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<document id="6CF8314DF13395440AD8E0023B0BB375" ID-DOI="10.11646/zootaxa.5353.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8427062" ID-ZooBank="7A24DCBF-BC93-4B25-BB8A-DEAE75167DB4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1696944081533" checkinUser="plazi" docAuthor="Fomichev, Alexander A., Omelko, Mikhail M. &amp; Marusik, Yuri M." docDate="2023" docId="0396878DF5566C773EA3A793F4FBF9AE" docLanguage="en" docName="zootaxa.5353.2.2.pdf" docOrigin="Zootaxa 5353 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5353.2.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acantheis dimidiatus Thorell 1890" docType="treatment" docVersion="3" lastPageNumber="128" masterDocId="FFAFFFF5F55D6C7C3E34A664F771FC06" masterDocTitle="A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species" masterLastPageNumber="130" masterPageNumber="117" pageNumber="128" updateTime="1696945185543" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CEA1E2B9D8976F283C7C54A058C7A470">A survey of the Sumatran Ctenidae (Araneae). 1. Two new Acantheis species</mods:title>
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<mods:namePart id="0A867DCF37E1F290EC8990B88B292659">Fomichev, Alexander A.</mods:namePart>
<mods:affiliation id="3D6AAA87B20249D3BE1D5BC3E395C08B">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia</mods:affiliation>
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<mods:namePart id="12EE4C217F023DDE40F788B02DD62F74">Omelko, Mikhail M.</mods:namePart>
<mods:affiliation id="66F50539376F7C1FDD32BAC33F09C3DB">Federal Scientific Center of East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok 690022, Russia</mods:affiliation>
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<mods:namePart id="0BB8DCECD1CE9427A78EEE5CC8485467">Marusik, Yuri M.</mods:namePart>
<mods:affiliation id="AD377887740122B94A4EB0B1BDDD2AC1">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa &amp; Institute for Biological Problems of the North, Magadan 685000, Russia</mods:affiliation>
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<mods:title id="65B9B53C1B0E3298468CF0A8806F77AF">Zootaxa</mods:title>
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<mods:date id="1B76AE0005802EA7756104048799E4BC">2023</mods:date>
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<treatment id="0396878DF5566C773EA3A793F4FBF9AE" ID-DOI="http://doi.org/10.5281/zenodo.8427172" ID-Zenodo-Dep="8427172" LSID="urn:lsid:plazi:treatment:0396878DF5566C773EA3A793F4FBF9AE" httpUri="http://treatment.plazi.org/id/0396878DF5566C773EA3A793F4FBF9AE" lastPageNumber="128" pageId="11" pageNumber="128">
<subSubSection id="C3256510F5566C773EA3A793F526FE17" box="[151,599,503,530]" pageId="11" pageNumber="128" type="nomenclature">
<paragraph id="8B80369BF5566C773EA3A793F526FE17" blockId="11.[151,599,503,566]" box="[151,599,503,530]" pageId="11" pageNumber="128">
<heading id="D0C881F7F5566C773EA3A793F526FE17" bold="true" box="[151,599,503,530]" fontSize="11" level="1" pageId="11" pageNumber="128" reason="1">
<taxonomicName id="4C3F4D18F5566C773EA3A793F526FE17" ID-CoL="8TFW" authority="(Thorell, 1890)" authorityName="Thorell" authorityYear="1890" baseAuthorityName="Thorell" baseAuthorityYear="1890" box="[151,599,503,530]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B94BEA89F5566C773EA3A793F526FE17" bold="true" box="[151,599,503,530]" pageId="11" pageNumber="128">
<emphasis id="B94BEA89F5566C773EA3A793F6E7FE14" bold="true" box="[151,406,503,530]" italics="true" pageId="11" pageNumber="128">Acantheis dimidiatus</emphasis>
(
