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<mods:title id="F28F685F7CD39E57FB4CFE2C96EFA98D">The genus Dicrania LePeletier &amp; Audinet-Serville, 1828 (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactylini) from Colombia with the description of a new species</mods:title>
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<subSubSection id="36B448ADFFB60101FF3CA070FD427725" box="[151,711,1087,1113]" pageId="1" pageNumber="573" type="nomenclature">
<paragraph id="7E111B26FFB60101FF3CA070FD427725" blockId="1.[151,711,1087,1113]" box="[151,711,1087,1113]" pageId="1" pageNumber="573">
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<taxonomicName id="B9AE60A5FFB60101FF3CA070FD427725" ID-CoL="44T2" authority="LePeletier &amp; Audinet-Serville, 1828" authorityName="LePeletier &amp; Audinet-Serville" authorityYear="1828" box="[151,711,1087,1113]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="4CDAC734FFB60101FF3CA070FD427725" bold="true" box="[151,711,1087,1113]" pageId="1" pageNumber="573">
<emphasis id="4CDAC734FFB60101FF3CA070FE867725" bold="true" box="[151,259,1087,1113]" italics="true" pageId="1" pageNumber="573">Dicrania</emphasis>
LePeletier &amp; Audinet-Serville, 1828
</emphasis>
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<subSubSection id="36B448ADFFB60101FF3CA0CBFDF677E0" box="[151,627,1156,1180]" pageId="1" pageNumber="573" type="reference_group">
<paragraph id="7E111B26FFB60101FF3CA0CBFDF677E0" blockId="1.[151,1436,1156,1210]" box="[151,627,1156,1180]" pageId="1" pageNumber="573">
<taxonomicName id="B9AE60A5FFB60101FF3CA0CBFDEA77E0" authority="LePeletier &amp; Audinet-Serville, 1828" authorityName="LePeletier &amp; Audinet-Serville" authorityYear="1828" box="[151,623,1156,1180]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="4CDAC734FFB60101FF3CA0CBFF7777E7" box="[151,242,1156,1179]" italics="true" pageId="1" pageNumber="573">Dicrania</emphasis>
LePeletier &amp; Audinet-Serville, 1828
</taxonomicName>
.
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<paragraph id="7E111B26FFB60101FDD2A0CBFDF477C6" blockId="1.[151,1436,1156,1210]" pageId="1" pageNumber="573">
<typeStatus id="A115A584FFB60101FDD2A0CBFD2E77E0" box="[633,683,1156,1180]" pageId="1" pageNumber="573">Type</typeStatus>
species:
<taxonomicName id="B9AE60A5FFB60101FCA1A0CBFA9977E0" authority="LePeletier &amp; Audinet-Serville, 1828" authorityName="LePeletier &amp; Audinet-Serville" authorityYear="1828" box="[778,1308,1156,1180]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="4CDAC734FFB60101FCA1A0CBFC2677E7" box="[778,931,1156,1179]" italics="true" pageId="1" pageNumber="573">Dicrania nigra</emphasis>
LePeletier &amp; Audinet-Serville, 1828
</taxonomicName>
(subsequent designation by
<bibRefCitation id="1A3F66D7FFB60101FEC9A0EDFD8377C6" author="Blanchard, E." box="[354,518,1186,1210]" pageId="1" pageNumber="573" pagination="44" refId="ref4735" refString="Blanchard, E. (1843) Insectes. In: Audouin, J. V., Blanchard, E., Doyere, L. &amp; Milne-Edwards, H., Le regne animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux, et d'introduction a l'anatomie comparee, par Georges Cuvier. Edition accompagnee de plaches gravees, representant les types de tous les genres, les caracteres distictifs des divers groupes et les modification de structure sur lesquelles repose cette classification; par une reunion de disciples de Cuvier. Fortin, Masson et Cie, Paris, pl. 44." type="book chapter" year="1843">Blanchard 1843</bibRefCitation>
: plate 44).
