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<document id="0CF35570B738CE20D0C333AF439DEA28" ID-DOI="10.11646/zootaxa.5020.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5223173" ID-ZooBank="4FF882DF-334F-49C8-A576-4192B5F2654C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629388374973" checkinUser="plazi" docAuthor="Ingrisch, Sigfrid" docDate="2021" docId="FA08E75DAA348052FDC03401932B6F95" docLanguage="en" docName="zootaxa.5020.2.1.pdf" docOrigin="Zootaxa 5020 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Habetia tuta Ingrisch 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="213" masterDocId="06319F25AA3F805EFD57366B91686D58" masterDocTitle="Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)" masterLastPageNumber="256" masterPageNumber="201" pageNumber="212" updateTime="1699071125630" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="0F9FE7DA0321501B0EC64D97545EE3CB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4B0FCC89750FEC3355AF895CB5DD7B5F">
<mods:title id="54FFA456AC10071359D1A4A53F8E82E3">Revision of the genera Habetia Kirby, 1906 and Parahabetia gen. nov. from New Guinea (Orthoptera, Tettigoniidae, Conocephalinae, Agraeciini)</mods:title>
</mods:titleInfo>
<mods:name id="8A5F7CB4435FC8E32DACF404955DDC8C" type="personal">
<mods:role id="C44944B69F8F6BD8296F902368530149">
<mods:roleTerm id="B8138D0E1F6A3EB95EC5B54B91E198C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9995DE5207B9CD96567AD2A57891642D">Ingrisch, Sigfrid</mods:namePart>
</mods:name>
<mods:typeOfResource id="ADD257008A3A12D620F750D18A10F126">text</mods:typeOfResource>
<mods:relatedItem id="3BAE3105396CCB5635B9A9C15BD20696" type="host">
<mods:titleInfo id="AF5B44D01406084E392305F430C0814F">
<mods:title id="D4A59E2D1435E21B8369356837B862B2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C34EBD5FD7F4BF452E6C84FDD3CE27D1">
<mods:date id="78182D5C8923FB4283B9A319A4861BB3">2021</mods:date>
<mods:detail id="1C34AE00A52CD8D6DF1480ABD1913BB1" type="pubDate">
<mods:number id="AC324FA3A55C94247D3BDE0952B57129">2021-08-12</mods:number>
</mods:detail>
<mods:detail id="06B08377D23C18B339C4FD2489DF57E3" type="volume">
<mods:number id="6A575282BD925806F24AC5218AA484EA">5020</mods:number>
</mods:detail>
<mods:detail id="11ECA325F7D77D95204B7D4AD79E45CE" type="issue">
<mods:number id="EC89B3839748453C3E6427E6335D1C70">2</mods:number>
</mods:detail>
<mods:extent id="6737B5450FE08968B8D56F755FF8CFD9" unit="page">
<mods:start id="DC84C570ACE5B6A17C780B474D64C6C8">201</mods:start>
<mods:end id="85D65163B5CE323DB28F7A8E441584EE">256</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B9865832E9ABEA49DB22F7450EA68FFD">journal article</mods:classification>
<mods:identifier id="D80EDDE38565350442C28C930273B610" type="DOI">10.11646/zootaxa.5020.2.1</mods:identifier>
<mods:identifier id="89D354537A6DDE78C3E94E8E0C76AB0A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D659B3BCD4BCC302DF64CB7B94CA36ED" type="Zenodo-Dep">5223173</mods:identifier>
<mods:identifier id="2CE57B27902D9DE17A61F0F0209EBB51" type="ZooBank">4FF882DF-334F-49C8-A576-4192B5F2654C</mods:identifier>
</mods:mods>
<treatment id="FA08E75DAA348052FDC03401932B6F95" ID-DOI="http://doi.org/10.5281/zenodo.5223221" ID-Zenodo-Dep="5223221" LSID="urn:lsid:plazi:treatment:FA08E75DAA348052FDC03401932B6F95" httpUri="http://treatment.plazi.org/id/FA08E75DAA348052FDC03401932B6F95" lastPageId="12" lastPageNumber="213" pageId="11" pageNumber="212">
<subSubSection id="3ABB05C0AA348055FDC0340190FB6FDD" box="[151,403,618,645]" pageId="11" pageNumber="212" type="nomenclature">
<paragraph id="721E564BAA348055FDC0340190FB6FDD" blockId="11.[151,812,618,681]" box="[151,403,618,645]" pageId="11" pageNumber="212">
<heading id="2956E127AA348055FDC0340190FB6FDD" bold="true" box="[151,403,618,645]" fontSize="11" level="1" pageId="11" pageNumber="212" reason="1">
<emphasis id="40D58A59AA348055FDC0340190FB6FDD" bold="true" box="[151,403,618,645]" pageId="11" pageNumber="212">
<taxonomicName id="B5A12DC8AA348055FDC0340190466FDD" ID-CoL="8QK86" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[151,302,618,645]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="tuta" status="sp. nov.">
