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<document id="D4733209BA2221165E55EC43F82ADFD7" ID-CLB-Dataset="24419" ID-DOI="10.11646/zootaxa.4718.3.6" ID-GBIF-Dataset="13c50c08-32cd-415a-818f-632ef4736aa6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3602528" ID-ZooBank="40857F32-81DA-4DFC-A3B9-F2AA1B485542" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578565019560" checkinUser="plazi" docAuthor="Zhang, Yan-Na, Du, Wei &amp; Hua, Bao-Zhen" docDate="2020" docId="03802B15E57FBF14FF0E729AFBFEFDF0" docLanguage="en" docName="zootaxa.4718.3.6.pdf" docOrigin="Zootaxa 4718 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bittacus shaoguanensis Zhang &amp; Du &amp; Hua 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="386" masterDocId="FFB9536DE57BBF11FF997168FFD8FF94" masterDocTitle="Three new species of the genus Bittacus Latreille, 1805 (Mecoptera: Bittacidae), with a key to the species of Bittacidae in South China" masterLastPageNumber="390" masterPageNumber="381" pageNumber="385" updateTime="1732566459927" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="057A898CF997274ED8AD253BFA08C67A">Three new species of the genus Bittacus Latreille, 1805 (Mecoptera: Bittacidae), with a key to the species of Bittacidae in South China</mods:title>
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<mods:name id="1BD763F2B74D8D4D5D1FEB86E19112B2" type="personal">
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<mods:namePart id="C8AF8DD9B2783FC7015B3D1926552094">Zhang, Yan-Na</mods:namePart>
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<mods:namePart id="515A04ADD393B855D43687E0E6FB52EC">Du, Wei</mods:namePart>
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<mods:namePart id="9E08E7858DA0F20FAC8572AE5E704BB5">Hua, Bao-Zhen</mods:namePart>
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<mods:title id="75D76F2F71F7D5C140D6ABDF6FD8CE1C">Zootaxa</mods:title>
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<mods:date id="647B4DFB17E6B1746A36C03634A41AE1">2020</mods:date>
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<treatment id="03802B15E57FBF14FF0E729AFBFEFDF0" ID-GBIF-Taxon="161671027" LSID="urn:lsid:plazi:treatment:03802B15E57FBF14FF0E729AFBFEFDF0" httpUri="http://treatment.plazi.org/id/03802B15E57FBF14FF0E729AFBFEFDF0" lastPageId="5" lastPageNumber="386" pageId="4" pageNumber="385" scope_order="Mecoptera">
<subSubSection id="C333C988E57FBF15FF0E729AFE27FB98" box="[151,511,1009,1036]" pageId="4" pageNumber="385" type="nomenclature">
<paragraph id="8B969A03E57FBF15FF0E729AFE27FB98" blockId="4.[151,511,1009,1072]" box="[151,511,1009,1036]" pageId="4" pageNumber="385">
<heading id="D0DE2D6FE57FBF15FF0E729AFE27FB98" bold="true" box="[151,511,1009,1036]" fontSize="11" level="1" pageId="4" pageNumber="385" reason="1">
<emphasis id="B95D4611E57FBF15FF0E729AFE27FB98" bold="true" box="[151,511,1009,1036]" pageId="4" pageNumber="385">
<taxonomicName id="4C29E180E57FBF15FF0E729AFE6DFB98" ID-CoL="C93NM" authority="Zhang &amp; Du &amp; Hua, 2020" authorityName="Zhang &amp; Du &amp; Hua" authorityYear="2020" box="[151,437,1009,1036]" class="Insecta" family="Bittacidae" genus="Bittacus" kingdom="Animalia" order="Mecoptera" pageId="4" pageNumber="385" phylum="Arthropoda" rank="species" species="shaoguanensis" status="sp. nov.">
<emphasis id="B95D4611E57FBF15FF0E729AFE6DFB98" bold="true" box="[151,437,1009,1036]" italics="true" pageId="4" pageNumber="385">Bittacus shaoguanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26EFB6AE57FBF15FE24729AFE27FB98" box="[445,511,1010,1036]" pageId="4" pageNumber="385" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C333C988E57FBF15FF0E757DFECAFBA4" box="[151,274,1045,1072]" pageId="4" pageNumber="385" type="description">
<paragraph id="8B969A03E57FBF15FF0E757DFECAFBA4" blockId="4.[151,511,1009,1072]" box="[151,274,1045,1072]" pageId="4" pageNumber="385">
(
