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<document id="20B6C56F75C54F4E435C36438714AD43" ID-DOI="10.11646/zootaxa.4742.1.5" ID-GBIF-Dataset="d9aaf71d-e347-4fc3-b5fd-cae0989cac05" ID-ISSN="1175-5326" ID-Zenodo-Dep="3674493" ID-ZooBank="3016F563-FDAC-44FD-B6A5-44E1E8DFCFB4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582096979764" checkinUser="plazi" docAuthor="Arimoto, Kôichi &amp; Suzuki, Wataru" docDate="2020" docId="FC3C87A4327FB31783CEBA3CBB7BFAE1" docLanguage="en" docName="zootaxa.4742.1.5.pdf" docOrigin="Zootaxa 4742 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limonius sapphirus Arimoto &amp; Suzuki 2020, sp. n." docType="treatment" docVersion="6" lastPageNumber="96" masterDocId="0005FFDC327CB3108359BB4DBF09FF9A" masterDocTitle="First record and two new species of the genus Limonius Eschscholtz (Coleoptera Elateridae, Dendrometrinae) from Taiwan" masterLastPageNumber="104" masterPageNumber="89" pageNumber="92" updateTime="1698719267030" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="9E8ADE744A95C6DC6B3E9239FC998274" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D13DF5170F4819090AD44880A89E639C">
<mods:title id="95F92650C5B3FA7B369C4C0AF32B5759">First record and two new species of the genus Limonius Eschscholtz (Coleoptera Elateridae, Dendrometrinae) from Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="E6946A3E4C2DE2725D68B4F8A79A2592" type="personal">
<mods:role id="011877A7BB1079622796D7CFCE9512DD">
<mods:roleTerm id="227307ACFEFBD3999AB09B0B324620BB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC8A4FA9CD0305ADC2241D22038D4986">Arimoto, Kôichi</mods:namePart>
</mods:name>
<mods:name id="2B0AA145BB5BE493AD3C3B2D58637727" type="personal">
<mods:role id="2DAC432E5E224F9DC9838EB664B403DA">
<mods:roleTerm id="D9CABF13C55636D1803C077B605C6C9F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C87C13C89E27AC48849A5C17060B7D21">Suzuki, Wataru</mods:namePart>
</mods:name>
<mods:typeOfResource id="019A74AF8721B3806FDD2AE37843A52A">text</mods:typeOfResource>
<mods:relatedItem id="075F81C9DBE82DE11F8BCA8622B657F0" type="host">
<mods:titleInfo id="71812E175A8437E2C95677F8DFA44DD6">
<mods:title id="79F3DC24896B8672DF80C600421C416E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="38C792E1B2D4D8B5AFB2F067672748C5">
<mods:date id="CFBD8F339EFF5E241DACFB6719503C65">2020</mods:date>
<mods:detail id="7189E0D2F477059CFA4156146308B57B" type="pubDate">
<mods:number id="03DE33634A7806083E770CC9E5D90D16">2020-02-19</mods:number>
</mods:detail>
<mods:detail id="16BF5EDF870C9588E1CB76D26D6A6E75" type="volume">
<mods:number id="0EB0DCC31778B06619EBB0D63EC67ECD">4742</mods:number>
</mods:detail>
<mods:detail id="AAB8C98BEF1A0529916D23CC956F5502" type="issue">
<mods:number id="825DAC77FDCA97908A4ACE70991B7C3E">1</mods:number>
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<mods:start id="B769E51002D7BC3015DCE565FC8A33A1">89</mods:start>
<mods:end id="4EEA469759920FB63845BD8D009AC512">104</mods:end>
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<mods:classification id="6E2D02C65B29242BE8F7554CA58FBF96">journal article</mods:classification>
<mods:identifier id="6D21E4AB0B0542DD311C06BB4AD8D99B" type="DOI">10.11646/zootaxa.4742.1.5</mods:identifier>
<mods:identifier id="0A4B98953909C2FA952D538191CE0DCC" type="GBIF-Dataset">d9aaf71d-e347-4fc3-b5fd-cae0989cac05</mods:identifier>
<mods:identifier id="6634C141A876712920F8C53468FA7D50" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CD12F346B528748A4ABCA14C33D5FC6B" type="Zenodo-Dep">3674493</mods:identifier>
<mods:identifier id="00395D035FC04918CB6336F4C51F2E91" type="ZooBank">3016F563-FDAC-44FD-B6A5-44E1E8DFCFB4</mods:identifier>
</mods:mods>
<treatment id="FC3C87A4327FB31783CEBA3CBB7BFAE1" ID-DOI="http://doi.org/10.5281/zenodo.3681231" ID-GBIF-Taxon="162163997" ID-Zenodo-Dep="3681231" LSID="urn:lsid:plazi:treatment:FC3C87A4327FB31783CEBA3CBB7BFAE1" httpUri="http://treatment.plazi.org/id/FC3C87A4327FB31783CEBA3CBB7BFAE1" lastPageId="7" lastPageNumber="96" pageId="3" pageNumber="92">
<subSubSection id="3C8F6539327FB31383CEBA3CBDCEFE11" box="[151,711,369,396]" pageId="3" pageNumber="92" type="nomenclature">
<paragraph id="742A36B2327FB31383CEBA3CBDCEFE11" blockId="3.[151,711,369,431]" box="[151,711,369,396]" pageId="3" pageNumber="92">
<heading id="2F6281DE327FB31383CEBA3CBDCEFE11" bold="true" box="[151,711,369,396]" fontSize="11" level="1" pageId="3" pageNumber="92" reason="1">
<emphasis id="46E1EAA0327FB31383CEBA3CBDCEFE11" bold="true" box="[151,711,369,396]" pageId="3" pageNumber="92">
<taxonomicName id="B3954D31327FB31383CEBA3CBD71FE11" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[151,632,369,396]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="92" phylum="Arthropoda" rank="species" species="sapphirus" status="sp. nov.">
<emphasis id="46E1EAA0327FB31383CEBA3CBE80FE16" bold="true" box="[151,393,369,396]" italics="true" pageId="3" pageNumber="92">Limonius sapphirus</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
,
<taxonomicNameLabel id="5DD257DB327FB31381DCBA3CBDCEFE11" box="[645,711,369,395]" pageId="3" pageNumber="92" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327FB31383CEBAD8BE68FE35" box="[151,353,405,431]" pageId="3" pageNumber="92" type="description">
<paragraph id="742A36B2327FB31383CEBAD8BE68FE35" blockId="3.[151,711,369,431]" box="[151,353,405,431]" pageId="3" pageNumber="92">