<emphasis id="40D58A59AA348055FDC0340190466FDD" bold="true" box="[151,302,618,645]" italics="true" pageId="11" pageNumber="212">Habetia tuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5BE63722AA348055FC62340090FB6FDD" box="[309,403,619,645]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA348055FDC034E492446FF1" box="[151,812,654,681]" pageId="11" pageNumber="212" type="description">
<paragraph id="721E564BAA348055FDC034E492446FF1" blockId="11.[151,812,618,681]" box="[151,812,654,681]" pageId="11" pageNumber="212">
<figureCitation id="EA9A4ACEAA348055FDC034E491996FF1" box="[151,241,655,681]" captionStart="FIGURE 1" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[163,1425,181,1920]" captionTargetId="figure-19@10.[162,1425,181,1920]" captionTargetPageId="10" captionText="FIGURE 1. Habetia sp. females: habitus lateral view; species with very long ovipositor: A. H. spada (Brunner, 1898) holotype; B. H. tuta sp. nov. (Keram River); C. H. pedala sp. nov. (Toricelli Mts); D. H. pallida sp. nov. (Sentani); E–F. H. multispinulosa Griffini, 1908 (E holotype, F Rigo Dorom). Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223175" httpUri="https://zenodo.org/record/5223175/files/figure.png" pageId="11" pageNumber="212">Figs 1B</figureCitation>
,
<figureCitation id="EA9A4ACEAA348055FDAA34E490486FF1" box="[253,288,655,681]" captionStart="FIGURE 3" captionStartId="17.[151,250,1860,1884]" captionTargetBox="[177,1409,182,1837]" captionTargetId="figure-15@17.[177,1409,181,1837]" captionTargetPageId="17" captionText="FIGURE 3. A, Habetia pilleata sp. nov. male (Wau 1200m); B, H. pallida sp. nov. female (Hollandia); C, H. tuta sp. nov. male (Quelllager); D–F, H. curvata sp. nov. (D, male Albatros Bivak, E–F, female Prauwenbivak); G, H. spada (Brunner, 1898) female holotype (Konstantinhafen); H, H. multispinulosa Griffini, 1908 female holotype (Dilo); I, H. pedala sp. nov. female (Toricelli Mountains).—Frons (A–E, G–I), female dorsal view (F). Scales 5 mm (A–E, G–I), 10 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5223179" httpUri="https://zenodo.org/record/5223179/files/figure.png" pageId="11" pageNumber="212">3C</figureCitation>
,
<figureCitation id="EA9A4ACEAA348055FC7B34E590016FF1" box="[300,361,654,681]" captionStart="FIGURE 5" captionStartId="19.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1620]" captionTargetId="figure-15@19.[151,1436,181,1621]" captionTargetPageId="19" captionText="FIGURE 5. Female subgenital plate of Habetia species in oblique ventral (A, J), ventral (C, E, G, L), apico-ventral (B, D, G, K), apical (H), and lateral (F, I) view.—A, H. spada holotype; B–C, H. multispinulosa (B holotype, C from Dorom), D, H. pedala sp. nov. (Toricelli Mts); E–F, H. tuta sp. nov. (Keram river); G, H. curvata sp. nov. (Prauwenbivak); H–L, H. pallida sp. nov. (H–I Jutefa Bay, J Bewani, K Ifar, L Hollandia).—Scales 1 mm; A, D, G not to scale." figureDoi="http://doi.org/10.5281/zenodo.5223183" httpUri="https://zenodo.org/record/5223183/files/figure.png" pageId="11" pageNumber="212">5E–F</figureCitation>
,
<figureCitation id="EA9A4ACEAA348055FC2334E490FF6FF1" box="[372,407,655,681]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (A–B, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); C–E, Habetia pedala sp. nov. (Toricelli Mts); F–H, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (A–C, F–G), 1 mm (D–E, H). Phallus removed in D, F–G." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="11" pageNumber="212">8B</figureCitation>
,
<figureCitation id="EA9A4ACEAA348055FCF434E490BC6FF1" box="[419,468,655,681]" captionStart="FIGURE 11" captionStartId="27.[151,250,1579,1603]" captionTargetBox="[151,1436,181,1555]" captionTargetId="figure-15@27.[151,1436,181,1555]" captionTargetPageId="27" captionText="FIGURE 11. Male subgenital plate and cerci of Habetia species in ventral view (J in ventro-apical view).—A, H. tuta sp. nov. (Rosensee); B, H. pallida sp. nov. (Ifar); C, H. pedala sp. nov. (Toricelli Mts); D, H. curvata sp. nov. (Albatros Bivak); E, H. imitatrix Karny, 1912 holotype (Bongu); F, H. dentata sp. nov. (Saidor); G, H. bivittata sp. nov. (Goroka); H, H. simbai sp. nov. (Simbai Valley); I, H. kondiu sp. nov. (Kondiu); J, H. lalibu sp. nov. (Lalibu); K, H. wau sp. nov. (Nami Creek). Scales 5 mm (A–E), 1 mm (F–K)." figureDoi="http://doi.org/10.5281/zenodo.5223195" httpUri="https://zenodo.org/record/5223195/files/figure.png" pageId="11" pageNumber="212">11A</figureCitation>