<bibRefCitation id="EFAE4B6AF5566C773F92A793F53FFE17" author="Thorell, T." box="[422,590,503,529]" pageId="11" pageNumber="128" pagination="132 - 172" refId="ref7277" refString="Thorell, T. (1890) Diagnoses aranearum aliquot novarum in Indo-Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30, 132 - 172." type="journal article" year="1890">Thorell, 1890</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3256510F5566C773EA3A478F664FE30" box="[151,277,540,566]" pageId="11" pageNumber="128" type="description">
<paragraph id="8B80369BF5566C773EA3A478F664FE30" blockId="11.[151,599,503,566]" box="[151,277,540,566]" pageId="11" pageNumber="128">
<figureCitation id="13042A1EF5566C773EA3A478F664FE30" box="[151,277,540,566]" captionStart-0="FIGURES 36–44" captionStart-1="FIGURES 46–48" captionStartId-0="10.[151,264,1943,1967]" captionStartId-1="12.[151,264,1922,1946]" captionTargetBox-0="[179,1408,181,1919]" captionTargetBox-1="[151,1436,617,1898]" captionTargetId-0="figure-21@10.[179,1408,181,1919]" captionTargetId-1="figure-119@12.[151,1436,617,1898]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionText-0="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." captionText-1="FIGURES 46–48. Collecting localities of A. sergeimishenini sp. n. (circle), A. andreimishenini sp. n. (square), and A. dimidiatus Thorell, 1890 (triangle). The frame on Fig. 48 refers to the content of Fig. 47 and the frame on Fig. 47 refers to the content of Fig. 46." figureDoi-0="http://doi.org/10.5281/zenodo.8427074" figureDoi-1="http://doi.org/10.5281/zenodo.8427076" httpUri-0="https://zenodo.org/record/8427074/files/figure.png" httpUri-1="https://zenodo.org/record/8427076/files/figure.png" pageId="11" pageNumber="128">Figs 36–48</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3256510F5566C773EA3A404F54BFE91" pageId="11" pageNumber="128" type="reference_group">
<paragraph id="8B80369BF5566C773EA3A404F5E0FE7D" blockId="11.[151,657,608,663]" box="[151,657,608,635]" pageId="11" pageNumber="128">
<treatmentCitationGroup id="AB2F11B5F5566C773EA3A404F5E0FE7D" box="[151,657,608,635]" pageId="11" pageNumber="128">
<treatmentCitation id="0A9E108AF5566C773EA3A404F510FE7E" author="Thorell, T." box="[151,609,608,632]" page="134" pageId="11" pageNumber="128" year="1890">
<taxonomicName id="4C3F4D18F5566C773EA3A404F510FE7E" authority="Thorell, 1890: 134" authorityName="Thorell" authorityPageNumber="134" authorityYear="1890" box="[151,609,608,632]" class="Arachnida" family="Ctenidae" genus="Acanthoctenus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B94BEA89F5566C773EA3A404F6EFFE7E" box="[151,414,608,632]" italics="true" pageId="11" pageNumber="128">Acanthoctenus dimidiatus</emphasis>
<bibRefCitation id="EFAE4B6AF5566C773F90A404F510FE7E" author="Thorell, T." box="[420,609,608,632]" pageId="11" pageNumber="128" pagination="132 - 172" refId="ref7277" refString="Thorell, T. (1890) Diagnoses aranearum aliquot novarum in Indo-Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30, 132 - 172." type="journal article" year="1890">Thorell, 1890: 134</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(♁).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B80369BF5566C773EA3A41BF54BFE91" blockId="11.[151,657,608,663]" box="[151,570,639,663]" pageId="11" pageNumber="128">
<treatmentCitationGroup id="AB2F11B5F5566C773EA3A41BF54BFE91" box="[151,570,639,663]" pageId="11" pageNumber="128">
<taxonomicName id="4C3F4D18F5566C773EA3A41BF61DFE91" ID-CoL="8TFW" authorityName="Thorell" authorityYear="1890" baseAuthorityName="Thorell" baseAuthorityYear="1890" box="[151,364,639,663]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B94BEA89F5566C773EA3A41BF61DFE91" box="[151,364,639,663]" italics="true" pageId="11" pageNumber="128">Acantheis dimidiatus</emphasis>
</taxonomicName>
—
<treatmentCitation id="0A9E108AF5566C773FB2A41BF547FE91" author="Simon, E." box="[390,566,639,663]" page="117" pageId="11" pageNumber="128" year="1897">