<figureCitation id="13128686E57FBF15FF06757DFED2FBA4" box="[159,266,1045,1072]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStartId-0="5.[151,250,1603,1628]" captionStartId-1="6.[151,250,1923,1948]" captionTargetBox-0="[151,1435,635,1583]" captionTargetBox-1="[151,1435,682,1892]" captionTargetId-0="figure@5.[151,1435,625,1583]" captionTargetId-1="figure@6.[151,1435,661,1904]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 4. Bittacus shaoguanensis sp. n. A. Male habitus in lateral view, holotype; B. articulation of basitarsus and tibia, to show two apical tibial spurs, holotype; C. tarsomeres IVV, paratype; D. right wings, paratype; E. head in frontal view, paratype; F. thorax in dorsal view, paratype. Abbreviations: see Fig. 2 caption. Scale bars: A = 5 mm; B, E and F = 0.5 mm; C = 0.2 mm; D = 2 mm." captionText-1="FIGURE 5. Bittacus shaoguanensis sp. n. A. Male terminalia in lateral view, holotype; B. male terminalia in dorsal view, holotype; C. female terminalia in lateral view, paratype; D. female terminalia in ventral view, paratype.Abbreviations: see Fig. 3 caption. Scale bars = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3602536" figureDoi-1="http://doi.org/10.5281/zenodo.3602538" httpUri-0="https://zenodo.org/record/3602536/files/figure.png" httpUri-1="https://zenodo.org/record/3602538/files/figure.png" pageId="4" pageNumber="385">Figs. 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C333C988E57FBF15FF0E7536FB5DFB0F" pageId="4" pageNumber="385" type="materials_examined">
<paragraph id="8B969A03E57FBF15FF0E7536FB5DFB0F" blockId="4.[151,1437,1117,2044]" pageId="4" pageNumber="385">
<emphasis id="B95D4611E57FBF15FF0E7536FE9CFBEC" bold="true" box="[151,324,1117,1144]" pageId="4" pageNumber="385">Type material.</emphasis>
<materialsCitation id="3B41905EE57FBF15FED37536FDE2FB0F" ID-GBIF-Occurrence="2545450325" collectingDate="2018-04-22" collectorName="Ji-Shen Wang" country="China" elevation="300" latitude="24.726389" location="Tsao Kok Wan" longLatPrecision="21" longitude="113.884995" municipality="Shaoguan" pageId="4" pageNumber="385" specimenCount="1" specimenCount-male="1" stateProvince="Guangdong" typeStatus="holotype">
<emphasis id="B95D4611E57FBF15FED37536FE66FBEC" bold="true" box="[330,446,1118,1144]" pageId="4" pageNumber="385">
<typeStatus id="549224A1E57FBF15FED37536FE61FBEC" box="[330,441,1118,1144]" pageId="4" pageNumber="385" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D2F518AE57FBF15FE5D7536FE03FBEC" box="[452,475,1118,1144]" pageId="4" pageNumber="385" type="male"></specimenCount>
,
<collectingCountry id="F33EDA93E57FBF15FE7F7536FD9BFBEC" box="[486,579,1118,1144]" name="China" pageId="4" pageNumber="385">CHINA</collectingCountry>
:
<collectingRegion id="49ED54E1E57FBF15FDD47535FD0BFBE3" box="[589,723,1117,1143]" country="China" name="Guangdong" pageId="4" pageNumber="385">Guangdong</collectingRegion>
,
<collectingMunicipality id="6BF20079E57FBF15FD457535FC95FBE3" box="[732,845,1117,1143]" pageId="4" pageNumber="385">Shaoguan</collectingMunicipality>
,
<location id="8EF6CCD8E57FBF15FCCE7536FC23FBEC" LSID="urn:lsid:plazi:treatment:03802B15E57FBF14FF0E729AFBFEFDF0:8EF6CCD8E57FBF15FCCE7536FC23FBEC" box="[855,1019,1117,1144]" country="China" latitude="24.726389" longLatPrecision="21" longitude="113.884995" municipality="Shaoguan" name="Tsao Kok Wan" pageId="4" pageNumber="385" stateProvince="Guangdong">Tsao Kok Wan</location>
(
<geoCoordinate id="EE1DFCC4E57FBF15FB907535FB4AFBEC" box="[1033,1170,1117,1144]" degrees="24" direction="north" minutes="43" orientation="latitude" pageId="4" pageNumber="385" precision="15" seconds="35" value="24.726389">24°4335″N</geoCoordinate>
,
<geoCoordinate id="EE1DFCC4E57FBF15FB057535FAF5FBEC" box="[1180,1325,1117,1144]" degrees="113" direction="east" minutes="53" orientation="longitude" pageId="4" pageNumber="385" precision="15" seconds="06" value="113.884995">113°5306″E</geoCoordinate>
, elev.