,
<figureCitation id="EA9A4ACEAA348055FCB734E4935C6FF1" box="[480,564,655,681]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: A–C, H. tuta sp. nov. (A Quelllager, B–C Töpferfluss); D–E, H. pedala sp. nov. (Toricelli Mts); F–I, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="11" pageNumber="212">12A–C</figureCitation>
,
<figureCitation id="EA9A4ACEAA348055FF6834E4931A6FF1" box="[575,626,655,681]" captionStart="FIGURE 18" captionStartId="35.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-15@35.[151,1436,181,1786]" captionTargetPageId="35" captionText="FIGURE 18. Stridulatory files on underside of left tegmen of males of Habetia species: A, H. tuta sp. nov. (Lake Chambri); B, H. pallida sp. nov. (Waris); C, H. pedala sp. nov. (Toricelli Mts); D, H. imitatrix Karny, 1912 (Bongu); E, H. dentata sp. nov. (Saidor); F, H. pilleata sp. nov. (Wau 1200 m); G–H, H. wau sp. nov. (G Nami Creek; H Wau).—Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223217" httpUri="https://zenodo.org/record/5223217/files/figure.png" pageId="11" pageNumber="212">18A</figureCitation>
,
<figureCitation id="EA9A4ACEAA348055FF2A34E493D86FF1" box="[637,688,655,681]" captionStart="FIGURE 20" captionStartId="40.[151,250,1843,1867]" captionTargetBox="[151,1436,181,1809]" captionTargetId="figure-19@40.[151,1436,181,1820]" captionTargetPageId="40" captionText="FIGURE 20. Habetia sp. males: Habitus in dorsal view with left wings spread.—A, H. bivittata sp. nov. (Moife); B, H. kondiu sp. nov. (Wabag); C, H. dentata sp. nov. (Saidor); D, H. lalibu sp. nov. (Lalibu); E, H. pilleata sp. nov. (Wau); F, H. simbai sp. nov. (Simbai Valley); G, H. wau sp. nov. (Wau); H, H. tuta sp. nov. (Lake Chambri).—All scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223223" httpUri="https://zenodo.org/record/5223223/files/figure.png" pageId="11" pageNumber="212">20H</figureCitation>
, Map 1(3).
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA348055FDC034BC938B6ED9" pageId="11" pageNumber="212" type="materials_examined">
<paragraph id="721E564BAA348055FDC034BC95EC6E4D" blockId="11.[151,1437,726,2013]" pageId="11" pageNumber="212">
<materialsCitation id="C2C95C16AA348055FDC034BC95EC6E4D" collectingDate="1913-02-16" collectionCode="ZMB" collectorName="Burgers &amp; Museum &amp; Naturkunde Berlin" country="Papua New Guinea" latitude="-4.3" location="Lager am Rosensee" longLatPrecision="1306" longitude="143.11667" pageId="11" pageNumber="212" specimenCount="1" stateProvince="East Sepik" typeStatus="holotype">
<emphasis id="40D58A59AA348055FDC034BC900C6FA8" bold="true" box="[151,356,726,753]" pageId="11" pageNumber="212">
<typeStatus id="AD1AE8E9AA348055FDC034BC906C6FA9" box="[151,260,727,753]" pageId="11" pageNumber="212" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="0AB616DBAA348055FC2734BC933D6FA9" box="[368,597,727,753]" name="Papua New Guinea" pageId="11" pageNumber="212">Papua New Guinea</collectingCountry>
:
<collectingRegion id="B06598A9AA348055FF3434BC938A6FA9" box="[611,738,727,753]" country="Papua New Guinea" name="East Sepik" pageId="11" pageNumber="212">East Sepik</collectingRegion>
,
<location id="777E0090AA348055FFA634BC92BD6FA9" LSID="urn:lsid:plazi:treatment:FA08E75DAA348052FDC03401932B6F95:777E0090AA348055FFA634BC92BD6FA9" box="[753,981,727,753]" country="Papua New Guinea" latitude="-4.3" longLatPrecision="1306" longitude="143.11667" name="Lager am Rosensee" pageId="11" pageNumber="212" stateProvince="East Sepik">Lager am Rosensee</location>
[= Lake Chambri], (
<geoCoordinate id="1795308CAA348055F99934BC94766FA9" box="[1230,1310,727,753]" degrees="4" direction="south" minutes="18" orientation="latitude" pageId="11" pageNumber="212" precision="925" value="-4.3">4°18’S</geoCoordinate>
,
<geoCoordinate id="1795308CAA348055F87934BC94E66FA9" box="[1326,1422,727,753]" degrees="143" direction="east" minutes="7" orientation="longitude" pageId="11" pageNumber="212" precision="925" value="143.11667">143°7’E</geoCoordinate>
),
<date id="061F708BAA348055FDC0349090636E4D" box="[151,267,763,789]" pageId="11" pageNumber="212" value="1913-02-16">
<collectingDate id="165B8963AA348055FDC0349090636E4D" box="[151,267,763,789]" pageId="11" pageNumber="212" value="1913-02-16">16.ii.1913</collectingDate>
</date>
, leg.