<bibRefCitation id="EFAE4B6AF5566C773FB2A41BF547FE91" author="Simon, E." box="[390,566,639,663]" pageId="11" pageNumber="128" pagination="1 - 192" refId="ref7191" refString="Simon, E. (1897) Histoire naturelle des araignees. Deuxieme edition, tome second. Roret, Paris, pp. 1 - 192. https: // doi. org / 10.5962 / bhl. title. 51973" type="book chapter" year="1897">Simon 1897: 117</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3256510F5566C773EA3A4DAF7AEFEFA" pageId="11" pageNumber="128" type="materials_examined">
<paragraph id="8B80369BF5566C773EA3A4DAF7AEFEFA" blockId="11.[151,1437,702,1449]" pageId="11" pageNumber="128">
<emphasis id="B94BEA89F5566C773EA3A4DAF6F0FEDE" bold="true" box="[151,385,702,728]" pageId="11" pageNumber="128">Material examined.</emphasis>
<materialsCitation id="3B573CC6F5566C773FB3A4A4F7ABFEFA" collectionCode="MSNG" collectorName="Singalong, O" country="Indonesia" location="Sumatra" pageId="11" pageNumber="128" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
1♁ (
<typeStatus id="54848839F5566C773F8DA4DAF56FFEDE" box="[441,542,702,728]" pageId="11" pageNumber="128">holotype</typeStatus>
) (
<collectionCode id="ED2EAE5EF5566C773C00A4DBF5FAFEDF" box="[564,651,703,729]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="11" pageNumber="128">MSNG</collectionCode>
),
<collectingCountry id="F328760BF5566C773CAAA4DBF44AFEDF" box="[670,827,703,729]" name="Indonesia" pageId="11" pageNumber="128">INDONESIA</collectingCountry>
:
<emphasis id="B94BEA89F5566C773D71A4DAF4D8FEDE" bold="true" box="[837,937,702,728]" italics="true" pageId="11" pageNumber="128">
<collectingRegion id="49FBF879F5566C773D71A4DAF4D8FEDE" box="[837,937,702,728]" country="Indonesia" name="Sumatera Utara" pageId="11" pageNumber="128">Sumatra</collectingRegion>
</emphasis>
Island:
<collectorName id="26CA534DF5566C773A37A4DAF3E8FEDF" box="[1027,1177,702,729]" pageId="11" pageNumber="128">Singalong, O</collectorName>
. Beccari. Examined by photo
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256510F5566C773EF3A562F269F8D7" pageId="11" pageNumber="128" type="diagnosis">
<paragraph id="8B80369BF5566C773EF3A562F269F8D7" blockId="11.[151,1437,702,1449]" pageId="11" pageNumber="128">
<emphasis id="B94BEA89F5566C773EF3A562F631FF26" bold="true" box="[199,320,774,800]" pageId="11" pageNumber="128">Diagnosis.</emphasis>
The species is similar to
<taxonomicName id="4C3F4D18F5566C773C4BA563F597FF26" authorityName="Thorell" authorityYear="1890" box="[639,742,774,801]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94BEA89F5566C773C4BA563F597FF26" box="[639,742,774,801]" italics="true" pageId="11" pageNumber="128">A. laetus</emphasis>
</taxonomicName>
from
<collectingRegion id="49FBF879F5566C773D02A562F4C6FF26" box="[822,951,774,800]" country="Indonesia" name="Kalimantan Utara" pageId="11" pageNumber="128">Kalimantan</collectingRegion>
and to
<taxonomicName id="4C3F4D18F5566C773A29A563F380FF26" authority="Fomichev &amp; Omelko &amp; Marusik, 2023" authorityName="Fomichev &amp; Omelko &amp; Marusik" authorityYear="2023" box="[1053,1265,774,801]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="sergeimishenini" status="sp. n.">
<emphasis id="B94BEA89F5566C773A29A563F380FF26" box="[1053,1265,774,801]" italics="true" pageId="11" pageNumber="128">A. sergeimishenini</emphasis>
</taxonomicName>
<emphasis id="B94BEA89F5566C773AC9A563F232FF27" bold="true" box="[1277,1347,775,801]" pageId="11" pageNumber="128">
<taxonomicNameLabel id="A27857F2F5566C773AC9A563F232FF27" box="[1277,1347,775,801]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3F4D18F5566C773BB0A563F63BFF42" authority="Fomichev &amp; Omelko &amp; Marusik, 2023" authorityName="Fomichev &amp; Omelko &amp; Marusik" authorityYear="2023" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="andreimishenini" status="sp. n.">
<emphasis id="B94BEA89F5566C773BB0A563F63BFF42" italics="true" pageId="11" pageNumber="128">A. andreimishenini</emphasis>