</paragraph>
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<subSubSection id="36B448ADFFB60101FF3CA143FDE9765B" box="[151,620,1292,1319]" pageId="1" pageNumber="573" type="multiple">
<paragraph id="7E111B26FFB60101FF3CA143FDE9765B" blockId="1.[151,620,1292,1355]" box="[151,620,1292,1319]" pageId="1" pageNumber="573">
<heading id="2559AC4AFFB60101FF3CA143FDE9765B" bold="true" box="[151,620,1292,1319]" fontSize="11" level="1" pageId="1" pageNumber="573" reason="1">
<emphasis id="4CDAC734FFB60101FF3CA143FDE9765B" bold="true" box="[151,620,1292,1319]" pageId="1" pageNumber="573">
<taxonomicName id="B9AE60A5FFB60101FF3CA143FE38765A" ID-CoL="35QZY" authorityName="Chevrolat" authorityYear="1844" box="[151,445,1292,1318]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="4CDAC734FFB60101FF3CA143FE38765A" bold="true" box="[151,445,1292,1318]" italics="true" pageId="1" pageNumber="573">Dicrania castaneipennis</emphasis>
</taxonomicName>
species group
</emphasis>
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<subSubSection id="36B448ADFFB60101FF3CA17FFBC875CF" pageId="1" pageNumber="573" type="description">
<paragraph id="7E111B26FFB60101FF3CA17FFE987637" blockId="1.[151,620,1292,1355]" box="[151,285,1328,1355]" pageId="1" pageNumber="573">
<figureCitation id="E69507A3FFB60101FF3CA17FFE987637" box="[151,285,1328,1355]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1335,1359]" captionStartId-1="4.[151,250,1618,1642]" captionStartId-2="5.[151,250,1400,1424]" captionStartId-3="6.[151,250,1768,1792]" captionTargetBox-0="[151,1436,183,1311]" captionTargetBox-1="[156,1436,202,1592]" captionTargetBox-2="[151,1437,181,1376]" captionTargetBox-3="[165,1431,396,1745]" captionTargetId-0="figure-24@3.[151,1436,181,1311]" captionTargetId-1="figure-21@4.[151,1436,181,1594]" captionTargetId-2="figure-24@5.[151,1437,181,1376]" captionTargetId-3="figure-119@6.[151,1436,381,1745]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Dicrania diagnostic morphological characters. A, Dicrania laevipennis Moser, 1921 male, head and pronotum in dorsal view; B, D. chimu new species male paratype, scutellum in dorsal view; C, D. laevipennis male, protibia; D, D. laevipennis male, metacoxa in dorsal view; E, D. chimu male paratype, metafemora in ventral view; F, D. chimu male paratype, lateral internal view of metatarsomeres 45." captionText-1="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." captionText-2="FIGURE 3. Dicrania chimu, new species, male paratype. A, Protarsal claw; B, mesotarsal claw; C, metatarsal claw; D, apex of protibial; E, apex of mesotibia; F, apex of metatibia; G, mesotarsomere 1 in ventral view; H, mesotarsomere 1 in lateral internal view." captionText-3="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.11548013" figureDoi-1="http://doi.org/10.5281/zenodo.11548017" figureDoi-2="http://doi.org/10.5281/zenodo.11548019" figureDoi-3="http://doi.org/10.5281/zenodo.11548021" httpUri-0="https://zenodo.org/record/11548013/files/figure.png" httpUri-1="https://zenodo.org/record/11548017/files/figure.png" httpUri-2="https://zenodo.org/record/11548019/files/figure.png" httpUri-3="https://zenodo.org/record/11548021/files/figure.png" pageId="1" pageNumber="573">Figures 14</figureCitation>
</paragraph>
<paragraph id="7E111B26FFB60101FF3CA137FBC875CF" blockId="1.[151,1437,1400,1715]" pageId="1" pageNumber="573">
The following five species comprise the
<taxonomicName id="B9AE60A5FFB60101FDD1A136FCC276EE" authorityName="Chevrolat" authorityYear="1844" box="[634,839,1401,1427]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="4CDAC734FFB60101FDD1A136FD0B76EE" box="[634,654,1401,1426]" italics="true" pageId="1" pageNumber="573">D</emphasis>
.
<emphasis id="4CDAC734FFB60101FD0AA136FCC276EE" box="[673,839,1401,1426]" italics="true" pageId="1" pageNumber="573">castaneipennis</emphasis>
</taxonomicName>
species group:
<taxonomicName id="B9AE60A5FFB60101FBAFA136FF5776CB" authority="Chevrolat, 1844" authorityName="Chevrolat" authorityYear="1844" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="4CDAC734FFB60101FBAFA136FA9E76EE" box="[1028,1307,1401,1426]" italics="true" pageId="1" pageNumber="573">Dicrania castaneipennis</emphasis>
Chevrolat, 1844
</taxonomicName>
,
<taxonomicName id="B9AE60A5FFB60101FF76A1D2FE0B76CA" authority="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann, 2024" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[221,398,1436,1462]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="chimu" status="sp. nov.">
<emphasis id="4CDAC734FFB60101FF76A1D2FE0B76CA" box="[221,398,1436,1462]" italics="true" pageId="1" pageNumber="573">Dicrania chimu</emphasis>
</taxonomicName>
,
<emphasis id="4CDAC734FFB60101FE37A1D2FDA376CA" bold="true" box="[412,550,1436,1463]" pageId="1" pageNumber="573">
<taxonomicNameLabel id="57E97A4FFFB60101FE37A1D2FDA376CA" box="[412,550,1436,1463]" pageId="1" pageNumber="573" rank="species">new species</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="B9AE60A5FFB60101FD9FA1D2FC3976CB" authority="Blanchard, 1850" authorityName="Blanchard" authorityYear="1850" box="[564,956,1436,1463]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="ebenina">
<emphasis id="4CDAC734FFB60101FD9FA1D2FD7C76CA" box="[564,761,1436,1462]" italics="true" pageId="1" pageNumber="573">Dicrania ebenina</emphasis>
Blanchard, 1850
</taxonomicName>
,
<taxonomicName id="B9AE60A5FFB60101FC6CA1D2FBEF76CA" authorityName="Moser" authorityYear="1921" box="[967,1130,1436,1463]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="4CDAC734FFB60101FC6CA1D2FC5E76CA" box="[967,987,1437,1462]" italics="true" pageId="1" pageNumber="573">D</emphasis>
.
