treatments-xml/data/03/B4/87/03B487C20048FFF7FF5BFC6FFC8717F0.xml
2024-06-21 12:22:17 +02:00

521 lines
71 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="20F4899C5FE3E0042CEC50B69C7A8E1D" ID-CLB-Dataset="7439" ID-DOI="10.11646/zootaxa.4934.1.1" ID-GBIF-Dataset="300bac18-55f5-4e34-90e1-7fccd02d68c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4558796" ID-ZooBank="79895443-4597-42A5-AF8A-023EACB20E10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614157470936" checkinUser="plazi" docAuthor="Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. &amp; Mathewes, Rolf W." docDate="2021" docId="03B487C20048FFF7FF5BFC6FFC8717F0" docLanguage="en" docName="zootaxa.4934.1.1.pdf" docOrigin="Zootaxa 4934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Okanopteryx macabeensis Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes 2021, new species" docType="treatment" docVersion="9" lastPageNumber="97" masterDocId="FF8DFFBA0012FF97FFCCFFE5FFAE1215" masterDocTitle="The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America" masterLastPageNumber="133" masterPageNumber="1" pageNumber="91" updateTime="1698931040708" updateUser="ExternalLinkService">
<mods:mods id="6701C21A683A4435DAA402E4AAC3F945" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AE7867DC323EEB84557A917D966166FA">
<mods:title id="A6937DEA358D023D7111032302561952">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
</mods:titleInfo>
<mods:name id="22C1A022DEFB05F13B2DE5D4D153887A" type="personal">
<mods:role id="65A49FDCECCC8E1CAC6AC24DBF55CADB">
<mods:roleTerm id="C25E3C14C257CA40882C8F1087453AEB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6CB45E6619769040CF07DF452DB7574">Archibald, Bruce</mods:namePart>
</mods:name>
<mods:name id="A8671856329BBC32516A39560039AF2C" type="personal">
<mods:role id="9CEB68A821D4D5998E18685AFDF48CD9">
<mods:roleTerm id="9EF11ACEE7BF09351E351BD11B8B1D76">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BE22F3A47D5CB91CB63397B4B18A520A">Cannings, Robert A.</mods:namePart>
<mods:nameIdentifier id="41B9B908DA6A90132930C746F1C2531A" type="ORCID">0000-0001-9491-8186</mods:nameIdentifier>
<mods:nameIdentifier id="736E8E0C1B7E57D61A028C4F05695FF5" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="60011FBB173A685E1FCC040ADBFF7848" type="personal">
<mods:role id="6A0585433B5D45C93949CE9F3352EC9F">
<mods:roleTerm id="D2A38C77436C62A2E52D60646DFC16DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4B6489F6FC1A24A6D94F6629432AE7CD">Erickson, Robert J.</mods:namePart>
<mods:nameIdentifier id="3B58AA74CFD03A5D1D8450492FA0AB36" type="ORCID">0000-0003-1162-0355</mods:nameIdentifier>
<mods:nameIdentifier id="A10B30E349D5080542502A58D531603F" type="email">1990platypus@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="6231038A171672BA86E0FF87232747FA" type="personal">
<mods:role id="68F5683B4C8198DE04EA2E383759D859">
<mods:roleTerm id="2648D642E46C170F6470374864DE15F8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EFBC52AE758F51502C78C80AB2A8736A">Bybee, Seth M.</mods:namePart>
<mods:nameIdentifier id="369EE2191F70D81E813CF8017171B905" type="ORCID">0000-0001-6224-6303</mods:nameIdentifier>
<mods:nameIdentifier id="C57B39188967699F0365F5C83DB7245E" type="email">seth.bybee@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="0835FADD9A773F1E8F4FA8C6DFD1E2D5" type="personal">
<mods:role id="E25983D441BD4F83CE701F89013A4706">
<mods:roleTerm id="BE59742F4029EF84E856FCFC9BAB9CE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4FD3ECB165B6C0AE23E678C35BA54CDC">Mathewes, Rolf W.</mods:namePart>
<mods:nameIdentifier id="FA1DB62F1D6F3092F4B7F172ED751257" type="ORCID">0000-0001-7637-199X</mods:nameIdentifier>