(
<figureCitation id="ECAE2A37327FB31383C6BAD8BE20FE35" box="[159,297,405,431]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[151,250,1819,1844]" captionStartId-1="5.[151,250,798,823]" captionStartId-2="5.[151,250,1975,2000]" captionStartId-3="6.[151,250,1473,1498]" captionTargetBox-0="[164,1423,187,1784]" captionTargetBox-1="[189,1399,187,787]" captionTargetBox-2="[189,1398,917,1950]" captionTargetBox-3="[182,1436,181,1448]" captionTargetId-0="figure@4.[164,1423,179,1792]" captionTargetId-1="figure@5.[189,1399,181,794]" captionTargetId-2="figure@5.[189,1398,917,1950]" captionTargetId-3="figure@6.[151,1436,181,1448]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." captionText-1="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." captionText-2="FIGURE 3. Limonius sapphirus Arimoto &amp; Suzuki sp. n., male, holotype.A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." captionText-3="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi-0="http://doi.org/10.5281/zenodo.3674495" figureDoi-1="http://doi.org/10.5281/zenodo.3674497" figureDoi-2="http://doi.org/10.5281/zenodo.3674499" figureDoi-3="http://doi.org/10.5281/zenodo.3674501" httpUri-0="https://zenodo.org/record/3674495/files/figure.png" httpUri-1="https://zenodo.org/record/3674497/files/figure.png" httpUri-2="https://zenodo.org/record/3674499/files/figure.png" httpUri-3="https://zenodo.org/record/3674501/files/figure.png" pageId="3" pageNumber="92">Figures 1–4</figureCitation>
,
<figureCitation id="ECAE2A37327FB313826CBAD8BE50FE35" box="[309,345,405,431]" captionStart="FIGURE 7" captionStartId="9.[151,250,1830,1855]" captionTargetBox="[158,1426,1166,1818]" captionTargetId="figure@9.[151,1436,1159,1826]" captionTargetPageId="9" captionText="FIGURE 7. Limonius spp., male, holotype, apical part of paramere A: Limonius sapphirus Arimoto &amp; Suzuki sp. n.; B: Limonius onyx Arimoto &amp; Suzuki sp. n." figureDoi="http://doi.org/10.5281/zenodo.3674507" httpUri="https://zenodo.org/record/3674507/files/figure.png" pageId="3" pageNumber="92">7A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327FB31383CEBA90BB43FE6D" box="[151,1098,477,503]" pageId="3" pageNumber="92" type="etymology">
<paragraph id="742A36B2327FB31383CEBA90BB43FE6D" blockId="3.[151,1437,477,2016]" box="[151,1098,477,503]" pageId="3" pageNumber="92">
<emphasis id="46E1EAA0327FB31383CEBA90BE17FE6D" bold="true" box="[151,286,477,503]" pageId="3" pageNumber="92">Etymology.</emphasis>
In reference to the body shining in blue and green like sapphire jewelry.
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327FB313839EB94CBA69FDA5" pageId="3" pageNumber="92" type="materials_examined">
<paragraph id="742A36B2327FB313839EB94CBA69FDA5" blockId="3.[151,1437,477,2016]" pageId="3" pageNumber="92">
<emphasis id="46E1EAA0327FB313839EB94CBE71FD81" bold="true" box="[199,376,512,539]" pageId="3" pageNumber="92">Type material.</emphasis>
<materialsCitation id="C4FD3CEF327FB31382DBB94CBE9BFDA5" ID-GBIF-Occurrence="2566071271" collectingDate="1978-05-01" collectionCode="LSA" collectorName="Toshio Kobayashi" country="Taiwan" county="City" location="Male" municipality="Mt. Lala" pageId="3" pageNumber="92" specimenCode="LSA001" specimenCount="1" stateProvince="Taoyuan" typeStatus="holotype">
<emphasis id="46E1EAA0327FB31382DBB94CBEE6FD81" bold="true" box="[386,495,513,539]" pageId="3" pageNumber="92">
<typeStatus id="AB2E8810327FB31382DBB94CBEE6FD81" box="[386,495,513,539]" pageId="3" pageNumber="92" type="holotype">Holotype</typeStatus>
</emphasis>
.
<location id="714A6069327FB3138159B94CBD33FD81" LSID="urn:lsid:plazi:treatment:FC3C87A4327FB31783CEBA3CBB7BFAE1:714A6069327FB3138159B94CBD33FD81" box="[512,570,513,539]" country="Taiwan" county="City" municipality="Mt. Lala" name="Male" pageId="3" pageNumber="92" stateProvince="Taoyuan">Male</location>
(
<specimenCode id="24339EC9327FB3138112B94CBDA6FD81" box="[587,687,513,539]" collectionCode="LSA" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13533" name="Lytham St. Annes Public Library" pageId="3" pageNumber="92" type="Herbarium">LSA001</specimenCode>
), near
<collectingMunicipality id="944EACC8327FB3138059B94CBC6CFD81" box="[768,869,513,539]" pageId="3" pageNumber="92">Mt. Lala</collectingMunicipality>
,
<collectingRegion id="B651F850327FB313802AB94CBB04FD81" box="[883,1037,513,539]" country="Taiwan" name="Taoyuan" pageId="3" pageNumber="92">Taoyuan City</collectingRegion>
to
<collectingRegion id="B651F850327FB313876EB94CBB74FD81" box="[1079,1149,513,539]" country="Taiwan" name="Taipei" pageId="3" pageNumber="92">Taipei</collectingRegion>
<collectingCounty id="9D4B4E3E327FB31387DFB94CBBBEFD81" box="[1158,1207,513,539]" pageId="3" pageNumber="92">City</collectingCounty>
,
<collectingCountry id="0C827622327FB313879CB94CBA13FD81" box="[1221,1306,513,539]" name="Taiwan" pageId="3" pageNumber="92">Taiwan</collectingCountry>
,
<date id="002B1072327FB3138671B94CBA91FD81" box="[1320,1432,513,539]" pageId="3" pageNumber="92" value="1978-05-01">
<collectingDate id="106FE99A327FB3138671B94CBA91FD81" box="[1320,1432,513,539]" pageId="3" pageNumber="92" value="1978-05-01">1 V 1978</collectingDate>
</date>
,
<collectorName id="D9605364327FB31383CEB968BE6BFDA5" box="[151,354,549,575]" pageId="3" pageNumber="92">Toshio Kobayashi</collectorName>
leg.