<quantity id="4CD137E6E57FBF15FAEB7536FF68FB08" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="385" unit="m" value="300.0">
<elevation id="00047D30E57FBF15FAEB7536FF68FB08" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="385" unit="m" value="300.0">300 m</elevation>
</quantity>
),
<date id="FF97BCC3E57FBF15FF5D75EAFEBBFB08" box="[196,355,1153,1180]" pageId="4" pageNumber="385" value="2018-04-22">
<collectingDate id="EFD3452BE57FBF15FF5D75EAFEBBFB08" box="[196,355,1153,1180]" pageId="4" pageNumber="385" value="2018-04-22">22 April 2018</collectingDate>
</date>
,
<collectorName id="26DCFFD5E57FBF15FEF775E9FDD1FB08" box="[366,521,1153,1180]" pageId="4" pageNumber="385">Ji-Shen Wang</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B41905EE57FBF15FDD975EAFB59FB0F" ID-GBIF-Occurrence="2545450330" box="[576,1153,1153,1180]" collectingDate="2018-04-22" collectorName="Ji-Shen Wang" country="China" elevation="300" latitude="24.726389" location="Tsao Kok Wan" longLatPrecision="21" longitude="113.884995" municipality="Shaoguan" pageId="4" pageNumber="385" specimenCount="22" specimenCount-female="8" specimenCount-male="14" stateProvince="Guangdong" typeStatus="paratype">
<emphasis id="B95D4611E57FBF15FDD975EAFD67FB08" bold="true" box="[576,703,1154,1180]" pageId="4" pageNumber="385">
<typeStatus id="549224A1E57FBF15FDD975EAFD62FB08" box="[576,698,1154,1180]" pageId="4" pageNumber="385" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D2F518AE57FBF15FD5F75EAFD21FB08" box="[710,761,1154,1180]" pageId="4" pageNumber="385" type="male">14♂</specimenCount>
,
<specimenCount id="9D2F518AE57FBF15FC9C75EAFCF1FB08" box="[773,809,1154,1180]" pageId="4" pageNumber="385" type="female">8♀</specimenCount>
, same data as for the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C333C988E57FBF15FF5E75CDFBEAFA9C" pageId="4" pageNumber="385" type="diagnosis">
<paragraph id="8B969A03E57FBF15FF5E75CDFBEAFA9C" blockId="4.[151,1437,1117,2044]" pageId="4" pageNumber="385">
<emphasis id="B95D4611E57FBF15FF5E75CDFE9AFB54" bold="true" box="[199,322,1189,1216]" pageId="4" pageNumber="385">Diagnosis.</emphasis>
The new species can be distinguished from its congeners by the following characters: 1) three brown single markings, one each at OM, ORs and Scv; pale brown clouding along apical crossveins, two middle streaks in a V-shape (
<figureCitation id="13128686E57FBF15FE8D7585FEB7FA9C" box="[276,367,1261,1288]" captionStart="FIGURE 4" captionStartId="5.[151,250,1603,1628]" captionTargetBox="[151,1435,635,1583]" captionTargetId="figure@5.[151,1435,625,1583]" captionTargetPageId="5" captionText="FIGURE 4. Bittacus shaoguanensis sp. n. A. Male habitus in lateral view, holotype; B. articulation of basitarsus and tibia, to show two apical tibial spurs, holotype; C. tarsomeres IVV, paratype; D. right wings, paratype; E. head in frontal view, paratype; F. thorax in dorsal view, paratype. Abbreviations: see Fig. 2 caption. Scale bars: A = 5 mm; B, E and F = 0.5 mm; C = 0.2 mm; D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3602536" httpUri="https://zenodo.org/record/3602536/files/figure.png" pageId="4" pageNumber="385">Fig. 4D</figureCitation>