<mods:nameIdentifier id="EBAC5A8F74DBE1A3C3CF817950E2664E" type="email">r_mathewes@sfu.ca</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="C6118C4604B437A9BE1181E1DD0BE2F1">text</mods:typeOfResource>
<mods:relatedItem id="19A3179841EDE9C51611684B931A5188" type="host">
<mods:titleInfo id="1323FF16C780EC63723EF7C8BBE447FC">
<mods:title id="78CC53271A803DB2B3824E699191B7C9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7BBECA90435F1F66DBC1F3B8E3DC943C">
<mods:date id="F1A8204B587410E19EF3AE683A9CC236">2021</mods:date>
<mods:detail id="4218C127BF226D0C14A691F4F78ED7E9" type="pubDate">
<mods:number id="B1EB61F27A3CE780132E66FC5DA4CF37">2021-02-24</mods:number>
</mods:detail>
<mods:detail id="BEAFAD34557244BDF8DC4B893143B1AD" type="volume">
<mods:number id="EAFA55B81EFA91990AE82A9EDA3F7150">4934</mods:number>
</mods:detail>
<mods:detail id="B407816D69B3CE1D1D0174B58D76959E" type="issue">
<mods:number id="681CAE568372787EE7A3550609602086">1</mods:number>
</mods:detail>
<mods:extent id="3C8EE9926444A4F04EA079C2DDEF1344" unit="page">
<mods:start id="515C6EBD5CCC576FDC8A67990D984745">1</mods:start>
<mods:end id="47BF8C2E330BA4CCE2E33B3C1269E3A2">133</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B023F6CC2E31B891023EADAFF76C93C5">journal article</mods:classification>
<mods:identifier id="DB6F8E874B583D8D58BE2A9243D8C840" type="CLB-Dataset">7439</mods:identifier>
<mods:identifier id="8A83F2AE7633CF1F3897C9C7EABFB6FD" type="DOI">10.11646/zootaxa.4934.1.1</mods:identifier>
<mods:identifier id="8E592BAC2A1D0988138608124F39843B" type="GBIF-Dataset">300bac18-55f5-4e34-90e1-7fccd02d68c1</mods:identifier>
<mods:identifier id="AA391DE0511187E33B5A2C97B24DFA11" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F0D66FAA8F779E88330563530CFB7859" type="Zenodo-Dep">4558796</mods:identifier>
<mods:identifier id="645F0AD26CCAE8F48EDFB7016E415349" type="ZooBank">79895443-4597-42A5-AF8A-023EACB20E10</mods:identifier>
</mods:mods>
<treatment id="03B487C20048FFF7FF5BFC6FFC8717F0" ID-DOI="http://doi.org/10.5281/zenodo.4559807" ID-GBIF-Taxon="180881799" ID-Zenodo-Dep="4559807" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0" httpUri="http://treatment.plazi.org/id/03B487C20048FFF7FF5BFC6FFC8717F0" lastPageId="96" lastPageNumber="97" pageId="90" pageNumber="91">
<subSubSection id="C307655F0048FFCDFF5BFC6FFC3B11B1" box="[151,917,906,933]" pageId="90" pageNumber="91" type="nomenclature">
<paragraph id="8BA236D40048FFCDFF5BFC6FFC3B11B1" blockId="90.[151,917,906,969]" box="[151,917,906,933]" pageId="90" pageNumber="91">
<heading id="D0EA81B80048FFCDFF5BFC6FFC3B11B1" bold="true" box="[151,917,906,933]" fontSize="11" level="2" pageId="90" pageNumber="91" reason="4">
<emphasis id="B969EAC60048FFCDFF5BFC6FFC3B11B1" bold="true" box="[151,917,906,933]" pageId="90" pageNumber="91">
<taxonomicName id="4C1D4D570048FFCDFF5BFC6FFD5711B1" authority="Archibald &amp; Cannings" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[151,761,906,933]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="macabeensis" status="sp. nov.">
<emphasis id="B969EAC60048FFCDFF5BFC6FFE7A11B0" bold="true" box="[151,468,906,933]" italics="true" pageId="90" pageNumber="91">Okanopteryx macabeensis</emphasis>
Archibald &amp; Cannings
</taxonomicName>
,
<taxonomicNameLabel id="A25A57BD0048FFCDFCC9FC6EFC3B11B1" box="[773,917,906,933]" pageId="90" pageNumber="91" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C307655F0048FFCDFF5BFC4AFE8D11DC" box="[151,291,942,969]" pageId="90" pageNumber="91" type="description">
<paragraph id="8BA236D40048FFCDFF5BFC4AFE8D11DC" blockId="90.[151,917,906,969]" box="[151,291,942,969]" pageId="90" pageNumber="91">