</materialsCitation>
<materialsCitation id="C4FD3CEF327FB31382C0B968BB0BFDA5" ID-GBIF-Occurrence="2566071261" box="[409,1026,549,575]" collectionCode="LSA" country="Taiwan" location="Mt. Lala" pageId="3" pageNumber="92" specimenCode="LSA002" specimenCount="1" typeStatus="paratype">
<emphasis id="46E1EAA0327FB31382C0B968BD0FFDA5" bold="true" box="[409,518,549,575]" pageId="3" pageNumber="92">
<typeStatus id="AB2E8810327FB31382C0B968BD0FFDA5" box="[409,518,549,575]" pageId="3" pageNumber="92" type="paratype">Paratype</typeStatus>
</emphasis>
. Female (
<specimenCode id="24339EC9327FB313812CB968BDD0FDA5" box="[629,729,549,575]" collectionCode="LSA" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13533" name="Lytham St. Annes Public Library" pageId="3" pageNumber="92" type="Herbarium">LSA002</specimenCode>
),
<location id="714A6069327FB31381B4B968BC46FDA5" LSID="urn:lsid:plazi:treatment:FC3C87A4327FB31783CEBA3CBB7BFAE1:714A6069327FB31381B4B968BC46FDA5" box="[749,847,549,575]" country="Taiwan" name="Mt. Lala" pageId="3" pageNumber="92">Mt. Lala</location>
,
<collectingCountry id="0C827622327FB3138002B968BCB9FDA5" box="[859,944,549,575]" name="Taiwan" pageId="3" pageNumber="92">Taiwan</collectingCountry>
,
<date id="002B1072327FB31380E2B968BB0BFDA5" box="[955,1026,549,575]" pageId="3" pageNumber="92" value="1980-05-20" valueMax="1980-05-24" valueMin="1980-05-20">20–24</date>
</materialsCitation>
<materialsCitation id="C4FD3CEF327FB3138750B968BA69FDA5" ID-GBIF-Occurrence="2566071264" box="[1033,1376,549,575]" collectionCode="V" collectorName="Hiroshi Makihara" country="Taiwan" pageId="3" pageNumber="92" specimenCount="1" typeStatus="paratype">
<collectionCode id="1284AE77327FB3138750B968BB14FDA5" box="[1033,1053,549,575]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="92" type="Museum">V</collectionCode>
1980,
<collectorName id="D9605364327FB3138733B968BA39FDA5" box="[1130,1328,549,575]" pageId="3" pageNumber="92">Hiroshi Makihara</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327FB313839EB905BB41FC3D" pageId="3" pageNumber="92" type="diagnosis">
<paragraph id="742A36B2327FB313839EB905BB41FC3D" blockId="3.[151,1437,477,2016]" pageId="3" pageNumber="92">
<emphasis id="46E1EAA0327FB313839EB905BE4BFDF9" bold="true" box="[199,322,584,611]" pageId="3" pageNumber="92">Diagnosis.</emphasis>
Body surface with metallic luster; elytra interspaces between punctures larger than puncture diameter. Body light blue, but elytra darker in male; body light blue, but head and prothorax light green in female. Frons in profile weakly depressed apically; supra-antennal carina in dorsal view broadly rounded, but almost straight apically. Antennae extending beyond the posterior apices of pronotum by apical three antennomeres in male, by apical antennomere in female; antennomere III slightly longer than II; IV distinctly longer than wide, longer than II–III combined, longer than V. Prothorax widening posteriad; lateral carina of prothorax weakly prominent just behind anterior angles. Anterior lobe of prosternum in profile weakly inclined. Mesial edge of hypomeron carinate anterolaterad. Prosternal spine with subapical tooth; sides at ventral view roundly expanded and anteriorly strongly prominent. Scutellum almost as long as wide or slightly shorter than wide, strongly convex; posterior margin broadly rounded, but apex almost transverse. Mesepisternum not reaching mesocoxal cavity.
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327FB317839EB8FCBE63FACD" lastPageId="7" lastPageNumber="96" pageId="3" pageNumber="92" type="description">
<paragraph id="742A36B2327FB313839EB8FCBCD1FB89" blockId="3.[151,1437,477,2016]" pageId="3" pageNumber="92">
<emphasis id="46E1EAA0327FB313839EB8FCBEC0FC51" bold="true" box="[199,457,945,971]" pageId="3" pageNumber="92">Measurements. Male.</emphasis>
BL: 8.35, BW: 2.39,
<collectingRegion id="B651F850327FB31381E7B8FCBDFFFC51" box="[702,758,945,971]" country="Japan" name="Mie" pageId="3" pageNumber="92">MIE</collectingRegion>
: 0.90, MAE: 1.24, OI: 139, PL: 2.30, PML: 2.18, PW: 2.07, PI: 111, EL: 5.44, EW: 2.39, EI: 228, BI: 237.
<emphasis id="46E1EAA0327FB31381F0B898BC0EFC75" bold="true" box="[681,775,981,1007]" pageId="3" pageNumber="92">Female.</emphasis>
BL: 9.35, BW: 2.87,
<collectingRegion id="B651F850327FB31380A7B898BB3FFC75" box="[1022,1078,981,1007]" country="Japan" name="Mie" pageId="3" pageNumber="92">MIE</collectingRegion>
: 1.00, MAE: 1.35, OI: 135, PL: 2.60, PML: 2.50; PW: 2.34, PI: 111, EL: 6.09, EW: 2.87, EI: 212, BI: 234.
