.3" ID-GBIF-Dataset="41b9d971-197b-41b4-89df-5160e209981f" ID-ISSN="1175-5326" ID-Zenodo-Dep="252252" ID-ZooBank="17D92CCD-AEC6-47A4-9D47-09756607048E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460397079819" checkinUser="plazi" docAuthor="Boucher, Stéphanie & Wheeler, Terry A." docDate="2014" docId="039687D1FF82FFC1FF47F9ACDBBEFEC6" docLanguage="en" docName="zt03779p176.pdf" docOrigin="Zootaxa 3779 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Melanagromyza pontis Boucher, sp. n." docType="treatment" docVersion="9" lastPageNumber="162" masterDocId="FFAFFFA9FF81FFC4FFD0FFC4D805FFDA" masterDocTitle="Neotropical Agromyzidae (Diptera) of the Mission Géodésique de l’Équateur: Becker (1920) revisited" masterLastPageNumber="176" masterPageNumber="157" pageNumber="160" updateTime="1698298796234" updateUser="ExternalLinkService">
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<mods:titleInfo id="1EA98C8EB7FEA70745B88811FB896167">
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<mods:title id="715A032A6060550534FCE1D1B6859EB0">Neotropical Agromyzidae (Diptera) of the Mission Géodésique de l’Équateur: Becker (1920) revisited</mods:title>
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<mods:name id="B362A5957906489811424D181B5CBBCB" type="personal">
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<mods:namePart id="D19B5E32A17F0D2F060BF7AC3F6CF037">Boucher, Stéphanie</mods:namePart>
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<mods:name id="BA8F095F6571E414CCD1475007379AB7" type="personal">
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<mods:roleTerm id="C99BAADCE485B98A3CB243551A18D7D3">Author</mods:roleTerm>
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<mods:namePart id="3CC6D7C4C179E037BE565BFE3EFC1FD9">Wheeler, Terry A.</mods:namePart>
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<mods:typeOfResource id="06E052FDB2F46343CF8250B073808F23">text</mods:typeOfResource>
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<mods:title id="3BB87BF6D89D9A5A14EDB47238748E4A">Zootaxa</mods:title>
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<mods:part id="8753E5DE85A63CAC639D317561584380">
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<mods:date id="D72CB9356D7A01D3FD6EE951AB69224D">2014</mods:date>
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<mods:number id="130362EC8F462A2CDA53A3DA16406A64">3779</mods:number>
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<mods:number id="F3EF88124DFCB90FD2728DA07F700249">2</mods:number>
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<mods:identifier id="3510298219D3E02B5E2EA2B34D1E05BD" type="DOI">10.11646/zootaxa.3779.2.3</mods:identifier>
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<treatment id="039687D1FF82FFC1FF47F9ACDBBEFEC6" ID-DOI="http://doi.org/10.5281/zenodo.6141285" ID-GBIF-Taxon="119418350" ID-Zenodo-Dep="6141285" LSID="urn:lsid:plazi:treatment:039687D1FF82FFC1FF47F9ACDBBEFEC6" httpUri="http://treatment.plazi.org/id/039687D1FF82FFC1FF47F9ACDBBEFEC6" lastPageId="5" lastPageNumber="162" pageId="3" pageNumber="160">
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<subSubSection id="C325654CFF82FFC7FF47F9ACDAD6F933" pageId="3" pageNumber="160" type="nomenclature">
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<paragraph id="8B8036C7FF82FFC7FF47F9ACDA60F959" blockId="3.[151,613,1640,1700]" box="[151,613,1640,1667]" pageId="3" pageNumber="160">
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<heading id="D0C881ABFF82FFC7FF47F9ACDA60F959" bold="true" box="[151,613,1640,1667]" fontSize="11" level="1" pageId="3" pageNumber="160" reason="1">
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<emphasis id="B94BEAD5FF82FFC7FF47F9ACDA60F959" bold="true" box="[151,613,1640,1667]" pageId="3" pageNumber="160">
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<taxonomicName id="4C3F4D44FF82FFC7FF47F9ACDA19F959" ID-CoL="9HZKV" authority="Boucher" authorityName="Boucher" box="[151,540,1640,1667]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="species" species="pontis" status="sp. nov.">
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<emphasis id="B94BEAD5FF82FFC7FF47F9ACD9AFF958" bold="true" box="[151,426,1640,1666]" italics="true" pageId="3" pageNumber="160">Melanagromyza pontis</emphasis>
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Boucher
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</taxonomicName>