); and 2) epandrial lobe sub-triangular in lateral view (
<figureCitation id="13128686E57FBF15FC527585FBFDFA9C" box="[971,1061,1261,1288]" captionStart="FIGURE 5" captionStartId="6.[151,250,1923,1948]" captionTargetBox="[151,1435,682,1892]" captionTargetId="figure@6.[151,1435,661,1904]" captionTargetPageId="6" captionText="FIGURE 5. Bittacus shaoguanensis sp. n. A. Male terminalia in lateral view, holotype; B. male terminalia in dorsal view, holotype; C. female terminalia in lateral view, paratype; D. female terminalia in ventral view, paratype.Abbreviations: see Fig. 3 caption. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3602538" httpUri="https://zenodo.org/record/3602538/files/figure.png" pageId="4" pageNumber="385">Fig. 5A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C333C988E57FBF15FF5E747AFC31FABF" box="[199,1001,1297,1324]" pageId="4" pageNumber="385" type="etymology">
<paragraph id="8B969A03E57FBF15FF5E747AFC31FABF" blockId="4.[151,1437,1117,2044]" box="[199,1001,1297,1324]" pageId="4" pageNumber="385">
<emphasis id="B95D4611E57FBF15FF5E747AFE96FAB8" bold="true" box="[199,334,1298,1324]" pageId="4" pageNumber="385">Etymology.</emphasis>
The new species is named after its
<typeStatus id="549224A1E57FBF15FD42747AFCD3FAB8" box="[731,779,1298,1324]" pageId="4" pageNumber="385">type</typeStatus>
locality, Shaoguan.
</paragraph>
</subSubSection>
<subSubSection id="C333C988E57FBF14FF5E745DFDCBFE1F" lastPageId="5" lastPageNumber="386" pageId="4" pageNumber="385" type="description">
<paragraph id="8B969A03E57FBF15FF5E745DFE76FA2F" blockId="4.[151,1437,1117,2044]" pageId="4" pageNumber="385">
<emphasis id="B95D4611E57FBF15FF5E745DFE46FAC4" bold="true" box="[199,414,1333,1360]" pageId="4" pageNumber="385">
Description.
<emphasis id="B95D4611E57FBF15FEF9745DFE46FAC4" bold="true" box="[352,414,1333,1360]" italics="true" pageId="4" pageNumber="385">Head</emphasis>
</emphasis>
(
<figureCitation id="13128686E57FBF15FE37745DFDDBFAC4" box="[430,515,1333,1360]" captionStart="FIGURE 4" captionStartId="5.[151,250,1603,1628]" captionTargetBox="[151,1435,635,1583]" captionTargetId="figure@5.[151,1435,625,1583]" captionTargetPageId="5" captionText="FIGURE 4. Bittacus shaoguanensis sp. n. A. Male habitus in lateral view, holotype; B. articulation of basitarsus and tibia, to show two apical tibial spurs, holotype; C. tarsomeres IVV, paratype; D. right wings, paratype; E. head in frontal view, paratype; F. thorax in dorsal view, paratype. Abbreviations: see Fig. 2 caption. Scale bars: A = 5 mm; B, E and F = 0.5 mm; C = 0.2 mm; D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3602536" httpUri="https://zenodo.org/record/3602536/files/figure.png" pageId="4" pageNumber="385">Fig. 4E</figureCitation>
). Vertex and frons yellowish; clypeus yellowish brown, darker than frons; labrum brown, darker basally and laterally; maxillary palp dark brown; fifth maxillary palpomere slightly shorter than the fourth. Ocellar triangle black. Antennae filiform and ciliated; scape and flagellum yellowish, pedicel pale brown; flagellum 17-segmented.