<emphasis id="4CDAC734FFB60101FC41A1D3FBEF76CA" box="[1002,1130,1436,1462]" italics="true" pageId="1" pageNumber="573">laevipennis</emphasis>
</taxonomicName>
, and
<taxonomicName id="B9AE60A5FFB60101FB0CA1D2FEAF76A7" authority="Moser, 1919" authorityName="Moser" authorityYear="1919" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="signaticollis">
<emphasis id="4CDAC734FFB60101FB0CA1D2FA1976CA" box="[1191,1436,1436,1462]" italics="true" pageId="1" pageNumber="573">Dicrania signaticollis</emphasis>
Moser, 1919
</taxonomicName>
. This species group is diagnosed by the following combination of characters (
<figureCitation id="E69507A3FFB60101FB16A18FFAA876A7" box="[1213,1325,1472,1499]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1335,1359]" captionStartId-1="4.[151,250,1618,1642]" captionStartId-2="5.[151,250,1400,1424]" captionStartId-3="6.[151,250,1768,1792]" captionTargetBox-0="[151,1436,183,1311]" captionTargetBox-1="[156,1436,202,1592]" captionTargetBox-2="[151,1437,181,1376]" captionTargetBox-3="[165,1431,396,1745]" captionTargetId-0="figure-24@3.[151,1436,181,1311]" captionTargetId-1="figure-21@4.[151,1436,181,1594]" captionTargetId-2="figure-24@5.[151,1437,181,1376]" captionTargetId-3="figure-119@6.[151,1436,381,1745]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Dicrania diagnostic morphological characters. A, Dicrania laevipennis Moser, 1921 male, head and pronotum in dorsal view; B, D. chimu new species male paratype, scutellum in dorsal view; C, D. laevipennis male, protibia; D, D. laevipennis male, metacoxa in dorsal view; E, D. chimu male paratype, metafemora in ventral view; F, D. chimu male paratype, lateral internal view of metatarsomeres 45." captionText-1="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." captionText-2="FIGURE 3. Dicrania chimu, new species, male paratype. A, Protarsal claw; B, mesotarsal claw; C, metatarsal claw; D, apex of protibial; E, apex of mesotibia; F, apex of metatibia; G, mesotarsomere 1 in ventral view; H, mesotarsomere 1 in lateral internal view." captionText-3="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.11548013" figureDoi-1="http://doi.org/10.5281/zenodo.11548017" figureDoi-2="http://doi.org/10.5281/zenodo.11548019" figureDoi-3="http://doi.org/10.5281/zenodo.11548021" httpUri-0="https://zenodo.org/record/11548013/files/figure.png" httpUri-1="https://zenodo.org/record/11548017/files/figure.png" httpUri-2="https://zenodo.org/record/11548019/files/figure.png" httpUri-3="https://zenodo.org/record/11548021/files/figure.png" pageId="1" pageNumber="573">Figs. 14</figureCitation>
): antenna with nine antennomeres and antennal club with three lamellae; pronotum with at least lateroposterior areas without punctures or setae, if surface evenly setose, then setae short and thick; males with elytra puncture usually indistinct or almost so, smooth between punctures, disc usually glabrous, when disc evidently punctate and setose, then setae are thick and homogeneously distributed; female with posterior half of elytral lateral margin not beaded abdomen of female with ventrite VI with a medial tubercle-like process; male mesotarsomere 1 with an inner spatulate tooth; metatarsomere 2 with sparse setae, medial setae shorter than the length of tarsomere.
</paragraph>
</subSubSection>
<subSubSection id="36B448ADFFB60101FF3CA34AFC88743E" pageId="1" pageNumber="573" type="multiple">
<paragraph id="7E111B26FFB60101FF3CA34AFD437463" blockId="1.[151,781,1796,1859]" box="[151,710,1796,1823]" pageId="1" pageNumber="573">
<heading id="2559AC4AFFB60101FF3CA34AFD437463" bold="true" box="[151,710,1796,1823]" fontSize="11" level="1" pageId="1" pageNumber="573" reason="1">
<emphasis id="4CDAC734FFB60101FF3CA34AFD437463" bold="true" box="[151,710,1796,1823]" pageId="1" pageNumber="573">
Key to
<taxonomicName id="B9AE60A5FFB60101FF5AA34BFD437463" ID-CoL="44T2" authority="LePeletier &amp; Audinet-Serville, 1828" authorityName="LePeletier &amp; Audinet-Serville" authorityYear="1828" box="[241,710,1796,1823]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="genus">
<emphasis id="4CDAC734FFB60101FF5AA34BFD927462" bold="true" box="[241,535,1796,1822]" italics="true" pageId="1" pageNumber="573">Dicrania castaneipennis</emphasis>
species group
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="7E111B26FFB60101FF3CA367FC88743E" blockId="1.[151,781,1796,1859]" box="[151,781,1832,1859]" pageId="1" pageNumber="573">
(Females of
<taxonomicName id="B9AE60A5FFB60101FE88A366FE0C743E" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[291,393,1832,1859]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="chimu">
<emphasis id="4CDAC734FFB60101FE88A366FEB2743E" box="[291,311,1833,1858]" italics="true" pageId="1" pageNumber="573">D</emphasis>
.