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<taxonomicNameLabel id="A27857AEFF82FFC7FDF3F9ADDA60F959" box="[547,613,1641,1667]" pageId="3" pageNumber="160" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8036C7FF82FFC7FF4EF94FD931F97E" blockId="3.[151,613,1640,1700]" box="[158,308,1675,1700]" pageId="3" pageNumber="160">
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(
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<figureCitation id="13042A42FF82FFC7FF76F94FD929F97E" box="[166,300,1675,1700]" captionStart="FIGURES 10 – 16" captionStartId="4.[151,264,1231,1254]" captionTargetBox="[344,1242,180,1194]" captionTargetId="figure@4.[329,1258,167,1211]" captionTargetPageId="4" captionText="FIGURES 10 – 16. Melanagromyza pontis Boucher, (10) phallus, lateral, (11) phallus, ventral. Inset shows freshly dissected specimen, (12) epandrium, arrow shows small spine at hind corner, (13) epandrium, posterior, arrow shows small patch of setae on surstylus, (14) surstylus, lateral, arrow shows small patch of setae, (15) hypandrium with apodeme, (16) ejaculatory apodeme. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/252255/files/figure.png" pageId="3" pageNumber="160">Figs. 10–16</figureCitation>
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)
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</paragraph>
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<paragraph id="8B8036C7FF82FFC7FF47F910DAD6F933" blockId="3.[151,723,1747,1769]" box="[151,723,1747,1769]" pageId="3" pageNumber="160">
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<taxonomicName id="4C3F4D44FF82FFC7FF47F910D9C2F933" authority="Fallen" authorityName="Fallen" box="[151,455,1747,1769]" class="Insecta" family="Agromyzidae" genus="Agromyza" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="species" species="aeneiventris">
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<emphasis id="B94BEAD5FF82FFC7FF47F910D97AF933" box="[151,383,1748,1769]" italics="true" pageId="3" pageNumber="160">Agromyza aeneiventris</emphasis>
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Fallén
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</taxonomicName>
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, of
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<bibRefCitation id="EFAE4B36FF82FFC7FE3CF917DA85F933" author="Becker" box="[492,640,1747,1769]" pageId="3" pageNumber="160" refString="Becker, T. (1920) Dipteres brachyceres. In: Mission du Service Geographique de l'Armee pour la mesure d'un arc de Meridien Equatoriel en Amerique du Sud sous le controle scientifique de l'Academie des Sciences, 1899 - 1906. Tome 10. Entomologie. - Botanique. Fascicule 2. - Opiliones. - Dipteres. - Myriapodes. [1919]. Gauthier-Villars, Paris. pp. 163 - 215." type="book chapter" year="1920">Becker (1920)</bibRefCitation>
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, in part.
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</paragraph>
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</subSubSection>
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<subSubSection id="C325654CFF82FFC7FF47F8D2DBE2F8F5" box="[151,999,1814,1839]" pageId="3" pageNumber="160" type="materials_examined">
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<paragraph id="8B8036C7FF82FFC7FF47F8D2DBE2F8F5" blockId="3.[151,1436,1814,2019]" box="[151,999,1814,1839]" pageId="3" pageNumber="160">
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<typeStatus id="54848865FF82FFC7FF47F8D2D901F8F5" box="[151,260,1814,1839]" pageId="3" pageNumber="160" type="holotype">
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<emphasis id="B94BEAD5FF82FFC7FF47F8D2D901F8F5" bold="true" box="[151,260,1814,1839]" pageId="3" pageNumber="160">Holotype</emphasis>
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</typeStatus>
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♂.
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<collectingCountry id="F3287657FF82FFC7FEFEF8D2D991F8F5" box="[302,404,1814,1839]" name="Ecuador" pageId="3" pageNumber="160">
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<emphasis id="B94BEAD5FF82FFC7FEFEF8D2D991F8F5" bold="true" box="[302,404,1814,1839]" pageId="3" pageNumber="160">Ecuador</emphasis>
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</collectingCountry>
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: La Rinconada, alt.