</paragraph>
<paragraph id="8B969A03E57FBF15FF5E74ADFEF2F907" blockId="4.[151,1437,1117,2044]" pageId="4" pageNumber="385">
<emphasis id="B95D4611E57FBF15FF5E74ADFEC5FA74" bold="true" box="[199,285,1477,1504]" italics="true" pageId="4" pageNumber="385">Thorax</emphasis>
(
<figureCitation id="13128686E57FBF15FEB774ADFE11FA74" box="[302,457,1477,1504]" captionStart="FIGURE 4" captionStartId="5.[151,250,1603,1628]" captionTargetBox="[151,1435,635,1583]" captionTargetId="figure@5.[151,1435,625,1583]" captionTargetPageId="5" captionText="FIGURE 4. Bittacus shaoguanensis sp. n. A. Male habitus in lateral view, holotype; B. articulation of basitarsus and tibia, to show two apical tibial spurs, holotype; C. tarsomeres IVV, paratype; D. right wings, paratype; E. head in frontal view, paratype; F. thorax in dorsal view, paratype. Abbreviations: see Fig. 2 caption. Scale bars: A = 5 mm; B, E and F = 0.5 mm; C = 0.2 mm; D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3602536" httpUri="https://zenodo.org/record/3602536/files/figure.png" pageId="4" pageNumber="385">Fig. 4B, C, F</figureCitation>
). Pronotum blackish brown, with two long setae along anterior and posterior margin, respectively. Anterior two-thirds of mesonotum blackish brown, remaining part and metanotum unevenly brown. Pleura unevenly pale brown. Legs yellowish brown, except for coxae pale brown, femora and tibiae darker apically; tibia with two apical spurs, the longer one about 1.25 times as long as the shorter one in hind leg. Tarsomere IV bearing 2427 prominent black ventral teeth, with two setae on each side subbasally; tarsomere V with 2629 ventral teeth.
</paragraph>
<paragraph id="8B969A03E57FBF15FF5E77F6FCD9F8D3" blockId="4.[151,1437,1117,2044]" pageId="4" pageNumber="385">
<emphasis id="B95D4611E57FBF15FF5E77F6FED7F92C" bold="true" box="[199,271,1694,1720]" italics="true" pageId="4" pageNumber="385">Wings</emphasis>
(
<figureCitation id="13128686E57FBF15FE8477F5FEACF92C" box="[285,372,1693,1720]" captionStart="FIGURE 4" captionStartId="5.[151,250,1603,1628]" captionTargetBox="[151,1435,635,1583]" captionTargetId="figure@5.[151,1435,625,1583]" captionTargetPageId="5" captionText="FIGURE 4. Bittacus shaoguanensis sp. n. A. Male habitus in lateral view, holotype; B. articulation of basitarsus and tibia, to show two apical tibial spurs, holotype; C. tarsomeres IVV, paratype; D. right wings, paratype; E. head in frontal view, paratype; F. thorax in dorsal view, paratype. Abbreviations: see Fig. 2 caption. Scale bars: A = 5 mm; B, E and F = 0.5 mm; C = 0.2 mm; D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3602536" httpUri="https://zenodo.org/record/3602536/files/figure.png" pageId="4" pageNumber="385">Fig. 4D</figureCitation>
). Wing membrane hyaline, tinged with yellowish brown. Forewing length
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, width
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. Pterostigma brown; two Pcv; Av absent; three distinct tawny brown markings, one each at OM, ORs, and Scv; pale brown clouding along apical crossveins, two middle streaks in a V-shape; apical margin slightly infuscate; thyridium at FM conspicuous; Sc ending beyond the level of FRs; 1A ending before FM. Hindwing length
<quantity id="4CD137E6E57FBF15FF0E7646FEE4F8DC" box="[151,316,1838,1864]" metricMagnitude="-2" metricUnit="m" metricValue="1.7799999999999998" metricValueMax="1.8399999999999999" metricValueMin="1.72" pageId="4" pageNumber="385" unit="mm" value="17.799999999999997" valueMax="18.4" valueMin="17.2">17.218.4 mm</quantity>
, width
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, similar to forewing.