<collectorName id="DF54339DAA348055FC10349090F76E4D" box="[327,415,763,789]" pageId="11" pageNumber="212">Bürgers</collectorName>
(KAE 267)—depository:
<collectorName id="DF54339DAA348055FF933490924E6E4D" box="[708,806,763,789]" pageId="11" pageNumber="212">Museum</collectorName>
für
<collectorName id="DF54339DAA348055FE023490954E6E4D" box="[853,1062,763,789]" pageId="11" pageNumber="212">Naturkunde Berlin</collectorName>
(
<collectionCode id="14B0CE8EAA348055F9623490951F6E4D" box="[1077,1143,763,789]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="11" pageNumber="212" type="Museum">ZMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="721E564BAA348055FD903574938B6ED9" blockId="11.[151,1437,726,2013]" pageId="11" pageNumber="212">
<materialsCitation id="C2C95C16AA348055FD90357493B76ED9" collectingDate="1912-08-13" collectingDateMax="1913-04-22" collectingDateMin="1912-08-13" collectionCode="ZMB" collectorName="Burgers" country="Papua New Guinea" latitude="-4.383333" location="Lager am Topferfluss" longLatPrecision="1306" longitude="142.78334" pageId="11" pageNumber="212" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="East Sepik" typeStatus="paratype">
Other specimens studied:
<collectingCountry id="0AB616DBAA348055FCA4357493BE6E61" box="[499,726,799,825]" name="Papua New Guinea" pageId="11" pageNumber="212">Papua New Guinea</collectingCountry>
:
<collectingRegion id="B06598A9AA348055FFB33574920A6E61" box="[740,866,799,825]" country="Papua New Guinea" name="East Sepik" pageId="11" pageNumber="212">East Sepik</collectingRegion>
,
<location id="777E0090AA348055FE27357495086E61" LSID="urn:lsid:plazi:treatment:FA08E75DAA348052FDC03401932B6F95:777E0090AA348055FE27357495086E61" box="[880,1120,799,825]" country="Papua New Guinea" latitude="-4.383333" longLatPrecision="1306" longitude="142.78334" name="Lager am Topferfluss" pageId="11" pageNumber="212" stateProvince="East Sepik">Lager am Töpferfluss</location>
[= Keram River], (
<geoCoordinate id="1795308CAA348055F810357494FF6E61" box="[1351,1431,799,825]" degrees="4" direction="south" minutes="27" orientation="latitude" pageId="11" pageNumber="212" precision="925" value="-4.45">4°27’S</geoCoordinate>
,
<geoCoordinate id="1795308CAA348055FDC03528906D6E05" box="[151,261,835,861]" degrees="144" direction="east" minutes="13" orientation="longitude" pageId="11" pageNumber="212" precision="925" value="144.21666">144°13’E</geoCoordinate>
),
<date id="061F708BAA348055FC4F352890F86E05" box="[280,400,835,861]" pageId="11" pageNumber="212" value="1913-04-22">
<collectingDate id="165B8963AA348055FC4F352890F86E05" box="[280,400,835,861]" pageId="11" pageNumber="212" value="1913-04-22">22.iv.1913</collectingDate>
</date>
, leg.
<collectorName id="DF54339DAA348055FC90352893776E05" box="[455,543,835,861]" pageId="11" pageNumber="212">Bürgers</collectorName>
(KAE 317)—
<specimenCount id="64A79DC2AA348055FFE8352892716E05" box="[703,793,835,861]" pageId="11" pageNumber="212" type="female">1 female</specimenCount>
(
<collectionCode id="14B0CE8EAA348055FE71352892006E05" box="[806,872,835,861]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="11" pageNumber="212" type="Museum">ZMB</collectionCode>
); Quelllager, (
<geoCoordinate id="1795308CAA348055F9543528953B6E05" box="[1027,1107,835,861]" degrees="4" direction="south" minutes="23" orientation="latitude" pageId="11" pageNumber="212" precision="925" value="-4.383333">4°23’S</geoCoordinate>
,
<geoCoordinate id="1795308CAA348055F90B352895A46E05" box="[1116,1228,835,861]" degrees="142" direction="east" minutes="47" orientation="longitude" pageId="11" pageNumber="212" precision="925" value="142.78334">142°47’E</geoCoordinate>
),
<date id="061F708BAA348055F98A352894F06E05" box="[1245,1432,835,861]" pageId="11" pageNumber="212" value="1912-08-13" valueMax="1912-08-16" valueMin="1912-08-13">
<collectingDate id="165B8963AA348055F98A352894F06E05" box="[1245,1432,835,861]" pageId="11" pageNumber="212" value="1912-08-13" valueMax="1912-08-16" valueMin="1912-08-13">13.–16.viii.1912</collectingDate>
</date>
, leg.