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<quantity id="4CC79B22FF82FFC7FDA7F8D3DACBF8F4" box="[631,718,1815,1839]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="160" unit="m" value="3100.0">3100 m</quantity>
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, P. Rivet 1903 (
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<collectionCode id="ED2EAE02FF82FFC7FCAFF8D3DBDEF8F5" LSID="urn:lsid:biocol.org:col:34988" box="[895,987,1815,1839]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="160">MNHN</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C325654CFF82FFC7FF17F8FEDBC5F8AD" pageId="3" pageNumber="160" type="etymology">
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<paragraph id="8B8036C7FF82FFC7FF17F8FEDBC5F8AD" blockId="3.[151,1436,1814,2019]" pageId="3" pageNumber="160">
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<emphasis id="B94BEAD5FF82FFC7FF17F8FED943F889" bold="true" box="[199,326,1850,1875]" pageId="3" pageNumber="160">Etymology</emphasis>
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. The species name is derived from the Latin
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<emphasis id="B94BEAD5FF82FFC7FCBCF8FFDBA4F888" box="[876,929,1851,1874]" italics="true" pageId="3" pageNumber="160">pons</emphasis>
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(bridge), referring to the membranous gap between the basiphallus and mesophallus complex of the male genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="C325654CFF82FFC7FF17F846D93EF839" pageId="3" pageNumber="160" type="diagnosis">
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<paragraph id="8B8036C7FF82FFC7FF17F846D93EF839" blockId="3.[151,1436,1814,2019]" pageId="3" pageNumber="160">
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<emphasis id="B94BEAD5FF82FFC7FF17F846D93FF841" bold="true" box="[199,314,1922,1947]" pageId="3" pageNumber="160">Diagnosis</emphasis>
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. Calypter brown, fringe deep black, gena deep, abdomen mostly reddish-bronze. Three well developed postsutural dc (anterior dc about 0.65 times length of second dc) in addition to 2 much smaller postsutural dc.
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</paragraph>
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</subSubSection>
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<caption id="DF40664FFF85FFC0FF47FB0BD9E6FA98" httpUri="https://zenodo.org/record/252255/files/figure.png" pageId="4" pageNumber="161" targetBox="[344,1242,180,1194]" targetPageId="4">
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<paragraph id="8B8036C7FF85FFC0FF47FB0BD9E6FA98" blockId="4.[151,1436,1231,1346]" pageId="4" pageNumber="161">
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<emphasis id="B94BEAD5FF85FFC0FF47FB0BD94AFB3F" bold="true" box="[151,335,1231,1254]" pageId="4" pageNumber="161">FIGURES 10–16</emphasis>
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.
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<taxonomicName id="4C3F4D44FF85FFC0FE8EFB14DAADFB3C" authority="Boucher" authorityName="Boucher" box="[350,680,1232,1254]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="species" species="pontis">
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<emphasis id="B94BEAD5FF85FFC0FE8EFB14DA4CFB3C" box="[350,585,1232,1254]" italics="true" pageId="4" pageNumber="161">Melanagromyza pontis</emphasis>
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Boucher
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</taxonomicName>
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, (10) phallus, lateral, (11) phallus, ventral. Inset shows freshly dissected specimen, (12) epandrium, arrow shows small spine at hind corner, (13) epandrium, posterior, arrow shows small patch of setae on surstylus, (14) surstylus, lateral, arrow shows small patch of setae, (15) hypandrium with apodeme, (16) ejaculatory apodeme. Scale bars = 0.05 mm.
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</paragraph>
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</caption>
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<subSubSection id="C325654CFF85FFC0FF17FAABDCFEF82B" pageId="4" pageNumber="161" type="description">
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<paragraph id="8B8036C7FF85FFC0FF17FAABDBCFF929" blockId="4.[151,1436,1391,2033]" pageId="4" pageNumber="161">
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<emphasis id="B94BEAD5FF85FFC0FF17FAABD954FA52" bold="true" box="[199,337,1391,1416]" pageId="4" pageNumber="161">Description</emphasis>
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. Ratio of frons width to eye width 1.5 (measured in dorsal view); orbit 0.22 times width of frons (including orbits) at midpoint; parafacial absent or slightly projecting, forming only narrow ring (cheek) below eye; 2 reclinate ors and 2 inclinate ori; orbital setulae erect or reclinate in 2–3 irregular rows; ocellar triangle extending anteriorly to upper ori; first flagellomere rounded with short pubescence apically; arista slightly shorter than maximum eye height and with very short pubescence; gena deep, projecting forward; gena height at midpoint about 0.26 times maximum eye height; antennae separated by flattened keel; upper margin of clypeus wide, more or less rounded, slightly pointed; eye with small patch of hairs dorsally near orbit. Three well developed postsutural dc (anterior dc about 0.65 times length of second dc) in addition to 2 much smaller postsutural dc on each side of welldeveloped anterior dc; acrostichal setulae in about 10 irregular rows; prescutellar acrostichal seta absent; notopleuron with 2 setae; postpronotum with 1 strong seta and 1 setula; wing length
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<quantity id="4CC79B22FF85FFC0FB81F977DCAFF911" box="[1105,1194,1715,1739]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="161" unit="mm" value="2.7">2.7 mm</quantity>
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; R4+5 ending closer to wing tip than M1+2; last section of CuA1 0.54 times length of penultimate.