</paragraph>
<paragraph id="8B969A03E57FBF14FF5E7639FD44FF6F" blockId="4.[151,1437,1117,2044]" lastBlockId="5.[151,1437,153,612]" lastPageId="5" lastPageNumber="386" pageId="4" pageNumber="385">
<emphasis id="B95D4611E57FBF15FF5E7639FE4DF8F8" bold="true" box="[199,405,1873,1900]" italics="true" pageId="4" pageNumber="385">Abdomen of male</emphasis>
(
<figureCitation id="13128686E57FBF15FE3C7639FDD2F8F8" box="[421,522,1873,1900]" captionStart="FIGURE 4" captionStartId="5.[151,250,1603,1628]" captionTargetBox="[151,1435,635,1583]" captionTargetId="figure@5.[151,1435,625,1583]" captionTargetPageId="5" captionText="FIGURE 4. Bittacus shaoguanensis sp. n. A. Male habitus in lateral view, holotype; B. articulation of basitarsus and tibia, to show two apical tibial spurs, holotype; C. tarsomeres IVV, paratype; D. right wings, paratype; E. head in frontal view, paratype; F. thorax in dorsal view, paratype. Abbreviations: see Fig. 2 caption. Scale bars: A = 5 mm; B, E and F = 0.5 mm; C = 0.2 mm; D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3602536" httpUri="https://zenodo.org/record/3602536/files/figure.png" pageId="4" pageNumber="385">Figs. 4A</figureCitation>
,
<figureCitation id="13128686E57FBF15FD8F7639FDFBF8F8" box="[534,547,1873,1900]" captionStart="FIGURE 5" captionStartId="6.[151,250,1923,1948]" captionTargetBox="[151,1435,682,1892]" captionTargetId="figure@6.[151,1435,661,1904]" captionTargetPageId="6" captionText="FIGURE 5. Bittacus shaoguanensis sp. n. A. Male terminalia in lateral view, holotype; B. male terminalia in dorsal view, holotype; C. female terminalia in lateral view, paratype; D. female terminalia in ventral view, paratype.Abbreviations: see Fig. 3 caption. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3602538" httpUri="https://zenodo.org/record/3602538/files/figure.png" pageId="4" pageNumber="385">5</figureCitation>
AB). Terga IIVI yellowish brown, each with a narrow black antecosta; terga VIIVIII blackish brown; tergum VIII emarginate distally in a V-shape. Epandrial lobe blackish brown, with distal margin pale, shorter than gonocoxites, sub-triangular in lateral view, truncated apically with a series of short black spines on inner side. Upper branch of proctiger yellowish brown, boot-shaped, bearing a bundle of yellowish setae distally; lower branch of proctiger broad basally and tapering toward apex, curved downward and shorter than up- per branch. Cerci pale, slightly shorter than half length of gonocoxites. Gonocoxites blackish brown, separated by a U-shaped membranous area caudally; gonostylus short and blunt distally, with a process on inner side. Aedeagus broad basally; penisfilum elongate and coiled.
</paragraph>
<paragraph id="8B969A03E57EBF14FF5E706CFDCBFE1F" blockId="5.[151,1437,153,612]" pageId="5" pageNumber="386">
<emphasis id="B95D4611E57EBF14FF5E706CFD9AFE8B" bold="true" box="[199,578,260,287]" pageId="5" pageNumber="386">
<emphasis id="B95D4611E57EBF14FF5E706CFE71FE8B" bold="true" box="[199,425,260,287]" italics="true" pageId="5" pageNumber="386">Abdomen of female</emphasis>
(
<figureCitation id="13128686E57EBF14FE21706CFE24FE8B" box="[440,508,260,287]" captionStart="FIGURE 5" captionStartId="6.[151,250,1923,1948]" captionTargetBox="[151,1435,682,1892]" captionTargetId="figure@6.[151,1435,661,1904]" captionTargetPageId="6" captionText="FIGURE 5. Bittacus shaoguanensis sp. n. A. Male terminalia in lateral view, holotype; B. male terminalia in dorsal view, holotype; C. female terminalia in lateral view, paratype; D. female terminalia in ventral view, paratype.Abbreviations: see Fig. 3 caption. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3602538" httpUri="https://zenodo.org/record/3602538/files/figure.png" pageId="5" pageNumber="386">Fig. 5</figureCitation>
CD).
</emphasis>
Terga IIIX unevenly dark brown, each with a narrow black antecosta. Subgenital plate yellowish brown, separated medially by a bullet-shaped membranous area, apically with a series of black setae. Tergum X yellowish brown, extending ventrally. Anal plates consistent in length, blunt apically. Cerci slightly longer than subanal plate.