</taxonomicName>
<emphasis id="B94BEA89F5566C773F60A54FF6E4FF43" bold="true" box="[340,405,811,837]" pageId="11" pageNumber="128">
<taxonomicNameLabel id="A27857F2F5566C773F60A54FF6E4FF43" box="[340,405,811,837]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from
<collectingRegion id="49FBF879F5566C773FE9A54FF548FF43" box="[477,569,811,837]" country="Indonesia" name="Sumatera Utara" pageId="11" pageNumber="128">Sumatra</collectingRegion>
in having the slender embolus (
<emphasis id="B94BEA89F5566C773D9EA54FF4BEFF42" box="[938,975,811,836]" italics="true" pageId="11" pageNumber="128">Em</emphasis>
), the medially located tegular apophysis (
<emphasis id="B94BEA89F5566C773E94A52BF7CFFF6F" box="[160,190,847,873]" italics="true" pageId="11" pageNumber="128">TA</emphasis>
) and the elongated cymbial tip. The male of
<taxonomicName id="4C3F4D18F5566C773C9FA52BF44EFF6E" authorityName="Thorell" authorityYear="1890" baseAuthorityName="Thorell" baseAuthorityYear="1890" box="[683,831,846,873]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B94BEA89F5566C773C9FA52BF44EFF6E" box="[683,831,846,873]" italics="true" pageId="11" pageNumber="128">A. dimidiatus</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="4C3F4D18F5566C773A29A52BF30CFF6E" authorityName="Thorell" authorityYear="1890" box="[1053,1149,846,873]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="laetus">
<emphasis id="B94BEA89F5566C773A29A52BF30CFF6E" box="[1053,1149,846,873]" italics="true" pageId="11" pageNumber="128">A. laetus</emphasis>
</taxonomicName>
by by the strongly curved embolus (
<emphasis id="B94BEA89F5566C773F37A517F66BFF8A" box="[259,282,883,908]" italics="true" pageId="11" pageNumber="128">vs</emphasis>
. smoothly rounded, cf.
<figureCitation id="13042A1EF5566C773C2DA516F52FFF8B" box="[537,606,882,909]" captionStart="FIGURES 36–44" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-21@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." figureDoi="http://doi.org/10.5281/zenodo.8427074" httpUri="https://zenodo.org/record/8427074/files/figure.png" pageId="11" pageNumber="128">Fig 43</figureCitation>
and
<bibRefCitation id="EFAE4B6AF5566C773CBBA516F442FF8A" author="Lehtinen, P. T." box="[655,819,882,908]" pageId="11" pageNumber="128" pagination="199 - 468" refId="ref6878" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen 1967</bibRefCitation>
: fig. 410).The male of
<taxonomicName id="4C3F4D18F5566C773A1CA517F3CBFF8A" authorityName="Thorell" authorityYear="1890" baseAuthorityName="Thorell" baseAuthorityYear="1890" box="[1064,1210,882,909]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B94BEA89F5566C773A1CA517F3CBFF8A" box="[1064,1210,882,909]" italics="true" pageId="11" pageNumber="128">A. dimidiatus</emphasis>
</taxonomicName>
can be distinguished from that of
<taxonomicName id="4C3F4D18F5566C773F15A5F3F69EFFB6" authority="Fomichev &amp; Omelko &amp; Marusik, 2023" authorityName="Fomichev &amp; Omelko &amp; Marusik" authorityYear="2023" box="[289,495,918,945]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="sergeimishenini" status="sp. n.">
<emphasis id="B94BEA89F5566C773F15A5F3F69EFFB6" box="[289,495,918,945]" italics="true" pageId="11" pageNumber="128">A. sergeimishenini</emphasis>
</taxonomicName>
<emphasis id="B94BEA89F5566C773FC1A5F3F545FFB7" bold="true" box="[501,564,919,945]" pageId="11" pageNumber="128">
<taxonomicNameLabel id="A27857F2F5566C773FC1A5F3F545FFB7" box="[501,564,919,945]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the non-bifurcated retrolateral tibial apophsis (
<emphasis id="B94BEA89F5566C773A6BA5F3F3FFFFB7" box="[1119,1166,919,945]" italics="true" pageId="11" pageNumber="128">RTA</emphasis>
) (
<emphasis id="B94BEA89F5566C773A93A5F3F3CFFFB6" box="[1191,1214,919,944]" italics="true" pageId="11" pageNumber="128">vs</emphasis>
. bifurcated, cf.