</paragraph>
<paragraph id="742A36B2327FB313839EBF51BD5FFA05" blockId="3.[151,1437,477,2016]" pageId="3" pageNumber="92">
<emphasis id="46E1EAA0327FB313839EBF51BE50FBAD" bold="true" box="[199,345,1052,1079]" pageId="3" pageNumber="92">Description.</emphasis>
Body surface generally with metallic luster and smooth, but ventral side of prosternal spine with micro-structure (
<figureCitation id="ECAE2A37327FB313820DBF0CBEA7FBC1" box="[340,430,1089,1115]" captionStart="FIGURE 2" captionStartId="5.[151,250,798,823]" captionTargetBox="[189,1399,187,787]" captionTargetId="figure@5.[189,1399,181,794]" captionTargetPageId="5" captionText="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="3" pageNumber="92">Fig. 2A</figureCitation>
), elytra partly rough; interspaces between punctures generally larger than puncture diameter, but head depression and scutellum interspaces between punctures smaller than puncture diameter (
<figureCitation id="ECAE2A37327FB313866BBF28BA9EFBE5" box="[1330,1431,1125,1151]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Figs. 1D</figureCitation>
,
<figureCitation id="ECAE2A37327FB31383CEBFC4BFD0FB39" box="[151,217,1161,1187]" captionStart="FIGURE 2" captionStartId="5.[151,250,798,823]" captionTargetBox="[189,1399,187,787]" captionTargetId="figure@5.[189,1399,181,794]" captionTargetPageId="5" captionText="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="3" pageNumber="92">2B, C</figureCitation>
). Color.
<emphasis id="46E1EAA0327FB3138262BFC4BE76FB39" bold="true" box="[315,383,1161,1187]" pageId="3" pageNumber="92">Male.</emphasis>
Body light blue, but elytra darker (
<figureCitation id="ECAE2A37327FB3138051BFC4BC88FB39" box="[776,897,1161,1187]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1A, C</figureCitation>
).
<emphasis id="46E1EAA0327FB31380CCBFC4BCFAFB39" bold="true" box="[917,1011,1161,1187]" pageId="3" pageNumber="92">Female.</emphasis>
Body light blue, but head and prothorax light green (
<figureCitation id="ECAE2A37327FB313821EBFE0BE95FB5D" box="[327,412,1197,1223]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1B</figureCitation>
). Labrum black. Antennae black, but external part of antennomeres IV–XI brown. Mandible black, but apical and internal parts orange. Maxilally palpomeres black, with brown tinge. Anterior edges of head and pronotum, mesial margin and posterior part of hypomeron, anterior lobe of prosternum, prosternal spine, mesothorax, posterior margins of each abdominal segment black. Scutellum black with blue tinge. Legs black, partly orange; claws orange. Tergites and sternites VIII‒X and aedeagus yellow-brown. Pubescence. Body covered with intermixed white and orange setae dorsally, with white setae ventrally. Antennae with yellow to brown setae. Setae of legs white becoming yellow apically.
</paragraph>
<paragraph id="742A36B2327FB313839EBEE4BA5AF925" blockId="3.[151,1437,477,2016]" pageId="3" pageNumber="92">
Head depressed longitudinally medially; the depression becoming narrow and shallow posteriad (
<figureCitation id="ECAE2A37327FB313864DBEE4BA84FA59" box="[1300,1421,1449,1475]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1D, E</figureCitation>
). Frons in profile weakly depressed apically (
<figureCitation id="ECAE2A37327FB3138122BE80BDC6FA7D" box="[635,719,1485,1511]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1F</figureCitation>
: green line); supra-antennal carina in dorsal view broadly round- ed, but almost transverse apically (
<figureCitation id="ECAE2A37327FB3138144BEBCBD7CF991" box="[541,629,1521,1547]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1D</figureCitation>
), in profile strongly prominent anteriad (
<figureCitation id="ECAE2A37327FB3138718BEBCBB9FF991" box="[1089,1174,1521,1547]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1F</figureCitation>
: arrow). Antennomeres I and IV–XI longer than wide; II globular, shortest; III obconical, slightly longer than II; IV–X serrate; IV longer than II–III combined, longer than V.
<emphasis id="46E1EAA0327FB3138165BD74BD89F9C9" bold="true" box="[572,640,1593,1619]" pageId="3" pageNumber="92">Male.</emphasis>
Antennae extending beyond pronotum posterior lateral apices by apical three antennomeres (
<figureCitation id="ECAE2A37327FB31382F2BD10BD0CF9ED" box="[427,517,1629,1655]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1A</figureCitation>
); antennomere IV 1.31 x as long as wide, 1.98 x as long as III; XI 2.70 x as long as wide.
<emphasis id="46E1EAA0327FB31383A4BDCCBE52F901" bold="true" box="[253,347,1665,1691]" pageId="3" pageNumber="92">Female.</emphasis>
Antennae extending beyond pronotum posterior lateral apices by apical antennomere (
<figureCitation id="ECAE2A37327FB313866CBDCCBA84F901" box="[1333,1421,1665,1691]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1B</figureCitation>
); antennomere IV 1.19 x as long as wide, 1.76 x as long as III; XI 2.01 x as long as wide. Mandible bidentate.