<figureCitation id="13262A510048FFCDFF5BFC4AFEB111DC" box="[151,287,942,969]" captionStart-0="FIGURE 53" captionStart-1="FIGURE 54" captionStart-2="FIGURE 55" captionStart-3="FIGURE 56" captionStart-4="FIGURE 57" captionStartId-0="91.[151,250,1868,1892]" captionStartId-1="92.[151,250,1491,1515]" captionStartId-2="93.[151,250,1835,1859]" captionStartId-3="94.[151,250,1688,1712]" captionStartId-4="95.[151,250,1964,1988]" captionTargetBox-0="[161,1426,181,1835]" captionTargetBox-1="[157,1436,181,1465]" captionTargetBox-2="[151,1423,782,1802]" captionTargetBox-3="[160,1436,181,1656]" captionTargetBox-4="[264,1322,182,1920]" captionTargetId-0="figure-17@91.[151,1436,181,1844]" captionTargetId-1="figure-19@92.[151,1436,181,1466]" captionTargetId-2="figure-330@93.[151,1436,777,1812]" captionTargetId-3="figure-19@94.[151,1436,181,1664]" captionTargetId-4="figure-17@95.[251,1337,181,1940]" captionTargetPageId-0="91" captionTargetPageId-1="92" captionTargetPageId-2="93" captionTargetPageId-3="94" captionTargetPageId-4="95" captionText-0="FIGURE 53. Okanopteryx macabeensis holotype, GSC 141100: A, photograph; B, drawing. Paratype 4: RBCM 3087: C, photograph; D, drawing. All to scale, 5 mm." captionText-1="FIGURE 54. Okanopteryx macabeensis paratype 1, GSC 141101. A, photograph of specimen; B close-up photograph of male anal appendages; C, drawing of male anal appendages labelling abdominal segments 810 and indicating paraprocts (red) and cerci (green). A, to 5mm scale, B, C, to 3 mm scale." captionText-2="FIGURE 55. Okanopteryx macabeensis and O. fraseri. A, O. macabeensis paratype 1, drawing of body; B, holotype O. fraseri drawing of body; abdominal segments labelled; CF, wings of O. macabeensis paratype 1. A, B and CF to scales, 5 mm." captionText-3="FIGURE 56. Okanopteryx macabeensis paratype 2, F-790. A, labelled drawing of wing; B, drawing of specimen; C, photograph of specimen. A to scale, B, C to scale, both scales 5 mm." captionText-4="FIGURE 57. Okanopteryx macabeensis paratype 3, GSC 141102. A, photograph of specimen; B, drawing of upper wing in A, (vertically inverted for anterior margin on top); C, drawing of bottom wing in A. All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4558930" figureDoi-1="http://doi.org/10.5281/zenodo.4558932" figureDoi-2="http://doi.org/10.5281/zenodo.4558936" figureDoi-3="http://doi.org/10.5281/zenodo.4558938" figureDoi-4="http://doi.org/10.5281/zenodo.4558940" httpUri-0="https://zenodo.org/record/4558930/files/figure.png" httpUri-1="https://zenodo.org/record/4558932/files/figure.png" httpUri-2="https://zenodo.org/record/4558936/files/figure.png" httpUri-3="https://zenodo.org/record/4558938/files/figure.png" httpUri-4="https://zenodo.org/record/4558940/files/figure.png" pageId="90" pageNumber="91">Figs. 5357</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307655F0048FFCDFF5BFC13FD2D1620" pageId="90" pageNumber="91" type="diagnosis">
<paragraph id="8BA236D40048FFCDFF5BFC13FD2D1620" blockId="90.[151,1437,1014,2013]" pageId="90" pageNumber="91">
<emphasis id="B969EAC60048FFCDFF5BFC13FEBC1605" bold="true" box="[151,274,1014,1040]" pageId="90" pageNumber="91">Diagnosis.</emphasis>
Wings distinct from those of
<taxonomicName id="4C1D4D570048FFCDFDAFFC12FD7F1604" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[611,721,1015,1041]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B969EAC60048FFCDFDAFFC12FD7F1604" box="[611,721,1015,1041]" italics="true" pageId="90" pageNumber="91">O. fraseri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570048FFCDFD13FC12FC311605" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[735,927,1015,1041]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="jeppesenorum">