<emphasis id="4CDAC734FFB60101FEEEA367FE0C743E" box="[325,393,1832,1858]" italics="true" pageId="1" pageNumber="573">chimu</emphasis>
</taxonomicName>
and
<taxonomicName id="B9AE60A5FFB60101FE6BA366FDEE743E" authorityName="Moser" authorityYear="1919" box="[448,619,1832,1859]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="signaticollis">
<emphasis id="4CDAC734FFB60101FE6BA366FE51743E" box="[448,468,1833,1858]" italics="true" pageId="1" pageNumber="573">D</emphasis>
.
<emphasis id="4CDAC734FFB60101FE49A367FDEE743E" box="[482,619,1832,1858]" italics="true" pageId="1" pageNumber="573">signaticollis</emphasis>
</taxonomicName>
are unknown)
</paragraph>
</subSubSection>
<subSubSection id="36B448ADFFB60102FF3CA321FA19713B" lastPageId="2" lastPageNumber="574" pageId="1" pageNumber="573" type="key">
<keyStep id="C55A0383FFB60101FF3CA321FA1974DD" pageId="1" pageNumber="573">
<paragraph id="7E111B26FFB60101FF3CA321FA1974FF" blockId="1.[151,1436,1901,2041]" box="[151,1436,1901,1923]" pageId="1" pageNumber="573">
<keyLead id="C55FB813FFB60101FF3CA321FA1974FF" box="[151,1436,1901,1923]" pageId="1" pageNumber="573">
<specimenCount id="68A8D0AFFFB60101FF3CA321FE9874FF" box="[151,285,1901,1923]" count="1" pageId="1" pageNumber="573" type="male">1 Males</specimenCount>
with venter of abdomen concave; pygidium directed downward (
<figureCitation id="E69507A3FFB60101FC3BA322FC6674FF" box="[912,995,1901,1923]" captionStart="FIGURE 1" captionStartId="3.[151,250,1335,1359]" captionTargetBox="[151,1436,183,1311]" captionTargetId="figure-24@3.[151,1436,181,1311]" captionTargetPageId="3" captionText="FIGURE 1. Dicrania diagnostic morphological characters. A, Dicrania laevipennis Moser, 1921 male, head and pronotum in dorsal view; B, D. chimu new species male paratype, scutellum in dorsal view; C, D. laevipennis male, protibia; D, D. laevipennis male, metacoxa in dorsal view; E, D. chimu male paratype, metafemora in ventral view; F, D. chimu male paratype, lateral internal view of metatarsomeres 45." figureDoi="http://doi.org/10.5281/zenodo.11548013" httpUri="https://zenodo.org/record/11548013/files/figure.png" pageId="1" pageNumber="573">Figs. 1E</figureCitation>
,
<figureCitation id="E69507A3FFB60101FC46A321FBAD74FF" box="[1005,1064,1902,1923]" captionStart="FIGURE 2" captionStartId="4.[151,250,1618,1642]" captionTargetBox="[156,1436,202,1592]" captionTargetId="figure-21@4.[151,1436,181,1594]" captionTargetPageId="4" captionText="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11548017" httpUri="https://zenodo.org/record/11548017/files/figure.png" pageId="1" pageNumber="573">2BC</figureCitation>
,
<figureCitation id="E69507A3FFB60101FB9AA321FBEF74FF" box="[1073,1130,1902,1923]" captionStart="FIGURE 4" captionStartId="6.[151,250,1768,1792]" captionTargetBox="[165,1431,396,1745]" captionTargetId="figure-119@6.[151,1436,381,1745]" captionTargetPageId="6" captionText="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi="http://doi.org/10.5281/zenodo.11548021" httpUri="https://zenodo.org/record/11548021/files/figure.png" pageId="1" pageNumber="573">4BC</figureCitation>
)....................... 2
</keyLead>
</paragraph>
<paragraph id="7E111B26FFB60101FF3CA3C4FA1974DD" blockId="1.[151,1436,1901,2041]" box="[151,1436,1931,1953]" pageId="1" pageNumber="573">
<keyLead id="C55FB813FFB60101FF3CA3C4FA1974DD" box="[151,1436,1931,1953]" pageId="1" pageNumber="573">
1 Females with venter of abdomen convex; pygidium directed backward (
<figureCitation id="E69507A3FFB60101FC3FA3C4FC7C74DC" box="[916,1017,1931,1953]" captionStart="FIGURE 4" captionStartId="6.[151,250,1768,1792]" captionTargetBox="[165,1431,396,1745]" captionTargetId="figure-119@6.[151,1436,381,1745]" captionTargetPageId="6" captionText="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi="http://doi.org/10.5281/zenodo.11548021" httpUri="https://zenodo.org/record/11548021/files/figure.png" pageId="1" pageNumber="573">Fig. 4FG</figureCitation>
)................................. 6
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</paragraph>
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<keyStep id="C55A0383FFB60101FF3CA3E7FA197484" pageId="1" pageNumber="573">
<paragraph id="7E111B26FFB60101FF3CA3E7FA1974A7" blockId="1.[151,1436,1901,2041]" pageId="1" pageNumber="573">
<keyLead id="C55FB813FFB60101FF3CA3E7FA1974A7" pageId="1" pageNumber="573">
2 Pronotum and elytra uniformly setose, disc of elytra homogeneously setose;
<collectingCountry id="06B95BB6FFB60101FC6AA3E7FC7A74C2" box="[961,1023,1960,1982]" name="Brazil" pageId="1" pageNumber="573">Brazil</collectingCountry>
:
<collectingRegion id="BC6AD5C4FFB60101FBA3A3E7FBB674C1" box="[1032,1075,1960,1981]" country="Brazil" name="Para" pageId="1" pageNumber="573">Pará</collectingRegion>
......................................................................................................