</paragraph>
<paragraph id="742A36B2327FB317839EBD84BE3AFF4D" blockId="3.[151,1437,477,2016]" lastBlockId="7.[151,1437,153,1403]" lastPageId="7" lastPageNumber="96" pageId="3" pageNumber="92">
Prothorax trapezoidal, longer than wide, widening posteriad; side weakly rounded; lateral carina extending from anterior angles to hind angles of prothorax, weakly prominent just behind anterior angles (
<figureCitation id="ECAE2A37327FB31387BDBDA0BA34F89D" box="[1252,1341,1773,1799]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1E</figureCitation>
: arrow). Pronotum posteriorly with a shallow median longitudinal depression; hind angles broad, rounded, unicarinate; the carina short, extending to the base. Hypomeron: mesial edge with elevated impunctate ridge next to pronotosternal suture, carinate anterolaterad (
<figureCitation id="ECAE2A37327FB31382B5BC14BD4FF8E9" box="[492,582,1881,1907]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1G</figureCitation>
: black arrow); posterior margin without large emargination, weakly sinuate. Anterior lobe of prosternum exceeding anterior angles of prothorax (
<figureCitation id="ECAE2A37327FB31380EABC30BB05F80D" box="[947,1036,1917,1943]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1C</figureCitation>
), in profile weakly inclined (at 15 degrees against prosternal horizontal line,
<figureCitation id="ECAE2A37327FB3138137BCECBDC8F821" box="[622,705,1953,1979]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1F</figureCitation>
: red line). Prosternal spine in profile weakly inclined (at 9–10 degrees against prosternal horizontal line,
<figureCitation id="ECAE2A37327FB313810ABC88BDB7F845" box="[595,702,1989,2015]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="3" pageNumber="92">Fig. 1H, I</figureCitation>
: red line), with subapical tooth; lateral lobe in profile with ventral margin almost straight to weakly emarginate (
<figureCitation id="ECAE2A37327AB31681C2BDF1BC02F94D" box="[667,779,1724,1751]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="6" pageNumber="95">Fig. 1H, I</figureCitation>
: yellow line), at ventral view with sides roundly expanded and anteriorly strongly prominent (
<figureCitation id="ECAE2A37327AB3168147BDADBD7EF961" box="[542,631,1760,1787]" captionStart="FIGURE 2" captionStartId="5.[151,250,798,823]" captionTargetBox="[189,1399,187,787]" captionTargetId="figure@5.[189,1399,181,794]" captionTargetPageId="5" captionText="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="6" pageNumber="95">Fig. 2A</figureCitation>
: arrows); apical margin at ventral view rounded, with weakly prominent apex (
<figureCitation id="ECAE2A37327AB3168382BC49BE3DF885" box="[219,308,1796,1823]" captionStart="FIGURE 2" captionStartId="5.[151,250,798,823]" captionTargetBox="[189,1399,187,787]" captionTargetId="figure@5.[189,1399,181,794]" captionTargetPageId="5" captionText="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="6" pageNumber="95">Fig. 2A</figureCitation>
). Pronotosternal sutures not grooved, anteriorly opened and appearing double (
<figureCitation id="ECAE2A37327AB31687FABC49BBF5F885" box="[1187,1276,1796,1823]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="6" pageNumber="95">Fig. 1G</figureCitation>
: white arrow). Scutellum 0.95–1.00 x as long as wide (
<figureCitation id="ECAE2A37327AB316813ABC65BDD6F8D9" box="[611,735,1832,1859]" captionStart="FIGURE 2" captionStartId="5.[151,250,798,823]" captionTargetBox="[189,1399,187,787]" captionTargetId="figure@5.[189,1399,181,794]" captionTargetPageId="5" captionText="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="6" pageNumber="95">Fig. 2B, C</figureCitation>
), widening posteriad, widest near posterior 1/5, strongly convex above (
<figureCitation id="ECAE2A37327AB3168243BC01BE7CF8FD" box="[282,373,1868,1895]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="6" pageNumber="95">Fig. 1H</figureCitation>
: arrow), strongly inclined anterior-downwards; anterior margin almost transverse or shallowly emarginate; posterior margin broadly rounded, but apex almost transverse. Mesosternum with a median longitudinal carina posterior to mesosternal cavity; borders of mesosternal cavity weakly sinuate followed by strongly declivous (
<figureCitation id="ECAE2A37327AB31683F9BCF5BE1BF849" box="[160,274,1976,2003]" captionStart="FIGURE 1" captionStartId="4.[151,250,1819,1844]" captionTargetBox="[164,1423,187,1784]" captionTargetId="figure@4.[164,1423,179,1792]" captionTargetPageId="4" captionText="FIGURE 1. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above)." figureDoi="http://doi.org/10.5281/zenodo.3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="6" pageNumber="95">Fig. 1H, I</figureCitation>
: green line). Mesepisternum not reaching mesocoxal cavity (
<figureCitation id="ECAE2A37327AB316809ABCF5BB17F849" box="[963,1054,1976,2003]" captionStart="FIGURE 2" captionStartId="5.[151,250,798,823]" captionTargetBox="[189,1399,187,787]" captionTargetId="figure@5.[189,1399,181,794]" captionTargetPageId="5" captionText="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="6" pageNumber="95">Fig. 2D</figureCitation>
). Mesepimeron with deep depression for reception of femora of mid-legs (
<figureCitation id="ECAE2A37327AB316812BBC91BDC4F86D" box="[626,717,2012,2039]" captionStart="FIGURE 2" captionStartId="5.[151,250,798,823]" captionTargetBox="[189,1399,187,787]" captionTargetId="figure@5.[189,1399,181,794]" captionTargetPageId="5" captionText="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="6" pageNumber="95">Fig. 2D</figureCitation>
). Metasternum sulcate medially and behind mesocoxae. Elytral striae I and II weakly grooved; remaining striae not grooved and defined only by rows of punctures. Tarsomeres simple.
</paragraph>
<caption id="20EA663A3278B31483CEBC56BB20F872" ID-DOI="http://doi.org/10.5281/zenodo.3674495" ID-Zenodo-Dep="3674495" httpUri="https://zenodo.org/record/3674495/files/figure.png" pageId="4" pageNumber="93" startId="4.[151,250,1819,1844]" targetBox="[164,1423,187,1784]" targetPageId="4">
<paragraph id="742A36B23278B31483CEBC56BB20F872" blockId="4.[151,1437,1819,2024]" pageId="4" pageNumber="93">
<emphasis id="46E1EAA03278B31483CEBC56BE1DF8AE" bold="true" box="[151,276,1819,1844]" pageId="4" pageNumber="93">FIGURE 1.</emphasis>
<taxonomicName id="B3954D313278B3148245BC50BDA3F8AE" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[284,682,1819,1844]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="sapphirus" status="sp. nov.">
<emphasis id="46E1EAA03278B3148245BC50BEEFF8AE" box="[284,486,1820,1844]" italics="true" pageId="4" pageNumber="93">Limonius sapphirus</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
<taxonomicNameLabel id="5DD257DB3278B31481EBBC51BDE3F8A9" box="[690,746,1820,1843]" pageId="4" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
A, C–H: holotype, male. B, I: paratype, female. D–G: mouth-parts and right antenna removed. A, B: habitus, dorsal view; C: habitus, ventral view; D: head, dorsal view; E: head and pronotum, laterodorsal view (arrow: lateral carina weakly prominent behind anterior angle); F: head and anterior half of prothorax, lateral view (arrow: supra-antennal carina strongly prominent anteriad); G: hypomeron and prosternum, lateroventral view (black arrow: mesial edge of hypomeron carinate anterolaterad; white arrow: pronotosternal sutures slightly grooved anteriorly); H, I: prosternal spine and mesosternum, lateral view (arrow: scutellum strongly convex above).