<figureCitation id="13042A1EF5566C773B58A5F2F7C2FFD3" captionStart="FIGURES 36–44" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-21@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." figureDoi="http://doi.org/10.5281/zenodo.8427074" httpUri="https://zenodo.org/record/8427074/files/figure.png" pageId="11" pageNumber="128">Figs 41</figureCitation>
and
<figureCitation id="13042A1EF5566C773EDDA5DEF676FFD3" box="[233,263,954,981]" captionStart="FIGURES 12–21" captionStartId="5.[151,264,1943,1967]" captionTargetBox="[186,1401,186,1920]" captionTargetId="figure-19@5.[186,1401,181,1920]" captionTargetPageId="5" captionText="FIGURES 12–21. Male palp of Acantheis sergeimishenini sp. n. (12–16), and A. andreimishenini sp. n. (17–21). 12, 17—whole palp, retrolateral; 13, 18—terminal part of the palp, prolateral; 14, 19—ditto, ventral; 15, 20—ditto, retrolateral; 16, 21—ditto, dorsal. Scale: 12, 17=1 mm; 13–16, 18–21=0.5 mm. Abbreviations: RTA—retrolateral tibial apophysis." figureDoi="http://doi.org/10.5281/zenodo.8427068" httpUri="https://zenodo.org/record/8427068/files/figure.png" pageId="11" pageNumber="128">15</figureCitation>
), the triangular tegular apophysis (
<emphasis id="B94BEA89F5566C773CBFA5DFF5D3FFD2" box="[651,674,955,980]" italics="true" pageId="11" pageNumber="128">vs</emphasis>
. comma-shaped, cf.
<figureCitation id="13042A1EF5566C773DBDA5DEF4ADFFD3" box="[905,988,954,981]" captionStart="FIGURES 36–44" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-21@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." figureDoi="http://doi.org/10.5281/zenodo.8427074" httpUri="https://zenodo.org/record/8427074/files/figure.png" pageId="11" pageNumber="128">Figs 43</figureCitation>
and
<figureCitation id="13042A1EF5566C773A26A5DFF340FFD3" box="[1042,1073,955,981]" captionStart="FIGURES 22–29" captionStartId="8.[151,264,1747,1771]" captionTargetBox="[151,1436,193,1724]" captionTargetId="figure-21@8.[151,1436,181,1724]" captionTargetPageId="8" captionText="FIGURES 22–29. Bulb of Acantheis sergeimishenini sp. n. (22–25), and A. andreimishenini sp. n. (26–29). 22, 26—prolateral; 23, 27—ventral; 24, 28—retrolateral; 25, 29—apical. Scale: 0.2 mm. Abbreviations: BP—basal process of embolus, Cn— conductor, DB—dorsal branch of embolus, EB—embolic base, Em—embolus, ES—embolic spike, ME—membranous extension of embolus, SD—sperm duct, SN—subtegular notch, St—subtegulum, TA—tegular apophysis, Te—tegulum, TP—tegular process, VB—ventral branch of embolus." figureDoi="http://doi.org/10.5281/zenodo.8427070" httpUri="https://zenodo.org/record/8427070/files/figure.png" pageId="11" pageNumber="128">23</figureCitation>
), and the ventral embolic branch twice thinner than dorsal one (
<emphasis id="B94BEA89F5566C773FDEA5BBF570FFFE" box="[490,513,991,1016]" italics="true" pageId="11" pageNumber="128">vs</emphasis>
. ventral embolic branch twice wider than dorsal one, cf.