<collectorName id="DF54339DAA348055FD90350C90776ED9" box="[199,287,871,897]" pageId="11" pageNumber="212">Bürgers</collectorName>
(KAE 128)—
<specimenCount id="64A79DC2AA348055FC92350C93636ED9" box="[453,523,871,897]" pageId="11" pageNumber="212" type="male">1 male</specimenCount>
(
<typeStatus id="AD1AE8E9AA348055FF4D350C93166ED9" box="[538,638,871,897]" pageId="11" pageNumber="212" type="paratype">paratype</typeStatus>
) (
<collectionCode id="14B0CE8EAA348055FFC3350C93BE6ED9" box="[660,726,871,897]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="11" pageNumber="212" type="Museum">ZMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA348055FD9035E1921D680D" pageId="11" pageNumber="212" type="diagnosis">
<paragraph id="721E564BAA348055FD9035E1921D680D" blockId="11.[151,1437,726,2013]" pageId="11" pageNumber="212">
<emphasis id="40D58A59AA348055FD9035E1902A6EFC" bold="true" box="[199,322,906,932]" pageId="11" pageNumber="212">Diagnosis.</emphasis>
The new species differs from its congeners by the titillators that form a pair of simple curved beams that are never markedly widened and have the apical area almost fully recurved (about 150–160°) against the truncate basal stem and are slightly narrowed but obtuse at tip; while in the other species with simple titillators the apical area is either less strongly recurved and has widened and/or narrowed sections in the apical area (
<taxonomicName id="B5A12DC8AA348055F87D359C94F46949" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[1322,1436,1015,1041]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="pallida" status="sp. nov.">
<emphasis id="40D58A59AA348055F87D359C94F46949" box="[1322,1436,1015,1041]" italics="true" pageId="11" pageNumber="212">H. pallida</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA348055FDC03270919A696D" bold="true" box="[151,242,1051,1077]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="5BE63722AA348055FDC03270919A696D" box="[151,242,1051,1077]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="B5A12DC8AA348055FC5632709018696D" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[257,368,1051,1077]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="pedala" status="sp. nov.">
<emphasis id="40D58A59AA348055FC5632709018696D" box="[257,368,1051,1077]" italics="true" pageId="11" pageNumber="212">H. pedala</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA348055FC2F327090BA696D" bold="true" box="[376,466,1051,1077]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="5BE63722AA348055FC2F327090BA696D" box="[376,466,1051,1077]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) or the apical area is recurved against the basal area for little more than 180° and has the tip broadly truncate (
<taxonomicName id="B5A12DC8AA348055FCE3325493506900" authorityName="Karny" authorityYear="1912" box="[436,568,1087,1112]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="40D58A59AA348055FCE3325493506900" box="[436,568,1087,1112]" italics="true" pageId="11" pageNumber="212">H. imitatrix</emphasis>
</taxonomicName>
), or the tip of the titillators is dentate (
<taxonomicName id="B5A12DC8AA348055FEAB3254951D6901" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" box="[1020,1141,1087,1113]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="dentata" status="sp. nov.">
<emphasis id="40D58A59AA348055FEAB3254951D6901" box="[1020,1141,1087,1113]" italics="true" pageId="11" pageNumber="212">H. dentata</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA348055F929325495B16901" bold="true" box="[1150,1241,1087,1113]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="5BE63722AA348055F929325495B16901" box="[1150,1241,1087,1113]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). The male cerci have the apico-internal process compressed and at tip divided into a spined dorsal and an obtuse ventral lobe in
<taxonomicName id="B5A12DC8AA348055F8D6320891AB69F8" authority="Ingrisch, 2021" authorityName="Ingrisch" authorityYear="2021" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="tuta" status="sp. nov.">
<emphasis id="40D58A59AA348055F8D6320891AB69F8" italics="true" pageId="11" pageNumber="212">H. tuta</emphasis>
</taxonomicName>
<emphasis id="40D58A59AA348055FD9D32EC904C69F9" bold="true" box="[202,292,1159,1185]" pageId="11" pageNumber="212">
<taxonomicNameLabel id="5BE63722AA348055FD9D32EC904C69F9" box="[202,292,1159,1185]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
while in
<taxonomicName id="B5A12DC8AA348055FCDA32EC909669F9" authorityName="Ingrisch" authorityYear="2021" box="[397,510,1159,1185]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="40D58A59AA348055FCDA32EC909669F9" box="[397,510,1159,1185]" italics="true" pageId="11" pageNumber="212">H. pallida</emphasis>
</taxonomicName>
, that has the most similar cerci, the dorsal lobe is truncate, only half as long as the ventral lobe and not really separated from it, the ventral lobe terminates into an acute tip and the internal surface of the cercus is provided with a little wavy projecting ridge, in