<taxonomicName id="B9AE60A5FFB60101FBE2A38AFA1974A7" ID-CoL="35R36" authority="Moser, 1919" authorityName="Moser" authorityYear="1919" box="[1097,1436,1989,2011]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="573" phylum="Arthropoda" rank="species" species="signaticollis">
<emphasis id="4CDAC734FFB60101FBE2A38AFA9B74A7" bold="true" box="[1097,1310,1989,2011]" italics="true" pageId="1" pageNumber="573">Dicrania signaticollis</emphasis>
Moser, 1919
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</keyLead>
</paragraph>
<paragraph id="7E111B26FFB60101FF3CA3ACFA197484" blockId="1.[151,1436,1901,2041]" box="[151,1436,2019,2041]" pageId="1" pageNumber="573">
<keyLead id="C55FB813FFB60101FF3CA3ACFA197484" box="[151,1436,2019,2041]" pageId="1" pageNumber="573">
2 At least posterolateral areas of pronotum without setae or punctures; disc of elytra glabrous (
<figureCitation id="E69507A3FFB60101FBF0A3ACFB347484" box="[1115,1201,2019,2041]" captionStart="FIGURE 1" captionStartId="3.[151,250,1335,1359]" captionTargetBox="[151,1436,183,1311]" captionTargetId="figure-24@3.[151,1436,181,1311]" captionTargetPageId="3" captionText="FIGURE 1. Dicrania diagnostic morphological characters. A, Dicrania laevipennis Moser, 1921 male, head and pronotum in dorsal view; B, D. chimu new species male paratype, scutellum in dorsal view; C, D. laevipennis male, protibia; D, D. laevipennis male, metacoxa in dorsal view; E, D. chimu male paratype, metafemora in ventral view; F, D. chimu male paratype, lateral internal view of metatarsomeres 45." figureDoi="http://doi.org/10.5281/zenodo.11548013" httpUri="https://zenodo.org/record/11548013/files/figure.png" pageId="1" pageNumber="573">Figs. 1A</figureCitation>
,
<figureCitation id="E69507A3FFB60101FB10A3ACFB5F7484" box="[1211,1242,2019,2040]" captionStart="FIGURE 2" captionStartId="4.[151,250,1618,1642]" captionTargetBox="[156,1436,202,1592]" captionTargetId="figure-21@4.[151,1436,181,1594]" captionTargetPageId="4" captionText="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11548017" httpUri="https://zenodo.org/record/11548017/files/figure.png" pageId="1" pageNumber="573">2A</figureCitation>
,
<figureCitation id="E69507A3FFB60101FB4FA3ACFA867484" box="[1252,1283,2019,2040]" captionStart="FIGURE 4" captionStartId="6.[151,250,1768,1792]" captionTargetBox="[165,1431,396,1745]" captionTargetId="figure-119@6.[151,1436,381,1745]" captionTargetPageId="6" captionText="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi="http://doi.org/10.5281/zenodo.11548021" httpUri="https://zenodo.org/record/11548021/files/figure.png" pageId="1" pageNumber="573">4A</figureCitation>
)........... 3
</keyLead>
</paragraph>
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<keyStep id="C55A0383FFB50102FF3CA4D7FA19739B" pageId="2" pageNumber="574">
<paragraph id="7E111B26FFB50102FF3CA4D7FA1873B7" blockId="2.[151,1437,151,584]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA4D7FA1873B7" pageId="2" pageNumber="574">
3 Eyes in dorsal view large and head about 1.8 times wider than narrowest interocular distance (
<figureCitation id="E69507A3FFB50102FBC1A4D8FB6073D1" box="[1130,1253,151,173]" captionStart="FIGURE 2" captionStartId="4.[151,250,1618,1642]" captionTargetBox="[156,1436,202,1592]" captionTargetId="figure-21@4.[151,1436,181,1594]" captionTargetPageId="4" captionText="FIGURE 2. Dicrania chimu, new species, male holotype. A, Dorsal view; B, lateral view; C, ventral view; D, labels; E, head and pronotum in dorsal view; F, aedeagus in lateral view; G, aedeagus in distal view; H, pygidium in caudal view; I, antenna in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.11548017" httpUri="https://zenodo.org/record/11548017/files/figure.png" pageId="2" pageNumber="574">Fig. 2A, E, I</figureCitation>
);
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:
<collectingRegion id="BC6AD5C4FFB50102FAC8A4D7FA1173D1" box="[1379,1428,152,173]" country="Colombia" name="Meta" pageId="2" pageNumber="574">Meta</collectingRegion>
.............................................................................
<emphasis id="4CDAC734FFB50102FB2FA4FAFA1873B7" bold="true" box="[1156,1437,180,203]" pageId="2" pageNumber="574">
<taxonomicName id="B9AE60A5FFB50102FB2FA4FAFAA473B6" authority="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann, 2024" authorityName="Clavijo-Bustos &amp; Moreno &amp; Fuhrmann" authorityYear="2024" box="[1156,1313,180,202]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="chimu" status="sp. nov.">
<emphasis id="4CDAC734FFB50102FB2FA4FAFAA473B6" bold="true" box="[1156,1313,180,202]" italics="true" pageId="2" pageNumber="574">Dicrania chimu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="57E97A4FFFB50102FA8DA4FAFA1873B7" box="[1318,1437,181,203]" pageId="2" pageNumber="574" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="7E111B26FFB50102FF3CA49DFA19739B" blockId="2.[151,1437,151,584]" box="[151,1436,210,232]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA49DFA19739B" box="[151,1436,210,232]" pageId="2" pageNumber="574">
3 Eyes in dorsal view small and head about 1.5 times wider than narrowest interocular distance (
<figureCitation id="E69507A3FFB50102FBC5A49DFB46739B" box="[1134,1219,210,232]" captionStart="FIGURE 1" captionStartId="3.[151,250,1335,1359]" captionTargetBox="[151,1436,183,1311]" captionTargetId="figure-24@3.[151,1436,181,1311]" captionTargetPageId="3" captionText="FIGURE 1. Dicrania diagnostic morphological characters. A, Dicrania laevipennis Moser, 1921 male, head and pronotum in dorsal view; B, D. chimu new species male paratype, scutellum in dorsal view; C, D. laevipennis male, protibia; D, D. laevipennis male, metacoxa in dorsal view; E, D. chimu male paratype, metafemora in ventral view; F, D. chimu male paratype, lateral internal view of metatarsomeres 45." figureDoi="http://doi.org/10.5281/zenodo.11548013" httpUri="https://zenodo.org/record/11548013/files/figure.png" pageId="2" pageNumber="574">Figs. 1A</figureCitation>
,
<figureCitation id="E69507A3FFB50102FB65A49DFB6F739B" box="[1230,1258,210,231]" captionStart="FIGURE 4" captionStartId="6.[151,250,1768,1792]" captionTargetBox="[165,1431,396,1745]" captionTargetId="figure-119@6.[151,1436,381,1745]" captionTargetPageId="6" captionText="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi="http://doi.org/10.5281/zenodo.11548021" httpUri="https://zenodo.org/record/11548021/files/figure.png" pageId="2" pageNumber="574">4E</figureCitation>
)............. 4
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</paragraph>
</keyStep>
<keyStep id="C55A0383FFB50102FF3CA4BFFA19723C" pageId="2" pageNumber="574">
<paragraph id="7E111B26FFB50102FF3CA4BFFA19725E" blockId="2.[151,1437,151,584]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA4BFFA19725E" pageId="2" pageNumber="574">
4 Protarsomere 1 with a truncate inner tooth, protarsomere 2 wider than long (
<figureCitation id="E69507A3FFB50102FC07A4A0FBBD7279" box="[940,1080,239,261]" captionStart="FIGURE 1" captionStartId="3.[151,250,1335,1359]" captionTargetBox="[151,1436,183,1311]" captionTargetId="figure-24@3.[151,1436,181,1311]" captionTargetPageId="3" captionText="FIGURE 1. Dicrania diagnostic morphological characters. A, Dicrania laevipennis Moser, 1921 male, head and pronotum in dorsal view; B, D. chimu new species male paratype, scutellum in dorsal view; C, D. laevipennis male, protibia; D, D. laevipennis male, metacoxa in dorsal view; E, D. chimu male paratype, metafemora in ventral view; F, D. chimu male paratype, lateral internal view of metatarsomeres 45." figureDoi="http://doi.org/10.5281/zenodo.11548013" httpUri="https://zenodo.org/record/11548013/files/figure.png" pageId="2" pageNumber="574">Figs. 1A, CD</figureCitation>
,
<figureCitation id="E69507A3FFB50102FBEAA4BFFBF97279" box="[1089,1148,240,261]" captionStart="FIGURE 4" captionStartId="6.[151,250,1768,1792]" captionTargetBox="[165,1431,396,1745]" captionTargetId="figure-119@6.[151,1436,381,1745]" captionTargetPageId="6" captionText="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi="http://doi.org/10.5281/zenodo.11548021" httpUri="https://zenodo.org/record/11548021/files/figure.png" pageId="2" pageNumber="574">4BC</figureCitation>
);
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:
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, Amazonas;
<collectingCountry id="06B95BB6FFB50102FF49A542FEEA725F" box="[226,367,269,291]" name="French Guiana" pageId="2" pageNumber="574">French Guiana</collectingCountry>
............................................................
<taxonomicName id="B9AE60A5FFB50102FBFBA542FA19725E" ID-CoL="35R2C" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[1104,1436,268,290]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="4CDAC734FFB50102FBFBA542FA9B725E" bold="true" box="[1104,1310,268,290]" italics="true" pageId="2" pageNumber="574">Dicrania laevipennis</emphasis>
<bibRefCitation id="1A3F66D7FFB50102FA8FA542FA19725E" author="Moser, J." box="[1316,1436,269,290]" pageId="2" pageNumber="574" pagination="133 - 182" refId="ref5317" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und Sud-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182." type="journal article" year="1921">Moser, 1921</bibRefCitation>
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</keyLead>
</paragraph>
<paragraph id="7E111B26FFB50102FF3CA565FA19723C" blockId="2.[151,1437,151,584]" box="[151,1436,298,320]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA565FA19723C" box="[151,1436,298,320]" pageId="2" pageNumber="574">4 Protarsomere 1 with or without an inner tooth, protarsomere 2 longer than wide.................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="C55A0383FFB50102FF3CA508FA1972C9" pageId="2" pageNumber="574">
<paragraph id="7E111B26FFB50102FF3CA508FA197206" blockId="2.[151,1437,151,584]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA508FA197206" pageId="2" pageNumber="574">
5 Pronotum with anterolateral areas with punctures and setae sparse or indistinct, disc with two evident tufts of setae;
<collectingCountry id="06B95BB6FFB50102FAF2A508FEAF7207" name="French Guiana" pageId="2" pageNumber="574">French Guiana</collectingCountry>
;
<collectingCountry id="06B95BB6FFB50102FE9FA52AFEF77207" box="[308,370,357,379]" name="Brazil" pageId="2" pageNumber="574">Brazil</collectingCountry>
:
<collectingRegion id="BC6AD5C4FFB50102FED0A52AFE237206" box="[379,422,357,378]" country="Brazil" name="Para" pageId="2" pageNumber="574">Pará</collectingRegion>
...................................................
<taxonomicName id="B9AE60A5FFB50102FBA0A52AFA197206" ID-CoL="35QZY" authority="Chevrolat, 1844" authorityName="Chevrolat" authorityYear="1844" box="[1035,1436,357,379]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="4CDAC734FFB50102FBA0A52AFB797206" bold="true" box="[1035,1276,357,378]" italics="true" pageId="2" pageNumber="574">Dicrania castaneipennis</emphasis>
Chevrolat, 1844
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7E111B26FFB50102FF3CA5CDFA1972C9" blockId="2.[151,1437,151,584]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA5CDFA1972C9" pageId="2" pageNumber="574">
5 Pronotum with anterolateral areas densely punctate and setose, disc without delimited setal tufts;
<collectingCountry id="06B95BB6FFB50102FBD4A5CDFA8B72E4" box="[1151,1294,386,408]" name="French Guiana" pageId="2" pageNumber="574">French Guiana</collectingCountry>
;
<collectingCountry id="06B95BB6FFB50102FAB3A5CDFAD372E4" box="[1304,1366,386,408]" name="Brazil" pageId="2" pageNumber="574">Brazil</collectingCountry>
:
<collectingRegion id="BC6AD5C4FFB50102FAF5A5CDFA1D72E4" box="[1374,1432,386,408]" country="Brazil" name="Goias" pageId="2" pageNumber="574">Goiás</collectingRegion>
,
<collectingRegion id="BC6AD5C4FFB50102FF49A5EFFE8872C9" box="[226,269,416,437]" country="Brazil" name="Para" pageId="2" pageNumber="574">Pará</collectingRegion>
.....................................................................
<taxonomicName id="B9AE60A5FFB50102FBE0A5D0FA1972C9" ID-CoL="35R23" authority="Blanchard, 1850" authorityName="Blanchard" authorityYear="1850" box="[1099,1436,415,437]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="ebenina">
<emphasis id="4CDAC734FFB50102FBE0A5D0FB7D72C9" bold="true" box="[1099,1272,415,437]" italics="true" pageId="2" pageNumber="574">Dicrania ebenina</emphasis>
Blanchard, 1850
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C55A0383FFB50102FF3CA5F2FA19728C" pageId="2" pageNumber="574">
<paragraph id="7E111B26FFB50102FF3CA5F2FA1972AE" blockId="2.[151,1437,151,584]" box="[151,1436,445,467]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA5F2FA1972AE" box="[151,1436,445,467]" pageId="2" pageNumber="574">
6 Frontoclypeal suture strongly sinuate;
<collectingCountry id="06B95BB6FFB50102FDF8A5F2FD6172AF" box="[595,740,445,467]" name="French Guiana" pageId="2" pageNumber="574">French Guiana</collectingCountry>
;
<collectingCountry id="06B95BB6FFB50102FD45A5F2FCA972AF" box="[750,812,445,467]" name="Brazil" pageId="2" pageNumber="574">Brazil</collectingCountry>
:
<collectingRegion id="BC6AD5C4FFB50102FC9EA5F2FCE572AE" box="[821,864,445,466]" country="Brazil" name="Para" pageId="2" pageNumber="574">Pará</collectingRegion>
..............
<taxonomicName id="B9AE60A5FFB50102FBA0A5F2FA1972AE" ID-CoL="35QZY" authority="Chevrolat, 1844" authorityName="Chevrolat" authorityYear="1844" box="[1035,1436,445,467]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="castaneipennis">
<emphasis id="4CDAC734FFB50102FBA0A5F2FB7972AE" bold="true" box="[1035,1276,445,466]" italics="true" pageId="2" pageNumber="574">Dicrania castaneipennis</emphasis>
Chevrolat, 1844
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7E111B26FFB50102FF3CA595FA19728C" blockId="2.[151,1437,151,584]" box="[151,1436,474,496]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA595FA19728C" box="[151,1436,474,496]" pageId="2" pageNumber="574">
6 Frontoclypeal suture slightly sinuate (
<figureCitation id="E69507A3FFB50102FDE6A595FD347293" box="[589,689,474,496]" captionStart="FIGURE 4" captionStartId="6.[151,250,1768,1792]" captionTargetBox="[165,1431,396,1745]" captionTargetId="figure-119@6.[151,1436,381,1745]" captionTargetPageId="6" captionText="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi="http://doi.org/10.5281/zenodo.11548021" httpUri="https://zenodo.org/record/11548021/files/figure.png" pageId="2" pageNumber="574">Fig. 4FG</figureCitation>
)............................................................ 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="C55A0383FFB50102FF3CA5B7FA19713B" pageId="2" pageNumber="574">
<paragraph id="7E111B26FFB50102FF3CA5B7FA197157" blockId="2.[151,1437,151,584]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA5B7FA197157" pageId="2" pageNumber="574">
7 Elytra smooth or finely and sparsely punctate, punctures indistinct under magnification of 120x;
<collectingCountry id="06B95BB6FFB50102FB2BA5B8FA8A7171" box="[1152,1295,503,525]" name="French Guiana" pageId="2" pageNumber="574">French Guiana</collectingCountry>
;
<collectingCountry id="06B95BB6FFB50102FAB3A5B8FAD37171" box="[1304,1366,503,525]" name="Brazil" pageId="2" pageNumber="574">Brazil</collectingCountry>
:
<collectingRegion id="BC6AD5C4FFB50102FAF5A5B8FA1D7171" box="[1374,1432,503,525]" country="Brazil" name="Goias" pageId="2" pageNumber="574">Goiás</collectingRegion>
,
<collectingRegion id="BC6AD5C4FFB50102FF49A65AFE887156" box="[226,269,533,554]" country="Brazil" name="Para" pageId="2" pageNumber="574">Pará</collectingRegion>
.....................................................................
<taxonomicName id="B9AE60A5FFB50102FBE0A65AFA197157" ID-CoL="35R23" authority="Blanchard, 1850" authorityName="Blanchard" authorityYear="1850" box="[1099,1436,532,555]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="ebenina">
<emphasis id="4CDAC734FFB50102FBE0A65AFB7D7156" bold="true" box="[1099,1272,532,554]" italics="true" pageId="2" pageNumber="574">Dicrania ebenina</emphasis>
Blanchard, 1850
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="7E111B26FFB50102FF3CA67DFA19713B" blockId="2.[151,1437,151,584]" box="[151,1436,562,584]" pageId="2" pageNumber="574">
<keyLead id="C55FB813FFB50102FF3CA67DFA19713B" box="[151,1436,562,584]" pageId="2" pageNumber="574">
7 Elytra distinctly punctate (
<figureCitation id="E69507A3FFB50102FE4BA67DFDCD713B" box="[480,584,562,584]" captionStart="FIGURE 4" captionStartId="6.[151,250,1768,1792]" captionTargetBox="[165,1431,396,1745]" captionTargetId="figure-119@6.[151,1436,381,1745]" captionTargetPageId="6" captionText="FIGURE 4. Dicrania laevipennis Moser, 1921. A, Male from Colombia, dorsal view; B, male from Colombia, lateral view; C, male from Colombia, ventral view; D, male from Colombia, aedeagus in lateral view; E, male from Colombia, aedeagus in distal view; F, female from French Guiana, dorsal view; G, female from French Guiana, lateral view." figureDoi="http://doi.org/10.5281/zenodo.11548021" httpUri="https://zenodo.org/record/11548021/files/figure.png" pageId="2" pageNumber="574">Fig. 4FG</figureCitation>
);
<collectingCountry id="06B95BB6FFB50102FDF2A67DFD397134" box="[601,700,562,584]" name="Colombia" pageId="2" pageNumber="574">Colombia</collectingCountry>
:
<collectingRegion id="BC6AD5C4FFB50102FD6FA67CFD7D7134" box="[708,760,563,584]" country="Colombia" name="Meta" pageId="2" pageNumber="574">Meta</collectingRegion>
, Amazonas;
<collectingCountry id="06B95BB6FFB50102FCD9A67DFC7A7134" box="[882,1023,562,584]" name="French Guiana" pageId="2" pageNumber="574">French Guiana</collectingCountry>
......
<taxonomicName id="B9AE60A5FFB50102FBFBA67DFA19713B" ID-CoL="35R2C" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[1104,1436,562,584]" class="Insecta" family="Scarabaeidae" genus="Dicrania" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="574" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="4CDAC734FFB50102FBFBA67DFA9B7134" bold="true" box="[1104,1310,562,584]" italics="true" pageId="2" pageNumber="574">Dicrania laevipennis</emphasis>
<bibRefCitation id="1A3F66D7FFB50102FA8FA67CFA19713B" author="Moser, J." box="[1316,1436,562,584]" pageId="2" pageNumber="574" pagination="133 - 182" refId="ref5317" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und Sud-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182." type="journal article" year="1921">Moser, 1921</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>