<figureCitation id="13042A1EF5566C773A4FA5BAF3BFFFFF" box="[1147,1230,990,1017]" captionStart="FIGURES 36–44" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-21@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." figureDoi="http://doi.org/10.5281/zenodo.8427074" httpUri="https://zenodo.org/record/8427074/files/figure.png" pageId="11" pageNumber="128">Figs 41</figureCitation>
and
<figureCitation id="13042A1EF5566C773B30A5BBF255FFFF" box="[1284,1316,991,1017]" captionStart="FIGURES 30–35" captionStartId="9.[151,264,1839,1863]" captionTargetBox="[183,1401,194,1814]" captionTargetId="figure-19@9.[183,1404,181,1815]" captionTargetPageId="9" captionText="FIGURES 30–35. Terminal part of the embolus (30–31) and female genitalia (32–35) of Acantheis sergeimeshenini sp. n. (30), and A. andreimishenini sp. n. (31–35). 30, 31—retrolateral; 32—intact epigyne, ventral; 33—macerated epigyne, ventral; 34—endogyne, dorsal; 35—epigyne, posterior. Scale: 0.2 mm. Abbreviations: Cn—conductor, DB—dorsal branch of embolus, Em—embolus, ET—epigynal tooth, Fo—fovea, ME—membranous extension of embolus, RI—receptacle’s chamber I, RII—receptacle’s chamber II, Re—receptacle, SB—septal base, SH—septal hump, SS—septal stem, VB—ventral branch of embolus." figureDoi="http://doi.org/10.5281/zenodo.8427072" httpUri="https://zenodo.org/record/8427072/files/figure.png" pageId="11" pageNumber="128">30</figureCitation>
). From the males of
<taxonomicName id="4C3F4D18F5566C773F35A267F6A5F81A" authority="Fomichev &amp; Omelko &amp; Marusik, 2023" authorityName="Fomichev &amp; Omelko &amp; Marusik" authorityYear="2023" box="[257,468,1026,1053]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="andreimishenini" status="sp. n.">
<emphasis id="B94BEA89F5566C773F35A267F6A5F81A" box="[257,468,1026,1053]" italics="true" pageId="11" pageNumber="128">A. andreimishenini</emphasis>
</taxonomicName>
<emphasis id="B94BEA89F5566C773FE9A267F56FF81B" bold="true" box="[477,542,1027,1053]" pageId="11" pageNumber="128">
<taxonomicNameLabel id="A27857F2F5566C773FE9A267F56FF81B" box="[477,542,1027,1053]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
those of
<taxonomicName id="4C3F4D18F5566C773CBDA267F46EF81A" authorityName="Thorell" authorityYear="1890" baseAuthorityName="Thorell" baseAuthorityYear="1890" box="[649,799,1026,1053]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B94BEA89F5566C773CBDA267F46EF81A" box="[649,799,1026,1053]" italics="true" pageId="11" pageNumber="128">A. dimidiatus</emphasis>
</taxonomicName>
differs by the dorsal surface of opisthosoma with a thin stripe reaching spinnerets (
<emphasis id="B94BEA89F5566C773FF4A243F6A6F846" box="[448,471,1063,1088]" italics="true" pageId="11" pageNumber="128">vs</emphasis>
. wide cardiac mark reaching middle part of opisthosoma, cf.
<figureCitation id="13042A1EF5566C773A4DA242F3BBF846" box="[1145,1226,1062,1088]" captionStart="FIGURES 36–44" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-21@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." figureDoi="http://doi.org/10.5281/zenodo.8427074" httpUri="https://zenodo.org/record/8427074/files/figure.png" pageId="11" pageNumber="128">Figs 36</figureCitation>
and
<figureCitation id="13042A1EF5566C773ACAA243F27CF847" box="[1278,1293,1063,1089]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[181,1407,191,1916]" captionTargetId="figure-19@3.[177,1409,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Acantheis sergeimishenini sp. n. (1–2), and A. andreimishenini sp. n. (3–6). 1, 3—male, dorsal; 2, 4—male, ventral; 5—female, dorsal; 6—female, ventral. Scale: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8427064" httpUri="https://zenodo.org/record/8427064/files/figure.png" pageId="11" pageNumber="128">3</figureCitation>
), the median band on the carapace as wide as distance between PME (
<emphasis id="B94BEA89F5566C773D16A22FF448F862" box="[802,825,1099,1124]" italics="true" pageId="11" pageNumber="128">vs</emphasis>
. as wide as the distance between PLE), the triangular tegular apophysis (
<emphasis id="B94BEA89F5566C773F5AA20BF6FDF88F" box="[366,396,1135,1161]" italics="true" pageId="11" pageNumber="128">TA</emphasis>
) (
<emphasis id="B94BEA89F5566C773F9CA20BF6CEF88E" box="[424,447,1135,1160]" italics="true" pageId="11" pageNumber="128">vs</emphasis>
. circular, cf.