<taxonomicName id="B5A12DC8AA348055FE6432A492C869B1" authorityName="Ingrisch" authorityYear="2021" box="[819,928,1231,1257]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="pedala">
<emphasis id="40D58A59AA348055FE6432A492C869B1" box="[819,928,1231,1257]" italics="true" pageId="11" pageNumber="212">H. pedala</emphasis>
</taxonomicName>
the apical internal tooth is divided at end into a pair of stout acute teeth of equal length, in
<taxonomicName id="B5A12DC8AA348055FFDA329892676854" authorityName="Karny" authorityYear="1912" box="[653,783,1267,1292]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="imitatrix">
<emphasis id="40D58A59AA348055FFDA329892676854" box="[653,783,1267,1292]" italics="true" pageId="11" pageNumber="212">H. imitatrix</emphasis>
</taxonomicName>
the apical projection of the male cercus is dorso-ventrally compressed and widened and both apical lobes carry a minute spinule at tip, and in
<taxonomicName id="B5A12DC8AA348055F905337C95A16869" authorityName="Ingrisch" authorityYear="2021" box="[1106,1225,1303,1329]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="dentata">
<emphasis id="40D58A59AA348055F905337C95A16869" box="[1106,1225,1303,1329]" italics="true" pageId="11" pageNumber="212">H. dentata</emphasis>
</taxonomicName>
the apical internal projection of the male cercus is prolonged and strongly modified.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA348052FD90333592086FDD" lastPageId="12" lastPageNumber="213" pageId="11" pageNumber="212" type="description">
<paragraph id="721E564BAA348055FD903335925A68BD" blockId="11.[151,1437,726,2013]" pageId="11" pageNumber="212">
<emphasis id="40D58A59AA348055FD90333590316820" bold="true" box="[199,345,1374,1400]" pageId="11" pageNumber="212">Description.</emphasis>
Coloration of face uniformly pale, of general color; lateral areas of face and genae little darker than central area of face, which might be due to conservation; antennal scrobae faintly darkened; mandibles black with few small yellow and dark red spots. Femora with the following number of spines on ventral margins of legs: (1) a 5–8, p 4–8; (2) a 7, p 2–4; (3) a 11–15, p 14–16 (n=3).
</paragraph>
<paragraph id="721E564BAA348052FD90338490C56CD1" blockId="11.[151,1437,726,2013]" lastBlockId="12.[151,1437,151,717]" lastPageId="12" lastPageNumber="213" pageId="11" pageNumber="212">
Male. Stridulatory file on underside of left tegmen with a distinct step in distal area, in
<typeStatus id="AD1AE8E9AA348055F9CE338495936B51" box="[1177,1275,1519,1545]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
with 66 teeth, of which 7 narrow teeth at base,
<quantity id="B559FBAEAA348055FF52307893576B75" box="[517,575,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="9.144" pageId="11" pageNumber="212" unit="in" value="36.0">36 in</quantity>
main part of file, 5 at a step, 13 clearly separated at end, and 6 of decreasing size at end (
<figureCitation id="EA9A4ACEAA348055FC4B305C90EB6B09" box="[284,387,1591,1617]" captionStart="FIGURE 18" captionStartId="35.[151,250,1810,1834]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-15@35.[151,1436,181,1786]" captionTargetPageId="35" captionText="FIGURE 18. Stridulatory files on underside of left tegmen of males of Habetia species: A, H. tuta sp. nov. (Lake Chambri); B, H. pallida sp. nov. (Waris); C, H. pedala sp. nov. (Toricelli Mts); D, H. imitatrix Karny, 1912 (Bongu); E, H. dentata sp. nov. (Saidor); F, H. pilleata sp. nov. (Wau 1200 m); G–H, H. wau sp. nov. (G Nami Creek; H Wau).—Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223217" httpUri="https://zenodo.org/record/5223217/files/figure.png" pageId="11" pageNumber="212">Fig. 18A</figureCitation>
), in
<typeStatus id="AD1AE8E9AA348055FCE4305C937B6B09" box="[435,531,1591,1617]" pageId="11" pageNumber="212" type="paratype">paratype</typeStatus>
with 69 teeth, of which 8 narrow teeth at base,
<quantity id="B559FBAEAA348055F972305D95356B09" box="[1061,1117,1590,1617]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="11" pageNumber="212" unit="in" value="35.0">35 in</quantity>
main part of file, 5 at a step, 13 clearly separated at end, and 6 of decreasing size at end. Mirror behind stridulatory file with straight anterior and posterior margins and convex lateral margins; slightly longer than wide, greatest width anteriorly. Tenth abdominal tergite transverse, in midline furrowed and at fore margin with an obtuse-angular incision filled by membrane, from hind margin with a narrow elongate triangular sub-membranous area; fully sclerotized areas with apical margin obtuse-angular but with rounded angle above bases of cerci (
<figureCitation id="EA9A4ACEAA348055FE6A308092FC6A5D" box="[829,916,1771,1797]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (A–B, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); C–E, Habetia pedala sp. nov. (Toricelli Mts); F–H, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (A–C, F–G), 1 mm (D–E, H). Phallus removed in D, F–G." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="11" pageNumber="212">Fig. 8B</figureCitation>