<emphasis id="B969EAC60048FFCDFD13FC12FC311605" box="[735,927,1015,1041]" italics="true" pageId="90" pageNumber="91">O. jeppesenorum</emphasis>
</taxonomicName>
as in their diagnoses; further from
<taxonomicName id="4C1D4D570048FFCDFAE3FC12FA331604" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1327,1437,1015,1041]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B969EAC60048FFCDFAE3FC12FA331604" box="[1327,1437,1015,1041]" italics="true" pageId="90" pageNumber="91">O. fraseri</emphasis>
</taxonomicName>
by abdomen/body length as in its diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="C307655F0048FFCDFF0BFBDBFCE617D5" pageId="90" pageNumber="91" type="materials_examined">
<paragraph id="8BA236D40048FFCDFF0BFBDBFCE617D5" blockId="90.[151,1437,1014,2013]" pageId="90" pageNumber="91">
<emphasis id="B969EAC60048FFCDFF0BFBDBFE98164D" bold="true" box="[199,310,1086,1112]" pageId="90" pageNumber="91">Material.</emphasis>
<materialsCitation id="3B753C890048FFCDFEF3FBDAFCE617D5" ID-GBIF-Occurrence="3069296312" collectingDate="1991-07" collectingDateMax="2000-07-10" collectingDateMin="1991-07" country="Canada" county="Geological Survey" location="Thompson Rivers University" municipality="McAbee Hoodoo Face" pageId="90" pageNumber="91" specimenCount="5" stateProvince="British Columbia" typeStatus="holotype">
<emphasis id="B969EAC60048FFCDFEF3FBDAFE08164C" box="[319,422,1087,1113]" italics="true" pageId="90" pageNumber="91">
<typeStatus id="54A688760048FFCDFEF3FBDAFE08164C" box="[319,422,1087,1113]" pageId="90" pageNumber="91" type="holotype">Holotype</typeStatus>
</emphasis>
: GSC 141100 (collector number SBA-2801) (
<figureCitation id="13262A510048FFCDFC7BFBDAFBA2164C" box="[951,1036,1086,1113]" captionStart="FIGURE 53" captionStartId="91.[151,250,1868,1892]" captionTargetBox="[161,1426,181,1835]" captionTargetId="figure-17@91.[151,1436,181,1844]" captionTargetPageId="91" captionText="FIGURE 53. Okanopteryx macabeensis holotype, GSC 141100: A, photograph; B, drawing. Paratype 4: RBCM 3087: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558930" httpUri="https://zenodo.org/record/4558930/files/figure.png" pageId="90" pageNumber="91">Fig. 53</figureCitation>
), collected at the McAbee Hoodoo Face beds, Geological Survey of
<collectingCountry id="F30A76440048FFCDFDDFFB86FDC91668" box="[531,615,1123,1149]" name="Canada" pageId="90" pageNumber="91">Canada</collectingCountry>
locality code V-016800, by SBA,
<date id="FFA310140048FFCDFC3DFB86FBDA1668" box="[1009,1140,1123,1149]" pageId="90" pageNumber="91" value="2000-07-10">
<collectingDate id="EFE7E9FC0048FFCDFC3DFB86FBDA1668" box="[1009,1140,1123,1149]" pageId="90" pageNumber="91" value="2000-07-10">10.vii.2000</collectingDate>
</date>
, housed in the Geological Survey of
<collectingCountry id="F30A76440048FFCDFEC7FB62FEF116B4" box="[267,351,1159,1185]" name="Canada" pageId="90" pageNumber="91">Canada</collectingCountry>
collections;
<emphasis id="B969EAC60048FFCDFE21FB62FDCB16B4" box="[493,613,1159,1185]" italics="true" pageId="90" pageNumber="91">
<typeStatus id="54A688760048FFCDFE21FB62FDFF16B5" box="[493,593,1159,1184]" pageId="90" pageNumber="91" type="paratype">paratype</typeStatus>
1
</emphasis>
: GSC 141101 (
<figureCitation id="13262A510048FFCDFCDEFB62FCCA16B4" box="[786,868,1158,1185]" captionStart="FIGURE 54" captionStartId="92.[151,250,1491,1515]" captionTargetBox="[157,1436,181,1465]" captionTargetId="figure-19@92.[151,1436,181,1466]" captionTargetPageId="92" captionText="FIGURE 54. Okanopteryx macabeensis paratype 1, GSC 141101. A, photograph of specimen; B close-up photograph of male anal appendages; C, drawing of male anal appendages labelling abdominal segments 810 and indicating paraprocts (red) and cerci (green). A, to 5mm scale, B, C, to 3 mm scale." figureDoi="http://doi.org/10.5281/zenodo.4558932" httpUri="https://zenodo.org/record/4558932/files/figure.png" pageId="90" pageNumber="91">Fig. 54</figureCitation>