<figureCitation id="13042A1EF5566C773C6CA20AF5DDF88F" box="[600,684,1134,1161]" captionStart="FIGURES 36–44" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-21@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." figureDoi="http://doi.org/10.5281/zenodo.8427074" httpUri="https://zenodo.org/record/8427074/files/figure.png" pageId="11" pageNumber="128">Figs 43</figureCitation>
and
<figureCitation id="13042A1EF5566C773CD2A20BF474F88F" box="[742,773,1135,1161]" captionStart="FIGURES 22–29" captionStartId="8.[151,264,1747,1771]" captionTargetBox="[151,1436,193,1724]" captionTargetId="figure-21@8.[151,1436,181,1724]" captionTargetPageId="8" captionText="FIGURES 22–29. Bulb of Acantheis sergeimishenini sp. n. (22–25), and A. andreimishenini sp. n. (26–29). 22, 26—prolateral; 23, 27—ventral; 24, 28—retrolateral; 25, 29—apical. Scale: 0.2 mm. Abbreviations: BP—basal process of embolus, Cn— conductor, DB—dorsal branch of embolus, EB—embolic base, Em—embolus, ES—embolic spike, ME—membranous extension of embolus, SD—sperm duct, SN—subtegular notch, St—subtegulum, TA—tegular apophysis, Te—tegulum, TP—tegular process, VB—ventral branch of embolus." figureDoi="http://doi.org/10.5281/zenodo.8427070" httpUri="https://zenodo.org/record/8427070/files/figure.png" pageId="11" pageNumber="128">27</figureCitation>
), and the ventral embolic branch twice thinner than dorsal one (
<emphasis id="B94BEA89F5566C773EFBA2F7F797F8AA" box="[207,230,1171,1196]" italics="true" pageId="11" pageNumber="128">vs</emphasis>
. dorsal and ventral embolic branches equal in size, cf.
<figureCitation id="13042A1EF5566C773D7DA2F6F4EAF8AB" box="[841,923,1170,1197]" captionStart="FIGURES 36–44" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-21@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." figureDoi="http://doi.org/10.5281/zenodo.8427074" httpUri="https://zenodo.org/record/8427074/files/figure.png" pageId="11" pageNumber="128">Figs 41</figureCitation>
and
<figureCitation id="13042A1EF5566C773DE5A2F7F481F8AB" box="[977,1008,1171,1197]" captionStart="FIGURES 30–35" captionStartId="9.[151,264,1839,1863]" captionTargetBox="[183,1401,194,1814]" captionTargetId="figure-19@9.[183,1404,181,1815]" captionTargetPageId="9" captionText="FIGURES 30–35. Terminal part of the embolus (30–31) and female genitalia (32–35) of Acantheis sergeimeshenini sp. n. (30), and A. andreimishenini sp. n. (31–35). 30, 31—retrolateral; 32—intact epigyne, ventral; 33—macerated epigyne, ventral; 34—endogyne, dorsal; 35—epigyne, posterior. Scale: 0.2 mm. Abbreviations: Cn—conductor, DB—dorsal branch of embolus, Em—embolus, ET—epigynal tooth, Fo—fovea, ME—membranous extension of embolus, RI—receptacle’s chamber I, RII—receptacle’s chamber II, Re—receptacle, SB—septal base, SH—septal hump, SS—septal stem, VB—ventral branch of embolus." figureDoi="http://doi.org/10.5281/zenodo.8427072" httpUri="https://zenodo.org/record/8427072/files/figure.png" pageId="11" pageNumber="128">31</figureCitation>
). For a complete list of the differences between the males of
<taxonomicName id="4C3F4D18F5566C773FBDA2D3F56CF8D6" authorityName="Thorell" authorityYear="1890" baseAuthorityName="Thorell" baseAuthorityYear="1890" box="[393,541,1206,1233]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="dimidiatus">
<emphasis id="B94BEA89F5566C773FBDA2D3F56CF8D6" box="[393,541,1206,1233]" italics="true" pageId="11" pageNumber="128">A. dimidiatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3F4D18F5566C773C1FA2D3F589F8D6" authority="Fomichev &amp; Omelko &amp; Marusik, 2023" authorityName="Fomichev &amp; Omelko &amp; Marusik" authorityYear="2023" box="[555,760,1206,1233]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="sergeimishenini" status="sp. n.">
<emphasis id="B94BEA89F5566C773C1FA2D3F589F8D6" box="[555,760,1206,1233]" italics="true" pageId="11" pageNumber="128">A. sergeimishenini</emphasis>
</taxonomicName>
<emphasis id="B94BEA89F5566C773CCBA2D3F44FF8D7" bold="true" box="[767,830,1207,1233]" pageId="11" pageNumber="128">