</paragraph>
</caption>
<caption id="20EA663A3279B31583CEB853BB40FCE4" ID-DOI="http://doi.org/10.5281/zenodo.3674497" ID-Zenodo-Dep="3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="5" pageNumber="94" startId="5.[151,250,798,823]" targetBox="[189,1399,187,787]" targetPageId="5">
<paragraph id="742A36B23279B31583CEB853BB40FCE4" blockId="5.[151,1436,798,895]" pageId="5" pageNumber="94">
<emphasis id="46E1EAA03279B31583CEB853BE1AFCAD" bold="true" box="[151,275,798,823]" pageId="5" pageNumber="94">FIGURE 2.</emphasis>
<taxonomicName id="B3954D313279B3158243B86DBDADFCAD" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[282,676,798,823]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="sapphirus" status="sp. nov.">
<emphasis id="46E1EAA03279B3158243B86DBEEBFCAD" box="[282,482,799,823]" italics="true" pageId="5" pageNumber="94">Limonius sapphirus</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
<taxonomicNameLabel id="5DD257DB3279B31581F2B852BDE8FCAC" box="[683,737,799,822]" pageId="5" pageNumber="94" rank="species">sp. n.</taxonomicNameLabel>
A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V.
</paragraph>
</caption>
<caption id="20EA663A3279B31583CEBCFABC2BF86E" ID-DOI="http://doi.org/10.5281/zenodo.3674499" ID-Zenodo-Dep="3674499" httpUri="https://zenodo.org/record/3674499/files/figure.png" pageId="5" pageNumber="94" startId="5.[151,250,1975,2000]" targetBox="[189,1398,917,1950]" targetPageId="5">
<paragraph id="742A36B23279B31583CEBCFABC2BF86E" blockId="5.[151,1437,1975,2036]" pageId="5" pageNumber="94">
<emphasis id="46E1EAA03279B31583CEBCFABE04F84A" bold="true" box="[151,269,1975,2000]" pageId="5" pageNumber="94">FIGURE 3</emphasis>
.
<taxonomicName id="B3954D313279B3158240BCF4BDA8F84A" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[281,673,1975,2000]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="94" phylum="Arthropoda" rank="species" species="sapphirus" status="sp. nov.">
<emphasis id="46E1EAA03279B3158240BCF4BEE8F84A" box="[281,481,1976,2000]" italics="true" pageId="5" pageNumber="94">Limonius sapphirus</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
<taxonomicNameLabel id="5DD257DB3279B31581FEBCF5BDD7F855" box="[679,734,1976,1999]" pageId="5" pageNumber="94" rank="species">sp. n.</taxonomicNameLabel>
, male, holotype. A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view.
</paragraph>
</caption>
<caption id="20EA663A327AB31683CEBE8CBB43F914" ID-DOI="http://doi.org/10.5281/zenodo.3674501" ID-Zenodo-Dep="3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="6" pageNumber="95" startId="6.[151,250,1473,1498]" targetBox="[182,1436,181,1448]" targetPageId="6">
<paragraph id="742A36B2327AB31683CEBE8CBB43F914" blockId="6.[151,1436,1473,1678]" pageId="6" pageNumber="95">
<emphasis id="46E1EAA0327AB31683CEBE8CBE1DFA40" bold="true" box="[151,276,1473,1498]" pageId="6" pageNumber="95">FIGURE 4.</emphasis>
<taxonomicName id="B3954D31327AB3168242BE8EBDA1FA40" authority="Arimoto &amp; Suzuki" authorityName="Arimoto &amp; Suzuki" authorityYear="2020" box="[283,680,1473,1498]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="95" phylum="Arthropoda" rank="species" species="sapphirus" status="sp. nov.">
<emphasis id="46E1EAA0327AB3168242BE8EBEECFA40" box="[283,485,1474,1498]" italics="true" pageId="6" pageNumber="95">Limonius sapphirus</emphasis>
Arimoto &amp; Suzuki
</taxonomicName>
<taxonomicNameLabel id="5DD257DB327AB31681E9BE8FBDE1FA43" box="[688,744,1474,1497]" pageId="6" pageNumber="95" rank="species">sp. n.</taxonomicNameLabel>
, female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented).
</paragraph>
</caption>
<paragraph id="742A36B2327BB317839EBBACBBEBFE11" blockId="7.[151,1437,153,1403]" pageId="7" pageNumber="96">
Abdomen with external margin micro-serrated (
<figureCitation id="ECAE2A37327BB3178184BBACBC3DFF61" box="[733,820,225,251]" captionStart="FIGURE 2" captionStartId="5.[151,250,798,823]" captionTargetBox="[189,1399,187,787]" captionTargetId="figure@5.[189,1399,181,794]" captionTargetPageId="5" captionText="FIGURE 2. Limonius sapphirus Arimoto &amp; Suzuki sp. n. A, B, D, E: holotype, male. C: paratype, female. A: prosternal spine, ventral view (arrows: sides at ventral view anteriorly strongly prominent); B, C: scutellum; D: posterior half of prothorax, mesothorax and anterior part of metathorax, lateroventral view; E: external margin of ventrite V." figureDoi="http://doi.org/10.5281/zenodo.3674497" httpUri="https://zenodo.org/record/3674497/files/figure.png" pageId="7" pageNumber="96">Fig. 2E</figureCitation>
); ventrite V 0.54–0.55 x as long as wide.