</paragraph>
</subSubSection>
<subSubSection id="C333C988E57EBF14FF5E70FCFD96FE24" box="[199,590,404,432]" pageId="5" pageNumber="386" type="distribution">
<paragraph id="8B969A03E57EBF14FF5E70FCFD96FE24" blockId="5.[151,1437,153,612]" box="[199,590,404,432]" pageId="5" pageNumber="386">
<emphasis id="B95D4611E57EBF14FF5E70FCFEB9FE3B" bold="true" box="[199,353,404,431]" pageId="5" pageNumber="386">Distribution.</emphasis>
<collectingCountry id="F33EDA93E57EBF14FEFE70FDFE72FE3B" box="[359,426,405,431]" name="China" pageId="5" pageNumber="386">China</collectingCountry>
(
<collectingRegion id="49ED54E1E57EBF14FE2070FDFD9AFE24" box="[441,578,405,432]" country="China" name="Guangdong" pageId="5" pageNumber="386">Guangdong</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C333C988E57EBF14FF5E70D1FBFEFDF0" pageId="5" pageNumber="386" type="discussion">
<paragraph id="8B969A03E57EBF14FF5E70D1FBFEFDF0" blockId="5.[151,1437,153,612]" pageId="5" pageNumber="386">
<emphasis id="B95D4611E57EBF14FF5E70D1FEE3FE47" bold="true" box="[199,315,441,467]" pageId="5" pageNumber="386">Remarks.</emphasis>
<taxonomicName id="4C29E180E57EBF14FEDA70D2FD94FE47" authority="Zhang &amp; Du &amp; Hua, 2020" authorityName="Zhang &amp; Du &amp; Hua" authorityYear="2020" box="[323,588,441,467]" class="Insecta" family="Bittacidae" genus="Bittacus" kingdom="Animalia" order="Mecoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="shaoguanensis" status="sp. nov.">
<emphasis id="B95D4611E57EBF14FEDA70D2FD94FE47" box="[323,588,441,467]" italics="true" pageId="5" pageNumber="386">Bittacus shaoguanensis</emphasis>
</taxonomicName>
<emphasis id="B95D4611E57EBF14FDCA70D1FD4CFE47" bold="true" box="[595,660,441,467]" pageId="5" pageNumber="386">
<taxonomicNameLabel id="A26EFB6AE57EBF14FDCA70D1FD4CFE47" box="[595,660,441,467]" pageId="5" pageNumber="386" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C29E180E57EBF14FC8A70D2FBADFE47" authority="Hua &amp; Chou, 1988" authorityName="Hua &amp; Chou" authorityYear="1988" box="[787,1141,441,467]" class="Insecta" family="Bittacidae" genus="Bittacus" kingdom="Animalia" order="Mecoptera" pageId="5" pageNumber="386" phylum="Arthropoda" rank="species" species="strigatus">
<emphasis id="B95D4611E57EBF14FC8A70D2FC4DFE47" box="[787,917,442,467]" italics="true" pageId="5" pageNumber="386">B. strigatus</emphasis>
Hua &amp; Chou,
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</taxonomicName>
in appearance, but can be readily separated from the latter by the following characters: 1) femora yellowish brown, darker apically (
<emphasis id="B95D4611E57EBF14FADB70B5FA8EFE63" box="[1346,1366,477,503]" italics="true" pageId="5" pageNumber="386">cf</emphasis>
. anterior 2/3 of fore femora blackish brown); 2) epandrial lobe shorter than gonocoxites, sub-triangular in lateral view (
<emphasis id="B95D4611E57EBF14FF38734DFF6DFDAB" box="[161,181,549,575]" italics="true" pageId="5" pageNumber="386">cf</emphasis>
. epandrial lobe longer than gonocoxites, curved downward); and 3) upper branch of proctiger yellowish brown, boot-shaped apically (
<emphasis id="B95D4611E57EBF14FE097321FE74FDF7" box="[400,428,585,611]" italics="true" pageId="5" pageNumber="386">cf.</emphasis>
upper branch of proctiger strongly sclerotized, digitate).
</paragraph>
</subSubSection>
</treatment>
</document>