<taxonomicNameLabel id="A27857F2F5566C773CCBA2D3F44FF8D7" box="[767,830,1207,1233]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C3F4D18F5566C773D47A2D3F33BF8D6" authority="Fomichev &amp; Omelko &amp; Marusik, 2023" authorityName="Fomichev &amp; Omelko &amp; Marusik" authorityYear="2023" box="[883,1098,1206,1233]" class="Arachnida" family="Ctenidae" genus="Acantheis" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="andreimeshenini" status="sp. n.">
<emphasis id="B94BEA89F5566C773D47A2D3F33BF8D6" box="[883,1098,1206,1233]" italics="true" pageId="11" pageNumber="128">A. andreimeshenini</emphasis>
</taxonomicName>
<emphasis id="B94BEA89F5566C773A65A2D3F3FEF8D7" bold="true" box="[1105,1167,1207,1233]" pageId="11" pageNumber="128">
<taxonomicNameLabel id="A27857F2F5566C773A65A2D3F3FEF8D7" box="[1105,1167,1207,1233]" pageId="11" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
see
<tableCitation id="C6BD0320F5566C773AF4A2D2F264F8D7" box="[1216,1301,1206,1233]" captionStart="TABLE 7" captionStartId="11.[152,244,1496,1520]" captionText="TABLE 7. Differences between males of Sumatran species of Acantheis.?—the state of the character is unclear." pageId="11" pageNumber="128">Table 7</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3256510F5566C773EF3A2BEF6EFF982" pageId="11" pageNumber="128" type="description">
<paragraph id="8B80369BF5566C773EF3A2BEF41CF967" blockId="11.[151,1437,702,1449]" pageId="11" pageNumber="128">
<emphasis id="B94BEA89F5566C773EF3A2BEF628F8F2" bold="true" box="[199,345,1242,1268]" pageId="11" pageNumber="128">Description.</emphasis>
Male palp as shown in
<figureCitation id="13042A1EF5566C773C57A2BEF592F8F3" box="[611,739,1242,1269]" captionStart="FIGURES 36–44" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-21@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURES 36–44. Habitus (36), whole palp (37–38), terminal part of the palp (39–42), bulb (43) and labels (44–45) of the holotype of Acantheis dimidiatus Thorell, 1890. 36, 42—dorsal; 37, 39—prolateral; 38, 41—retrolateral; 40, 43—ventral." figureDoi="http://doi.org/10.5281/zenodo.8427074" httpUri="https://zenodo.org/record/8427074/files/figure.png" pageId="11" pageNumber="128">Figs 37–43</figureCitation>
. Tibia ca. 4.6 longer than wide, 4 very long spines, longest ca. 0.7 of tibia length. Retrolateral tibial apophysis small, non-bifurcated. Cymbial tip about 0.42 of its length, slightly shorter than bulb. Sperm duct clearly visible only in retrolateral view. Tegular apophysis triangular. Embolus strongly curved. Ventral branch of embolus twice thinner than dorsal one.
</paragraph>
<paragraph id="8B80369BF5566C773EF3A30EF6EFF982" blockId="11.[151,1437,702,1449]" box="[199,414,1386,1412]" pageId="11" pageNumber="128">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3256510F5566C773EF3A3EAF4FBF9AE" box="[199,906,1422,1449]" pageId="11" pageNumber="128" type="distribution">
<paragraph id="8B80369BF5566C773EF3A3EAF4FBF9AE" blockId="11.[151,1437,702,1449]" box="[199,906,1422,1449]" pageId="11" pageNumber="128">
<emphasis id="B94BEA89F5566C773EF3A3EAF610F9AE" bold="true" box="[199,353,1422,1448]" pageId="11" pageNumber="128">Distribution.</emphasis>
Only known from the
<typeStatus id="54848839F5566C773C54A3EBF5E1F9AF" box="[608,656,1423,1449]" pageId="11" pageNumber="128">type</typeStatus>
locality (
<figureCitation id="13042A1EF5566C773CCEA3EAF40FF9AE" box="[762,894,1422,1448]" captionStart="FIGURES 46–48" captionStartId="12.[151,264,1922,1946]" captionTargetBox="[151,1436,617,1898]" captionTargetId="figure-119@12.[151,1436,617,1898]" captionTargetPageId="12" captionText="FIGURES 46–48. Collecting localities of A. sergeimishenini sp. n. (circle), A. andreimishenini sp. n. (square), and A. dimidiatus Thorell, 1890 (triangle). The frame on Fig. 48 refers to the content of Fig. 47 and the frame on Fig. 47 refers to the content of Fig. 46." figureDoi="http://doi.org/10.5281/zenodo.8427076" httpUri="https://zenodo.org/record/8427076/files/figure.png" pageId="11" pageNumber="128">Figs 46–48</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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