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</paragraph>
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<paragraph id="8B8036C7FF85FFC0FF17F8C4D91DF8BB" blockId="4.[151,1436,1391,2033]" pageId="4" pageNumber="161">
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<emphasis id="B94BEAD5FF85FFC0FF17F8C4D91FF8C3" bold="true" box="[199,282,1792,1817]" pageId="4" pageNumber="161">Colour</emphasis>
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. Body brownish-black with abdomen mostly reddish-bronze; mesonotum with slight metallic reflection when viewed from posterior; calypter brown with fringe deep black. Orbit and ocellar triangle subshining.
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</paragraph>
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<paragraph id="8B8036C7FF85FFC0FF17F8A8DCFEF82B" blockId="4.[151,1436,1391,2033]" pageId="4" pageNumber="161">
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<emphasis id="B94BEAD5FF85FFC0FF17F8A8D970F85F" bold="true" box="[199,373,1900,1925]" pageId="4" pageNumber="161">Male genitalia</emphasis>
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. Basiphallus U-shaped, widely separated from mesophallus complex by a membranous gap (
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<figureCitation id="13042A42FF85FFC0FF4FF854D8F7F873" box="[159,242,1936,1961]" captionStart="FIGURES 10 – 16" captionStartId="4.[151,264,1231,1254]" captionTargetBox="[344,1242,180,1194]" captionTargetId="figure@4.[329,1258,167,1211]" captionTargetPageId="4" captionText="FIGURES 10 – 16. Melanagromyza pontis Boucher, (10) phallus, lateral, (11) phallus, ventral. Inset shows freshly dissected specimen, (12) epandrium, arrow shows small spine at hind corner, (13) epandrium, posterior, arrow shows small patch of setae on surstylus, (14) surstylus, lateral, arrow shows small patch of setae, (15) hypandrium with apodeme, (16) ejaculatory apodeme. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/252255/files/figure.png" pageId="4" pageNumber="161">Fig. 11</figureCitation>
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); epandrium with patch of short spines internally and one small spine at hind corner (
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<figureCitation id="13042A42FF85FFC0FB1CF854DD24F873" box="[1228,1313,1936,1961]" captionStart="FIGURES 10 – 16" captionStartId="4.[151,264,1231,1254]" captionTargetBox="[344,1242,180,1194]" captionTargetId="figure@4.[329,1258,167,1211]" captionTargetPageId="4" captionText="FIGURES 10 – 16. Melanagromyza pontis Boucher, (10) phallus, lateral, (11) phallus, ventral. Inset shows freshly dissected specimen, (12) epandrium, arrow shows small spine at hind corner, (13) epandrium, posterior, arrow shows small patch of setae on surstylus, (14) surstylus, lateral, arrow shows small patch of setae, (15) hypandrium with apodeme, (16) ejaculatory apodeme. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/252255/files/figure.png" pageId="4" pageNumber="161">Fig. 12</figureCitation>
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); surstylus extended ventrally and bearing patch of short setae at apex (arrows,
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<figureCitation id="13042A42FF85FFC0FC1FF870DC59F817" box="[975,1116,1972,1997]" captionStart="FIGURES 10 – 16" captionStartId="4.[151,264,1231,1254]" captionTargetBox="[344,1242,180,1194]" captionTargetId="figure@4.[329,1258,167,1211]" captionTargetPageId="4" captionText="FIGURES 10 – 16. Melanagromyza pontis Boucher, (10) phallus, lateral, (11) phallus, ventral. Inset shows freshly dissected specimen, (12) epandrium, arrow shows small spine at hind corner, (13) epandrium, posterior, arrow shows small patch of setae on surstylus, (14) surstylus, lateral, arrow shows small patch of setae, (15) hypandrium with apodeme, (16) ejaculatory apodeme. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/252255/files/figure.png" pageId="4" pageNumber="161">Figs. 13–14</figureCitation>
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); hypandrial apodeme well developed (