<emphasis id="46E1EAA0327BB317865DBBACBA41FF61" bold="true" box="[1284,1352,225,251]" pageId="7" pageNumber="96">Male.</emphasis>
Tergite VIII 0.81 x as long as wide (
<figureCitation id="ECAE2A37327BB317828EBA48BD23FEBA" box="[471,554,261,288]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,2000]" captionTargetBox="[189,1398,917,1950]" captionTargetId="figure@5.[189,1398,917,1950]" captionTargetPageId="5" captionText="FIGURE 3. Limonius sapphirus Arimoto &amp; Suzuki sp. n., male, holotype.A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674499" httpUri="https://zenodo.org/record/3674499/files/figure.png" pageId="7" pageNumber="96">Fig.3A</figureCitation>
). Sternite VIII with median notch rounded (
<figureCitation id="ECAE2A37327BB317874CBA48BB65FE85" box="[1045,1132,261,287]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,2000]" captionTargetBox="[189,1398,917,1950]" captionTargetId="figure@5.[189,1398,917,1950]" captionTargetPageId="5" captionText="FIGURE 3. Limonius sapphirus Arimoto &amp; Suzuki sp. n., male, holotype.A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674499" httpUri="https://zenodo.org/record/3674499/files/figure.png" pageId="7" pageNumber="96">Fig. 3B</figureCitation>
). Tergite IX almost as long as length of tergite X (
<figureCitation id="ECAE2A37327BB31782C8BA64BEEAFEDE" box="[401,483,297,324]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,2000]" captionTargetBox="[189,1398,917,1950]" captionTargetId="figure@5.[189,1398,917,1950]" captionTargetPageId="5" captionText="FIGURE 3. Limonius sapphirus Arimoto &amp; Suzuki sp. n., male, holotype.A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674499" httpUri="https://zenodo.org/record/3674499/files/figure.png" pageId="7" pageNumber="96">Fig.3C</figureCitation>
). Tergite X 2.1 x as long as wide (
<figureCitation id="ECAE2A37327BB3178007BA64BCB9FEDE" box="[862,944,297,324]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,2000]" captionTargetBox="[189,1398,917,1950]" captionTargetId="figure@5.[189,1398,917,1950]" captionTargetPageId="5" captionText="FIGURE 3. Limonius sapphirus Arimoto &amp; Suzuki sp. n., male, holotype.A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674499" httpUri="https://zenodo.org/record/3674499/files/figure.png" pageId="7" pageNumber="96">Fig.3C</figureCitation>
). Sternite IX rounded apically (
<figureCitation id="ECAE2A37327BB3178656BA64BA6AFEDE" box="[1295,1379,297,324]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,2000]" captionTargetBox="[189,1398,917,1950]" captionTargetId="figure@5.[189,1398,917,1950]" captionTargetPageId="5" captionText="FIGURE 3. Limonius sapphirus Arimoto &amp; Suzuki sp. n., male, holotype.A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674499" httpUri="https://zenodo.org/record/3674499/files/figure.png" pageId="7" pageNumber="96">Fig.3D</figureCitation>
).
<emphasis id="46E1EAA0327BB317862CBA64BFD1FEFD" bold="true" pageId="7" pageNumber="96">Female.</emphasis>
Terigite VIII 0.98 x as long as wide (
<figureCitation id="ECAE2A37327BB3178125BA00BDDDFEFD" box="[636,724,333,359]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4B</figureCitation>
). Sternite VIII with side almost parallel on basal half (
<figureCitation id="ECAE2A37327BB317866FBA00BA86FEFD" box="[1334,1423,333,359]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4C</figureCitation>
); spiculum ventrale 1.9 x as long as sternite VIII (
<figureCitation id="ECAE2A37327BB31781EABA3CBC04FE11" box="[691,781,369,395]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4A</figureCitation>
), curved twice apically (
<figureCitation id="ECAE2A37327BB3178779BA3CBB73FE11" box="[1056,1146,369,395]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4A</figureCitation>
: arrows).
</paragraph>
<paragraph id="742A36B2327BB317839EBAD8BE47FD4A" blockId="7.[151,1437,153,1403]" pageId="7" pageNumber="96">
Genitalia.
<emphasis id="46E1EAA0327BB3178267BAD8BE88FE35" bold="true" box="[318,385,405,431]" pageId="7" pageNumber="96">Male.</emphasis>
Aedeagus (
<figureCitation id="ECAE2A37327BB3178151BAD8BD88FE35" box="[520,641,405,431]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,2000]" captionTargetBox="[189,1398,917,1950]" captionTargetId="figure@5.[189,1398,917,1950]" captionTargetPageId="5" captionText="FIGURE 3. Limonius sapphirus Arimoto &amp; Suzuki sp. n., male, holotype.A: tergite VIII; B: sternite VIII; C: tergites IX–X; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674499" httpUri="https://zenodo.org/record/3674499/files/figure.png" pageId="7" pageNumber="96">Fig. 3E, F</figureCitation>
). Basal piece 0.38 x total length of aedeagus. Median lobe exceeding apices of parameres; basal struts 0.31 x total length of median lobe. Parameres separated ventrally; apex beyond lateral subapical barb longer sub-triangular (
<figureCitation id="ECAE2A37327BB31781D4BA90BDE1FE6D" box="[653,744,477,503]" captionStart="FIGURE 7" captionStartId="9.[151,250,1830,1855]" captionTargetBox="[158,1426,1166,1818]" captionTargetId="figure@9.[151,1436,1159,1826]" captionTargetPageId="9" captionText="FIGURE 7. Limonius spp., male, holotype, apical part of paramere A: Limonius sapphirus Arimoto &amp; Suzuki sp. n.; B: Limonius onyx Arimoto &amp; Suzuki sp. n." figureDoi="http://doi.org/10.5281/zenodo.3674507" httpUri="https://zenodo.org/record/3674507/files/figure.png" pageId="7" pageNumber="96">Fig. 7A</figureCitation>
), rounded apically, 0.11 x length of dorsal side of parameres, without setae dorsally, with some long setae ventrally.