). Cerci in about basal half roughly cylindrical, in apical half with an obtuse dorsal carina and strongly concave internal surface; at end of cercus, at dorsal margin from nearly tip, at ventral margin from beginning of apical third with a large internal projection with concave proximal surface that is before end divided into a short rounded dorsal process with narrowing margins that ends into an acute black spine, and into a small compressed ventral flap that is curved posteriorly, surpasses the dorsal spine, and carries a minute black spinule at tip (
<figureCitation id="EA9A4ACEAA348055FF3431F493A36AE1" box="[611,715,1951,1977]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: A–C, H. tuta sp. nov. (A Quelllager, B–C Töpferfluss); D–E, H. pedala sp. nov. (Toricelli Mts); F–I, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="11" pageNumber="212">Fig. 12A</figureCitation>
). Subgenital plate with convex lateral margins that form in basal two thirds a wide, dorso-ventrally compressed rim that frame the strongly convex central area that is dived from base to nearly apical third into a pair of ovoid hemispheres; apical area short and narrow, flattened and with a faint median carina, with two short styli (
<figureCitation id="EA9A4ACEAA338052FF6236D093F56D8D" box="[565,669,187,213]" captionStart="FIGURE 11" captionStartId="27.[151,250,1579,1603]" captionTargetBox="[151,1436,181,1555]" captionTargetId="figure-15@27.[151,1436,181,1555]" captionTargetPageId="27" captionText="FIGURE 11. Male subgenital plate and cerci of Habetia species in ventral view (J in ventro-apical view).—A, H. tuta sp. nov. (Rosensee); B, H. pallida sp. nov. (Ifar); C, H. pedala sp. nov. (Toricelli Mts); D, H. curvata sp. nov. (Albatros Bivak); E, H. imitatrix Karny, 1912 holotype (Bongu); F, H. dentata sp. nov. (Saidor); G, H. bivittata sp. nov. (Goroka); H, H. simbai sp. nov. (Simbai Valley); I, H. kondiu sp. nov. (Kondiu); J, H. lalibu sp. nov. (Lalibu); K, H. wau sp. nov. (Nami Creek). Scales 5 mm (A–E), 1 mm (F–K)." figureDoi="http://doi.org/10.5281/zenodo.5223195" httpUri="https://zenodo.org/record/5223195/files/figure.png" pageId="12" pageNumber="213">Fig. 11A</figureCitation>
); above the subgenital plate there is a semi-tube-shaped sclerotized phallus sheath that holds the phallus when at rest, it has the dorsal margins faintly granular and terminates into a pair of separate rounded lobes (
<figureCitation id="EA9A4ACEAA338052FCAA3768933D6C45" box="[509,597,259,285]" captionStart="FIGURE 8" captionStartId="24.[151,250,1841,1865]" captionTargetBox="[151,1436,181,1817]" captionTargetId="figure-19@24.[151,1436,181,1817]" captionTargetPageId="24" captionText="FIGURE 8. Male abdominal apex with cerci in dorsal view (A–B, F) dorso-apical view (C) and apical view (D, G); male cercus in roughly dorsal view (E); apical view of subgenital plate with styli (H).—A, Habetia imitatrix Karny, 1912 (Bongu); B, Habetia tuta sp. nov. (Quelllager); C–E, Habetia pedala sp. nov. (Toricelli Mts); F–H, Habetia pallida sp. nov. (Ifar).—The arrows aim at dorsal modifications of the subgential plate. Abbreviations: ce cercus, ph phallus, ps phallus sheath, sg subgenital plate, st stylus, ti titillator. Scales 5 mm (A–C, F–G), 1 mm (D–E, H). Phallus removed in D, F–G." figureDoi="http://doi.org/10.5281/zenodo.5223189" httpUri="https://zenodo.org/record/5223189/files/figure.png" pageId="12" pageNumber="213">Fig. 8B</figureCitation>
). Its function is unknown. Titillators simple, laterally compressed, dark or red brown, forming a pair of simple, curved sclerites; at base faintly curved, then straight, followed by an about 150–160° curve, in apical area faintly narrowed, tip rounded (
<figureCitation id="EA9A4ACEAA338052FE023720928B6C3D" box="[853,995,331,357]" captionStart="FIGURE 12" captionStartId="29.[151,250,1784,1808]" captionTargetBox="[152,1433,182,1758]" captionTargetId="figure-15@29.[151,1436,181,1760]" captionTargetPageId="29" captionText="FIGURE 12. Male titillators and other abdominal appendages of Habetia species: A–C, H. tuta sp. nov. (A Quelllager, B–C Töpferfluss); D–E, H. pedala sp. nov. (Toricelli Mts); F–I, H. imitatrix Karny, 1912 holotype.—Abdominal apex with titillators in situ (A); titillators in lateral (B), dorso-proximal (C, I), oblique ventro-apical (D, H), and strict ventro-apical views (E, G); abdominal apex with dorsal projection of subgenital plate in lateral view (F).—Abbreviations: ls lateral sclerite, ph phallus, pr apical projection of subgenital plate, ps phallus sheath, sg subgenital plate.All scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5223199" httpUri="https://zenodo.org/record/5223199/files/figure.png" pageId="12" pageNumber="213">Figs 12B–C</figureCitation>
). With a pair of spindle-shaped narrow elongate lateral sclerites.