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<figureCitation id="13042A42FF85FFC0FEC9F81CD969F82B" box="[281,364,2008,2033]" captionStart="FIGURES 10 – 16" captionStartId="4.[151,264,1231,1254]" captionTargetBox="[344,1242,180,1194]" captionTargetId="figure@4.[329,1258,167,1211]" captionTargetPageId="4" captionText="FIGURES 10 – 16. Melanagromyza pontis Boucher, (10) phallus, lateral, (11) phallus, ventral. Inset shows freshly dissected specimen, (12) epandrium, arrow shows small spine at hind corner, (13) epandrium, posterior, arrow shows small patch of setae on surstylus, (14) surstylus, lateral, arrow shows small patch of setae, (15) hypandrium with apodeme, (16) ejaculatory apodeme. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/252255/files/figure.png" pageId="4" pageNumber="161">Fig. 15</figureCitation>
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); ejaculatory apodeme with narrow blade and well-defined medial vein (
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<figureCitation id="13042A42FF85FFC0FB4AF81CDCEBF82B" box="[1178,1262,2008,2033]" captionStart="FIGURES 10 – 16" captionStartId="4.[151,264,1231,1254]" captionTargetBox="[344,1242,180,1194]" captionTargetId="figure@4.[329,1258,167,1211]" captionTargetPageId="4" captionText="FIGURES 10 – 16. Melanagromyza pontis Boucher, (10) phallus, lateral, (11) phallus, ventral. Inset shows freshly dissected specimen, (12) epandrium, arrow shows small spine at hind corner, (13) epandrium, posterior, arrow shows small patch of setae on surstylus, (14) surstylus, lateral, arrow shows small patch of setae, (15) hypandrium with apodeme, (16) ejaculatory apodeme. Scale bars = 0.05 mm." httpUri="https://zenodo.org/record/252255/files/figure.png" pageId="4" pageNumber="161">Fig. 16</figureCitation>
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).
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C325654CFF84FFC1FF17FF53DBBEFEC6" pageId="5" pageNumber="162" type="discussion">
|
||
<paragraph id="8B8036C7FF84FFC1FF17FF53DBBEFEC6" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="162">
|
||
<emphasis id="B94BEAD5FF84FFC1FF17FF53D94DFF6A" bold="true" box="[199,328,151,176]" pageId="5" pageNumber="162">Comments</emphasis>
|
||
. This species is most similar to
|
||
<taxonomicName id="4C3F4D44FF84FFC1FD7DFF53DC84FF6A" authority="Spencer 1973" authorityName="Spencer" authorityYear="1973" box="[685,1153,151,176]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="meracula">
|
||
<emphasis id="B94BEAD5FF84FFC1FD7DFF53DBD6FF6A" box="[685,979,151,176]" italics="true" pageId="5" pageNumber="162">Melanagromyza meracula</emphasis>
|
||
<bibRefCitation id="EFAE4B36FF84FFC1FC09FF5CDC84FF6A" author="Spencer" box="[985,1153,152,176]" pageId="5" pageNumber="162" refString="Spencer, K. A. (1973 a) The Agromyzidae of Venezuela. Revista de la Facultad de Agronomia (Maracay), 7 (2), 5 - 107." type="journal article" year="1973" yearSuffix="a">Spencer 1973a</bibRefCitation>
|
||
</taxonomicName>
|
||
, but can be differentiated from it by the presence of 3 strong postsutural dc in addition to 2 much smaller postsutural dc (only 2 strong dc in
|
||
<taxonomicName id="4C3F4D44FF84FFC1FF47FF25D92FFF22" box="[151,298,223,248]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="meracula">
|
||
<emphasis id="B94BEAD5FF84FFC1FF47FF25D92FFF22" box="[151,298,223,248]" italics="true" pageId="5" pageNumber="162">M. meracula</emphasis>
|
||
</taxonomicName>
|
||
as in most other
|
||
<taxonomicName id="4C3F4D44FF84FFC1FE2EFF1BDAB6FF22" box="[510,691,223,248]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94BEAD5FF84FFC1FE2EFF1BDAB6FF22" box="[510,691,223,248]" italics="true" pageId="5" pageNumber="162">Melanagromyza</emphasis>
|
||
</taxonomicName>
|
||
spp.). The male genitalia are also distinct, with the basiphallus widely separated from the mesophallus (approximated in
|
||
<taxonomicName id="4C3F4D44FF84FFC1FCCBFEC1DBAEFEC7" box="[795,939,260,285]" class="Insecta" family="Agromyzidae" genus="Melanagromyza" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="meracula">
|
||
<emphasis id="B94BEAD5FF84FFC1FCCBFEC1DBAEFEC7" box="[795,939,260,285]" italics="true" pageId="5" pageNumber="162">M. meracula</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |