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<document id="57C8C7FAB696B03D3FA258A31C930047" ID-DOI="10.1080/00222930902894096" ID-ISSN="1464-5262" ID-Zenodo-Dep="4651894" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617211343156" checkinUser="carolina" docAuthor="Hong, Yong &amp; James, Samuel W." docDate="2009" docId="03B487E85555FF9BFE75FE72FD6C56C2" docLanguage="en" docName="JNATHIST.43.21-22.1229-1256.pdf" docOrigin="Journal of Natural History 43 (21 - 22)" docSource="http://dx.doi.org/10.1080/00222930902894096" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Amynthas quinqueconvexus Hong &amp; James 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="1238" masterDocId="FF8DFF90555CFF90FF99FFA5FF95573D" masterDocTitle="Some new Korean Megascolecoid earthworms (Oligochaeta)" masterLastPageNumber="1256" masterPageNumber="1229" pageNumber="1237" updateTime="1698897605693" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="8AAE3BABB3F6FAB4DD138BFBA7806158" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EF9E5A3DC4092A112892C463281F3A8E">
<mods:title id="A460F01F0FC102FACD8EA2FBDC9ED669">Some new Korean Megascolecoid earthworms (Oligochaeta)</mods:title>
</mods:titleInfo>
<mods:name id="3C91AD04756171DA2F3B92C2275CE507" type="personal">
<mods:role id="23728F2E76CF42E3DD08000D28472773">
<mods:roleTerm id="AA1BD58E4A54450F17D192E2F7A51915">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="71799761D70D6F7C819DB0D9474511B7">Hong, Yong</mods:namePart>
</mods:name>
<mods:name id="BE81ADE977BD78AEA524BD5C144376D7" type="personal">
<mods:role id="CE7EB36157D8EF86BD7DC97560F5F72E">
<mods:roleTerm id="7071785C1FCF4E0809A9A681A39D77A4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FFD477176F0DAE3D02D59971B3EC4D28">James, Samuel W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5C517A7F532DC1C4FF893E7A868E9B0D">text</mods:typeOfResource>
<mods:relatedItem id="A124E4F085596CE2292F8750F32E31F5" type="host">
<mods:titleInfo id="2E48C0EBB7CBEE2C2C93205416C11765">
<mods:title id="0CA90C46A6F0CC0BB94AF35CA0F9F288">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="D5F07C98F2A8D9C1B72FF0E5812BF6C5">
<mods:date id="0FA551EAFD8D85B31459C2E7F4258923">2009</mods:date>
<mods:detail id="58C7AADB315A4B0D31398E9EE993C9A1" type="pubDate">
<mods:number id="E00633D610C791143BC474ED151A5E09">2009-05-21</mods:number>
</mods:detail>
<mods:detail id="584CF90470727ABA3DDC1340C42F57D4" type="volume">
<mods:number id="083E990209BE293F3E57E114BF56D32F">43</mods:number>
</mods:detail>
<mods:detail id="A0D614A95BABBCF39759E8711257F937" type="issue">
<mods:number id="3E56FE587719EF798FE30219948863F5">21 - 22</mods:number>
</mods:detail>
<mods:extent id="561CD98EE4F97B0159C570346F4A7EBC" unit="page">
<mods:start id="5B2C11D34E5BA265D0586C9FAE4361E1">1229</mods:start>
<mods:end id="926F0E32EF621630B7B7024F1A0AD508">1256</mods:end>
</mods:extent>
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<mods:location id="E6EE8CC1278522E0C74D53593B3E975F">
<mods:url id="8DACA003AAAE8CF97DF33F886945C999">http://dx.doi.org/10.1080/00222930902894096</mods:url>
</mods:location>
<mods:classification id="EFBF5E21F016BFB5990239A6352D5983">journal article</mods:classification>
<mods:identifier id="A0E190337CF1FB6D0AFFFBD0565E9B6A" type="DOI">10.1080/00222930902894096</mods:identifier>
<mods:identifier id="1F9876319255010479D4B5867C2ED1D2" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="BBA4023396DE4FE89FB3404B0C7B401A" type="Zenodo-Dep">4651894</mods:identifier>
</mods:mods>
<treatment id="03B487E85555FF9BFE75FE72FD6C56C2" LSID="urn:lsid:plazi:treatment:03B487E85555FF9BFE75FE72FD6C56C2" httpUri="http://treatment.plazi.org/id/03B487E85555FF9BFE75FE72FD6C56C2" lastPageId="11" lastPageNumber="1238" pageId="9" pageNumber="1237">
<subSubSection id="C30765755555FF99FE75FE72FCE956D2" box="[492,892,471,495]" pageId="9" pageNumber="1237" type="nomenclature">
<paragraph id="8BA236FE5555FF99FE75FE72FCE956D2" blockId="9.[492,892,471,527]" box="[492,892,471,495]" pageId="9" pageNumber="1237">
<heading id="D0EA81925555FF99FE75FE72FCE956D2" box="[492,892,471,495]" centered="true" fontSize="36" level="2" pageId="9" pageNumber="1237" reason="7">
<taxonomicName id="4C1D4D7D5555FF99FE75FE72FC8956D2" authority="Hong &amp; James, 2009" authorityName="Hong &amp; James" authorityYear="2009" box="[492,796,471,495]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="1237" phylum="Annelida" rank="species" species="quinqueconvexus" status="sp. nov.">
<emphasis id="B969EAEC5555FF99FE75FE72FC8956D2" bold="true" box="[492,796,471,495]" italics="true" pageId="9" pageNumber="1237">Amynthas quinqueconvexus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57975555FF99FCBAFE7DFCE956D2" box="[803,892,472,495]" pageId="9" pageNumber="1237" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30765755555FF9BFDE2FE52FD6C56C2" lastPageId="11" lastPageNumber="1239" pageId="9" pageNumber="1237" type="description">
<paragraph id="8BA236FE5555FF99FDE2FE52FD795532" blockId="9.[492,892,471,527]" box="[635,748,503,527]" pageId="9" pageNumber="1237">
(
<figureCitation id="13262A7B5555FF99FD1AFE52FD715532" box="[643,740,503,527]" captionStart="Figure 5" captionStartId="10.[144,211,1194,1216]" captionTargetBox="[454,793,169,1162]" captionTargetId="figure-174@10.[453,793,169,1163]" captionTargetPageId="10" captionText="Figure 5. Amynthas quinqueconvexus sp. nov.: ventral view and spermathecae." figureDoi="http://doi.org/10.5281/zenodo.4651910" httpUri="https://zenodo.org/record/4651910/files/figure.png" pageId="9" pageNumber="1237">Figure 5</figureCitation>
)
</paragraph>
<paragraph id="8BA236FE5555FF99FF4DFD9AFEFA556A" blockId="9.[212,367,575,599]" box="[212,367,575,599]" pageId="9" pageNumber="1237">
<emphasis id="B969EAEC5555FF99FF4DFD9AFEFA556A" box="[212,367,575,599]" italics="true" pageId="9" pageNumber="1237">Type material</emphasis>
</paragraph>
<paragraph id="8BA236FE5555FF99FF4DFDC2FD1D55A2" blockId="9.[212,1172,615,671]" pageId="9" pageNumber="1237">
<materialsCitation id="3B753CA35555FF99FF4DFDC2FD1D55A2" collectingDate="1971-08-05" collectorName="G. S. Kim" country="South Korea" location="Gujosa" municipality="Clitellate" pageId="9" pageNumber="1237" specimenCount="1" stateProvince="Gyeongbuk" typeStatus="holotype">
<typeStatus id="54A6885C5555FF99FF4DFDC2FED65542" box="[212,323,615,639]" pageId="9" pageNumber="1237" type="holotype">Holotype</typeStatus>
:
<collectingMunicipality id="6BC6AC845555FF99FECDFDC2FE2E5542" box="[340,443,615,639]" pageId="9" pageNumber="1237">Clitellate</collectingMunicipality>
(NIBRIV0000120466) specimen:
<collectingCountry id="F30A766E5555FF99FCD6FDCDFC0F5542" box="[847,922,616,639]" name="South Korea" pageId="9" pageNumber="1237">Korea</collectingCountry>
,
<collectingRegion id="49D9F81C5555FF99FC34FDC2FB055542" box="[941,1168,615,639]" country="South Korea" name="Gyeongbuk" pageId="9" pageNumber="1237">Gyeongsangbuk-do</collectingRegion>
,
<location id="8EC260255555FF99FF4DFD22FEBC55A2" LSID="urn:lsid:plazi:treatment:03B487E85555FF9BFE75FE72FD6C56C2:8EC260255555FF99FF4DFD22FEBC55A2" box="[212,297,647,671]" country="South Korea" municipality="Clitellate" name="Gujosa" pageId="9" pageNumber="1237" stateProvince="Gyeongbuk">Gujosa</location>
,
<date id="FFA3103E5555FF99FEADFD22FE4C55A2" box="[308,473,647,671]" pageId="9" pageNumber="1237" value="1971-08-05">
<collectingDate id="EFE7E9D65555FF99FEADFD22FE4C55A2" box="[308,473,647,671]" pageId="9" pageNumber="1237" value="1971-08-05">5 August 1971</collectingDate>
</date>
,
<collectorName id="26E853285555FF99FE7DFD22FDC455A2" box="[484,593,647,671]" pageId="9" pageNumber="1237">G.S. Kim</collectorName>
coll.
</materialsCitation>
</paragraph>
<paragraph id="8BA236FE5555FF99FF4DFD6AFED855DA" blockId="9.[212,333,719,743]" box="[212,333,719,743]" pageId="9" pageNumber="1237">
<emphasis id="B969EAEC5555FF99FF4DFD6AFED855DA" box="[212,333,719,743]" italics="true" pageId="9" pageNumber="1237">Etymology</emphasis>
</paragraph>
<paragraph id="8BA236FE5555FF99FF4DFD52FE775412" blockId="9.[212,1172,759,815]" pageId="9" pageNumber="1237">
The epithet
<taxonomicName id="4C1D4D7D5555FF99FECEFD52FDC65432" authority=", Latin" authorityName="Latin" box="[343,595,759,783]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="1237" phylum="Annelida" rank="species" species="quinqueconvexus">
<emphasis id="B969EAEC5555FF99FECEFD52FD9C5432" box="[343,521,759,783]" italics="true" pageId="9" pageNumber="1237">quinqueconvexus</emphasis>
, Latin
</taxonomicName>
for “containing five”, with regard to the appearance of the male pore region.
</paragraph>
<paragraph id="8BA236FE5555FF99FF4DFCFAFED4544A" blockId="9.[212,321,863,887]" box="[212,321,863,887]" pageId="9" pageNumber="1237">
<emphasis id="B969EAEC5555FF99FF4DFCFAFED4544A" box="[212,321,863,887]" italics="true" pageId="9" pageNumber="1237">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BA236FE5555FF99FF4DFC22FD6454E2" blockId="9.[212,1172,903,991]" pageId="9" pageNumber="1237">
Spermathecal pores in VI, VII, above lateral margin; male pores equatorial at centres of large hardened male discs, discs ovate to nearly circular; four other pairs of pads in XVII, XIX–XXI,
<quantity id="4CE59B1B5555FF99FE39FC62FD9754E2" box="[416,514,967,991]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="1237" unit="mm" value="0.23">0.23 mm</quantity>
circumference apart.
</paragraph>
<paragraph id="8BA236FE5555FF99FF4DFBAAFEC1531A" blockId="9.[212,340,1039,1063]" box="[212,340,1039,1063]" pageId="9" pageNumber="1237">
<emphasis id="B969EAEC5555FF99FF4DFBAAFEC1531A" box="[212,340,1039,1063]" italics="true" pageId="9" pageNumber="1237">Description</emphasis>
</paragraph>
<paragraph id="8BA236FE5555FF99FF4DFB9DFC2E53F2" blockId="9.[212,1172,1079,1679]" pageId="9" pageNumber="1237">
Brown dorsal pigment. Dimensions 182 by
<quantity id="4CE59B1B5555FF99FD50FB92FCB55372" box="[713,800,1079,1103]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="1237" unit="mm" value="8.5">8.5 mm</quantity>
at segment X,
<quantity id="4CE59B1B5555FF99FC48FB92FBB25372" box="[977,1063,1079,1103]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="9" pageNumber="1237" unit="mm" value="6.7">6.7 mm</quantity>
at XXX,
<quantity id="4CE59B1B5555FF99FF4DFBF2FEBF5352" box="[212,298,1111,1135]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="9" pageNumber="1237" unit="mm" value="7.1">7.1 mm</quantity>
at clitellum; body cylindrical throughout, segments 133. Setae regularly distributed around segmental equators, numbering 66 at VII, 87 at XX; 15 between male pores, regular spacing; setal formula AA: AB: YZ: ZZ = 8.5: 3: 2.5: 5 at XIII. Female pore single in XIV, 1.2-mm circle. First dorsal pore 12/13.
</paragraph>
<paragraph id="8BA236FE5555FF99FF67FB72FE0C52F2" blockId="9.[212,1172,1079,1679]" pageId="9" pageNumber="1237">
Male pores equatorial at centres of large hardened male discs, discs ovate to nearly circular, each with curved transverse groove posterior to male pore, shorter semi-circular groove anterior to male pore; paired male pads in XVII, XIX, XX, XXI; XVII, XIX slightly smaller than XVIII, decreasing in size posteriorly,
<quantity id="4CE59B1B5555FF99FBFAFA92FE955252" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="1237" unit="mm" value="0.23">0.23 mm</quantity>
circumference apart. Male pads appear to be darkened areas roughly triangular with apex towards mid-ventral line. Spermathecal pores in VI, VII, above lateral margin, inconspicuous, spermathecal pores region swelled slightly. Genital markings absent.
</paragraph>
<paragraph id="8BA236FE5555FF99FF67FA72FCD351B2" blockId="9.[212,1172,1079,1679]" pageId="9" pageNumber="1237">
Septa 5/6, 6/7, 7/8 thick, 8/9
<emphasis id="B969EAEC5555FF99FDDCFA72FDC752D2" bold="true" box="[581,594,1495,1519]" pageId="9" pageNumber="1237">–</emphasis>
9/10 absent, 10/11
<emphasis id="B969EAEC5555FF99FCBBFA72FCBA52D2" bold="true" box="[802,815,1495,1519]" pageId="9" pageNumber="1237">–</emphasis>
13/14 thick, muscular. Gizzard globular in VIII
<emphasis id="B969EAEC5555FF99FE0CFA52FE375132" bold="true" box="[405,418,1527,1551]" pageId="9" pageNumber="1237">–</emphasis>
X. Vascularized low oesophageal lamellae XIII, XIV. Intestine begins XV, lymph glands from XXVII. Typhlosole medium height from XXVII. Intestinal caeca simple, originating in XXVII, extending anteriorly to about XXIII, each consisting of a finger-shaped sac, each finger with 10 serriform outgrowths on ventral margin. Hearts X
<emphasis id="B969EAEC5555FF99FE69F9D2FE6851B2" bold="true" box="[496,509,1655,1679]" pageId="9" pageNumber="1237">–</emphasis>
XIII oesophageal, IX lateral.
</paragraph>
<caption id="DF6266765556FF9AFF09FB0FFC3953FD" ID-DOI="http://doi.org/10.5281/zenodo.4651910" ID-Zenodo-Dep="4651910" box="[144,940,1194,1216]" httpUri="https://zenodo.org/record/4651910/files/figure.png" pageId="10" pageNumber="1238" startId="10.[144,211,1194,1216]" targetBox="[454,793,169,1162]" targetPageId="10">
<paragraph id="8BA236FE5556FF9AFF09FB0FFC3953FD" blockId="10.[144,940,1194,1216]" box="[144,940,1194,1216]" pageId="10" pageNumber="1238">
Figure 5.
<taxonomicName id="4C1D4D7D5556FF9AFF6EFB0FFD9C53FD" authority="Hong &amp; James, 2009" authorityName="Hong &amp; James" authorityYear="2009" box="[247,521,1194,1216]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="1238" phylum="Annelida" rank="species" species="quinqueconvexus" status="sp. nov.">
<emphasis id="B969EAEC5556FF9AFF6EFB0FFD9C53FD" box="[247,521,1194,1216]" italics="true" pageId="10" pageNumber="1238">Amynthas quinqueconvexus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57975556FF9AFD96FB0EFDF653FD" box="[527,611,1195,1216]" pageId="10" pageNumber="1238" rank="species">sp. nov.</taxonomicNameLabel>
: ventral view and spermathecae.
</paragraph>
</caption>
<paragraph id="8BA236FE5556FF9AFF23FB5FFBB6512F" blockId="10.[144,1104,1274,1554]" pageId="10" pageNumber="1238">Ovaries in XIII. Paired spermathecae in VI, VII; each ampulla a pear-shaped small pouch with apical appendix; ducts as long as ampulla, diverticula club-shaped, stalked; no nephridia on spermathecal ducts. Male sexual system holandric, testes sacs of X enclose hearts of X, testes sacs XI enclosing seminal vesicles, hearts of XI; both pairs of sacs joined dorsally, ventrally; however, dorsal connection section of X not filled with sperm. Seminal vesicles small, two pairs in XI, XII; sperm ducts slightly muscular. Prostates just in XVIII small; thick short ducts, both glandular portions consisting of one main lobe, each lobe divided into seven or eight leaflets. Large paired muscle bands traversing coelom from mid-ventral to near mid-dorsal in XIV–XII.</paragraph>
<paragraph id="8BA236FE5556FF9AFF09F9E7FF665167" blockId="10.[144,243,1602,1626]" box="[144,243,1602,1626]" pageId="10" pageNumber="1238">
<emphasis id="B969EAEC5556FF9AFF09F9E7FF665167" box="[144,243,1602,1626]" italics="true" pageId="10" pageNumber="1238">Remarks</emphasis>
</paragraph>
<paragraph id="8BA236FE5556FF9BFF09F9CFFD6C56C2" blockId="10.[144,1104,1642,1698]" lastBlockId="11.[212,1172,167,511]" lastPageId="11" lastPageNumber="1239" pageId="10" pageNumber="1238">
The present species has a peculiar male pore region, with paired pads within XVII– XXI. The ampullae of the spermathecae are different from other Korean
<taxonomicName id="4C1D4D7D5556FF9AFC79F92FFBC5519F" baseAuthorityName="Hong and James" baseAuthorityYear="2004" box="[992,1104,1674,1698]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="1238" phylum="Annelida" rank="genus">
<emphasis id="B969EAEC5556FF9AFC79F92FFBC5519F" box="[992,1104,1674,1698]" italics="true" pageId="10" pageNumber="1238">Amynthas</emphasis>
</taxonomicName>
because there is an apical appendix. Also this species is large-bodied, but it has only two pairs of spermathecae, unlike the other large species, which have four pairs. The species has different testes sac structure from the other Korean species with spermathecae in VI and VII, and also different from the
<emphasis id="B969EAEC5557FF9BFCE6FEA2FC5B5622" box="[895,974,263,287]" italics="true" pageId="11" pageNumber="1239">morrisi</emphasis>
-group members. Compared to the
<taxonomicName id="4C1D4D7D5557FF9BFE3BFE82FDA75602" baseAuthorityName="Gates" baseAuthorityYear="1932" box="[418,562,295,319]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="1239" phylum="Annelida" rank="species" species="canaliculatus">
<emphasis id="B969EAEC5557FF9BFE3BFE82FDA75602" box="[418,562,295,319]" italics="true" pageId="11" pageNumber="1239">canaliculatus</emphasis>
</taxonomicName>
-group of species (intrasegmental spermathecal pores in VI, VII), this species has simple rather than manicate caeca (as in
<taxonomicName id="4C1D4D7D5557FF9BFC40FEE2FB1E5662" baseAuthorityName="Gates" baseAuthorityYear="1932" box="[985,1163,327,351]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="1239" phylum="Annelida" rank="species" species="canaliculatus">
<emphasis id="B969EAEC5557FF9BFC40FEE2FB1E5662" box="[985,1163,327,351]" italics="true" pageId="11" pageNumber="1239">A. canaliculatus</emphasis>
</taxonomicName>
) and has genital markings unlike
<taxonomicName id="4C1D4D7D5557FF9BFDD3FEC2FCD15642" authority="(Gates 1936)" baseAuthorityName="Gates" baseAuthorityYear="1936" box="[586,836,359,383]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="1239" phylum="Annelida" rank="species" species="rallus">
<emphasis id="B969EAEC5557FF9BFDD3FEC2FD3F5642" box="[586,682,359,383]" italics="true" pageId="11" pageNumber="1239">A. rallus</emphasis>
(
<bibRefCitation id="EF8C4B0F5557FF9BFD23FEC2FCA95642" author="Gates GE" box="[698,828,359,383]" pageId="11" pageNumber="1239" pagination="87 - 117" refId="ref10838" refString="Gates GE. 1936. On some earthworms from the Cameron Highlands, Pahang. Bull Raffles Mus, Singapore 12: 87 - 117." type="journal article" year="1936">Gates 1936</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1D4D7D5557FF9BFCE7FEC2FB055642" authority="(Gates 1936)" baseAuthorityName="Gates" baseAuthorityYear="1936" box="[894,1168,359,383]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="1239" phylum="Annelida" rank="species" species="rallidus">
<emphasis id="B969EAEC5557FF9BFCE7FEC2FC665642" box="[894,1011,359,383]" italics="true" pageId="11" pageNumber="1239">A. rallidus</emphasis>
(
<bibRefCitation id="EF8C4B0F5557FF9BFB9AFEC2FB1D5642" author="Gates GE" box="[1027,1160,359,383]" pageId="11" pageNumber="1239" pagination="87 - 117" refId="ref10838" refString="Gates GE. 1936. On some earthworms from the Cameron Highlands, Pahang. Bull Raffles Mus, Singapore 12: 87 - 117." type="journal article" year="1936">Gates 1936</bibRefCitation>
)
</taxonomicName>
. On the other hand the last two share some similarities of the testes sacs with the present species, but they are also considerably smaller (
<quantity id="4CE59B1B5557FF9BFCFAFE02FC775682" box="[867,994,423,447]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" metricValueMax="6.6" metricValueMin="3.0" pageId="11" pageNumber="1239" unit="mm" value="48.0" valueMax="66.0" valueMin="30.0">30–66 mm</quantity>
), and have red pigmentation.
<taxonomicName id="4C1D4D7D5557FF9BFEE5FE62FD6756E2" authority="(Chen 1946)" baseAuthorityName="Chen" baseAuthorityYear="1946" box="[380,754,455,479]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="11" pageNumber="1239" phylum="Annelida" rank="species" species="benigmus">
<emphasis id="B969EAEC5557FF9BFEE5FE62FDC956E2" box="[380,604,455,479]" italics="true" pageId="11" pageNumber="1239">Amynthas benigmus</emphasis>
(
<bibRefCitation id="EF8C4B0F5557FF9BFDF4FE62FD7E56E2" author="Chen Y" box="[621,747,455,479]" pageId="11" pageNumber="1239" pagination="83 - 141" refId="ref10776" refString="Chen Y. 1946. On the terrestrial Oligochaeta from Szechwan III. J West China Border Res Soc. 16: 83 - 141." type="journal article" year="1946">Chen 1946</bibRefCitation>
)
</taxonomicName>
has low simple testes sacs and only one papilla pair placed medial to the male pores.
</paragraph>
</subSubSection>
</treatment>
</document>
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