</paragraph>
<paragraph id="721E564BAA338052FD9037F893246F41" blockId="12.[151,1437,151,717]" pageId="12" pageNumber="213">
Female. Subgenital plate rhombic with up curved lateral margins that are in basal area strongly prolonged dorsad; at apico-lateral angles with up-bent acute cones (
<figureCitation id="EA9A4ACEAA338052FE7A37DC92C26C89" box="[813,938,438,465]" captionStart="FIGURE 5" captionStartId="19.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1620]" captionTargetId="figure-15@19.[151,1436,181,1621]" captionTargetPageId="19" captionText="FIGURE 5. Female subgenital plate of Habetia species in oblique ventral (A, J), ventral (C, E, G, L), apico-ventral (B, D, G, K), apical (H), and lateral (F, I) view.—A, H. spada holotype; B–C, H. multispinulosa (B holotype, C from Dorom), D, H. pedala sp. nov. (Toricelli Mts); E–F, H. tuta sp. nov. (Keram river); G, H. curvata sp. nov. (Prauwenbivak); H–L, H. pallida sp. nov. (H–I Jutefa Bay, J Bewani, K Ifar, L Hollandia).—Scales 1 mm; A, D, G not to scale." figureDoi="http://doi.org/10.5281/zenodo.5223183" httpUri="https://zenodo.org/record/5223183/files/figure.png" pageId="12" pageNumber="213">Figs 5E–F</figureCitation>
). Ovipositor almost twice as long as body; behind basal constriction straight with dorsal valves in basal half faintly higher than in apical half; dorsal margin faintly convex but not raised (
<figureCitation id="EA9A4ACEAA338052FCB0379493566F41" box="[487,574,511,537]" captionStart="FIGURE 1" captionStartId="10.[151,250,1944,1968]" captionTargetBox="[163,1425,181,1920]" captionTargetId="figure-19@10.[162,1425,181,1920]" captionTargetPageId="10" captionText="FIGURE 1. Habetia sp. females: habitus lateral view; species with very long ovipositor: A. H. spada (Brunner, 1898) holotype; B. H. tuta sp. nov. (Keram River); C. H. pedala sp. nov. (Toricelli Mts); D. H. pallida sp. nov. (Sentani); E–F. H. multispinulosa Griffini, 1908 (E holotype, F Rigo Dorom). Scales 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5223175" httpUri="https://zenodo.org/record/5223175/files/figure.png" pageId="12" pageNumber="213">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="721E564BAA338052FD90344892086FDD" blockId="12.[151,1437,151,717]" pageId="12" pageNumber="213">
Measurements (
<specimenCount id="64A79DC2AA338052FC2C344890BE6F65" box="[379,470,547,573]" pageId="12" pageNumber="213" type="male">2 males</specimenCount>
,
<specimenCount id="64A79DC2AA338052FCB5344893206F66" box="[482,584,547,574]" pageId="12" pageNumber="213" type="female">1 female</specimenCount>
).—Body w wings: male 44–45, female 53; w/o wings: male 33–37, female 39; pronotum: male 9.2–9.5, female 10; tegmen: male 33–34, female 40; hind femur: male 23–24, female 29; ovipositor length: female 69; ovipositor height: female
<quantity id="B559FBAEAA338052FE54340092346FDD" box="[771,860,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="12" pageNumber="213" unit="mm" value="2.7">2.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="3ABB05C0AA338052FD9034E4932B6F95" pageId="12" pageNumber="213" type="etymology">
<paragraph id="721E564BAA338052FD9034E4932B6F95" blockId="12.[151,1437,151,717]" pageId="12" pageNumber="213">
<emphasis id="40D58A59AA338052FD9034E490266FF1" bold="true" box="[199,334,655,681]" pageId="12" pageNumber="213">Etymology.</emphasis>
The name of the new species refers to the cover encompassing the phallus when at rest; from the Latin adjective
<emphasis id="40D58A59AA338052FC1234D890DE6F94" box="[325,438,691,716]" italics="true" pageId="12" pageNumber="213">
tutus,
<taxonomicName id="B5A12DC8AA338052FCDD34D890DE6F94" authorityName="Ingrisch" authorityYear="2021" box="[394,438,691,716]" class="Insecta" family="Tettigoniidae" genus="Habetia" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="tuta">tuta</taxonomicName>
</emphasis>
= protected.
</paragraph>
</subSubSection>
</treatment>
</document>
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