<emphasis id="46E1EAA0327BB31781A4B94CBC52FD81" bold="true" box="[765,859,513,539]" pageId="7" pageNumber="96">Female.</emphasis>
Coxites two segmented at ventral side (
<figureCitation id="ECAE2A37327BB3178644B94CBA7AFD81" box="[1309,1395,513,539]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4F</figureCitation>
: arrow), with some setae; stylus with setae near apex. Vagina long; uterus with dome-shaped sac at each dorsal and ventral side (
<figureCitation id="ECAE2A37327BB3178273B904BE8FFDF9" box="[298,390,585,611]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4D</figureCitation>
: white arrow;
<figureCitation id="ECAE2A37327BB3178170B904BD88FDF9" box="[553,641,585,611]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4E</figureCitation>
: white triangles), with two dome-shaped bulges at right side (
<figureCitation id="ECAE2A37327BB3178664B904BA90FDF9" box="[1341,1433,585,611]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4D</figureCitation>
: black arrow;
<figureCitation id="ECAE2A37327BB3178272B920BE8BFD1D" box="[299,386,621,647]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4E</figureCitation>
: black triangles); bursa copulatrix thick tubular, without sclerotized structures, with two short thin sacs (
<figureCitation id="ECAE2A37327BB3178250B9DCBE6CFD31" box="[265,357,657,683]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4D</figureCitation>
: white triangles), with a long thin sac extending from apex of bursa copulatrix (
<figureCitation id="ECAE2A37327BB31787B0B9DCBA4CFD31" box="[1257,1349,657,683]" captionStart="FIGURE 4" captionStartId="6.[151,250,1473,1498]" captionTargetBox="[182,1436,181,1448]" captionTargetId="figure@6.[151,1436,181,1448]" captionTargetPageId="6" captionText="FIGURE 4. Limonius sapphirus Arimoto &amp; Suzuki sp. n., female, paratype. A: tergite VIII and sternite VIII, dorsal view (arrows: spiculum ventrale curved twice apically); B: tergite VIII; C: sternite VIII; D: genitalia, dorsal view (white arrow: uterus with dome-shaped sac at each side; black arrow: uterus with two dome-shaped bulges at right side; white triangles: bursa copulatrix with two short thin sacs; double white triangles: long thin sac extending from apex of bursa copulatrix); E: genitalia, lateral side (white triangles: uterus with dome-shaped bulges at dorsal and ventral sides; black triangles: uterus with two dome-shaped bulges at right side); F; apical part of ovipositor, ventral side (arrow: coxites two segmented)." figureDoi="http://doi.org/10.5281/zenodo.3674501" httpUri="https://zenodo.org/record/3674501/files/figure.png" pageId="7" pageNumber="96">Fig. 4D</figureCitation>
: double white triangles).
</paragraph>
<paragraph id="742A36B2327BB317839EB995BE63FACD" blockId="7.[151,1437,153,1403]" pageId="7" pageNumber="96">
<emphasis id="46E1EAA0327BB317839EB995BEB9FD69" bold="true" box="[199,432,728,755]" pageId="7" pageNumber="96">Comparative notes.</emphasis>
This species is distinct from the congeners by the whole blue to green body with metallic luster. It is relatively similar to
<taxonomicName id="B3954D31327BB31782AFB9B3BC56FC8D" authority="Lewis, 1894" authorityName="Lewis" authorityYear="1894" box="[502,863,765,791]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="ignicollis">
<emphasis id="46E1EAA0327BB31782AFB9B3BDC7FC8D" box="[502,718,765,791]" italics="true" pageId="7" pageNumber="96">Limonius ignicollis</emphasis>
<bibRefCitation id="10044B43327BB317818CB9B0BC56FC8D" author="Lewis, G." box="[725,863,765,791]" pageId="7" pageNumber="96" pagination="311 - 320" refId="ref9779" refString="Lewis, G. (1894) On the Elateridae of Japan. Annals and Magazine of Natural History, Series 6, 13, 182 - 201, 311 - 320. https: // doi. org / 10.1080 / 00222939408677706" type="journal article" year="1894">
Lewis,
<quantity id="B36D9B57327BB317807EB9B0BC56FC8D" box="[807,863,765,791]" metricMagnitude="1" metricUnit="m" metricValue="4.81076" pageId="7" pageNumber="96" unit="in" value="1894.0">1894</quantity>
</bibRefCitation>
</taxonomicName>
in having the following characters: frons in profile weakly depressed apically; supra-antennal carina in profile strongly prominent anteriad above clypeus; antennomere III slightly longer than II; antennomere IV longer than II–III combined and longer than V; prothorax widening posteriad; prosternal spine with subapical tooth; mesial edge of hypomeron carinate anterolaterad; sides of prosternal spine at ventral view anteriorly strongly prominent; scutellum strongly convex. However it is distinguished by the following contrasting characters (
<taxonomicName id="B3954D31327BB317816EB8FFBDB5FC51" authorityName="Lewis" authorityYear="1894" box="[567,700,945,971]" class="Insecta" family="Elateridae" genus="Limonius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="ignicollis">
<emphasis id="46E1EAA0327BB317816EB8FFBDB5FC51" box="[567,700,945,971]" italics="true" pageId="7" pageNumber="96">L. ignicollis</emphasis>
</taxonomicName>
in parentheses), except for body color: elytra interspaces between punctures lager than puncture diameter (elytra interspaces between punctures smaller than puncture diameter); supra-antennal carina in dorsal view broadly rounded but almost straight apically (supra-antennal carina in dorsal view broadly rounded but weakly emarginate apically); antennae extending beyond posterior apices of pronotum by apical three antennomeres in male and by apical antennomere in female (antennae extending beyond posterior apices of pronotum by apical two antennomeres in male and by apical antennomere in female); antennomere IV distinctly longer than wide (antennomere IV almost as long as wide); lateral carina of prothorax weakly prominent just behind anterior angles (lateral carina of prothorax strongly prominent just behind anterior angles); anterior lobe of prosternum in profile weakly inclined (anterior lobe of prosternum in profile strongly inclined); scutellum almost as long as wide or slightly shorter than wide (scutellum distinctly longer than wide); posterior margin of scutellum broadly rounded, but apex almost transverse (posterior margin of scutellum broadly rounded, but apex often shallowly emarginate).
</paragraph>
</subSubSection>
<subSubSection id="3C8F6539327BB317839EBE2DBB7BFAE1" box="[199,1138,1376,1403]" pageId="7" pageNumber="96" type="distribution">
<paragraph id="742A36B2327BB317839EBE2DBB7BFAE1" blockId="7.[151,1437,153,1403]" box="[199,1138,1376,1403]" pageId="7" pageNumber="96">
<emphasis id="46E1EAA0327BB317839EBE2DBE68FAE1" bold="true" box="[199,353,1376,1403]" pageId="7" pageNumber="96">Distribution.</emphasis>
<collectingCountry id="0C827622327BB317823EBE2CBEB4FAE1" box="[359,445,1377,1403]" name="Taiwan" pageId="7" pageNumber="96">Taiwan</collectingCountry>
: around Mt. Lala, located between
<collectingRegion id="B651F850327BB317801FBE2CBCAFFAE1" box="[838,934,1377,1403]" country="Taiwan" name="Taoyuan" pageId="7" pageNumber="96">Taoyuan</collectingRegion>
and
<collectingRegion id="B651F850327BB3178085BE2CBB2BFAE1" box="[988,1058,1377,1403]" country="Taiwan" name="Taipei" pageId="7" pageNumber="96">Taipei</collectingRegion>
Cities.
</paragraph>
</subSubSection>
</treatment>
</document>
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