,
<figureCitation id="13262A510048FFCDFCA3FB63FC6716B4" box="[879,969,1158,1185]" captionStart="FIGURE 55" captionStartId="93.[151,250,1835,1859]" captionTargetBox="[151,1423,782,1802]" captionTargetId="figure-330@93.[151,1436,777,1812]" captionTargetPageId="93" captionText="FIGURE 55. Okanopteryx macabeensis and O. fraseri. A, O. macabeensis paratype 1, drawing of body; B, holotype O. fraseri drawing of body; abdominal segments labelled; CF, wings of O. macabeensis paratype 1. A, B and CF to scales, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558936" httpUri="https://zenodo.org/record/4558936/files/figure.png" pageId="90" pageNumber="91">55A, 55</figureCitation>
C55F), collected at the McAbee Hoodoo Face beds, Geological Survey of
<collectingCountry id="F30A76440048FFCDFDC9FB4EFDF716D0" box="[517,601,1195,1221]" name="Canada" pageId="90" pageNumber="91">Canada</collectingCountry>
locality code V-016801 by Rod Bartlett,
<date id="FFA310140048FFCDFBD3FB4EFBD016D0" box="[1055,1150,1195,1221]" pageId="90" pageNumber="91" value="1991-07">
<collectingDate id="EFE7E9FC0048FFCDFBD3FB4EFBD016D0" box="[1055,1150,1195,1221]" pageId="90" pageNumber="91" value="1991-07">vii.1991</collectingDate>
</date>
, in the
<collectingCounty id="62C34E580048FFCDFB00FB4EFA3216D0" box="[1228,1436,1195,1221]" pageId="90" pageNumber="91">Geological Survey</collectingCounty>
of
<collectingCountry id="F30A76440048FFCDFF7AFB2AFEA416FC" box="[182,266,1231,1257]" name="Canada" pageId="90" pageNumber="91">Canada</collectingCountry>
collections;
<emphasis id="B969EAC60048FFCDFE55FB2AFDBC16FC" box="[409,530,1231,1257]" italics="true" pageId="90" pageNumber="91">
<typeStatus id="54A688760048FFCDFE55FB2AFE5316FD" box="[409,509,1231,1256]" pageId="90" pageNumber="91" type="paratype">paratype</typeStatus>
2
</emphasis>
: F-790 (
<figureCitation id="13262A510048FFCDFDB8FB2AFD6616FC" box="[628,712,1230,1257]" captionStart="FIGURE 56" captionStartId="94.[151,250,1688,1712]" captionTargetBox="[160,1436,181,1656]" captionTargetId="figure-19@94.[151,1436,181,1664]" captionTargetPageId="94" captionText="FIGURE 56. Okanopteryx macabeensis paratype 2, F-790. A, labelled drawing of wing; B, drawing of specimen; C, photograph of specimen. A to scale, B, C to scale, both scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558938" httpUri="https://zenodo.org/record/4558938/files/figure.png" pageId="90" pageNumber="91">Fig. 56</figureCitation>
), collected at the
<collectingMunicipality id="6BC6ACAE0048FFCDFC42FB2AFB2216FC" box="[910,1164,1231,1257]" pageId="90" pageNumber="91">McAbee Hoodoo Face</collectingMunicipality>
beds, date and collector unknown (this is the much less complete counterpart; the part, with the almost entire insect preserved, was last seen in the inaccessible private collection of a commercial fossil dealer), in the
<location id="8EC2600F0048FFCDFC1DFAF2FABA1724" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F0048FFCDFC1DFAF2FABA1724" box="[977,1300,1303,1329]" country="Canada" county="Geological Survey" municipality="McAbee Hoodoo Face" name="Thompson Rivers University" pageId="90" pageNumber="91" stateProvince="British Columbia">Thompson Rivers University</location>
collections;
<emphasis id="B969EAC60048FFCDFF5BFADEFEBD1740" box="[151,275,1339,1365]" italics="true" pageId="90" pageNumber="91">
<typeStatus id="54A688760048FFCDFF5BFADEFF551741" box="[151,251,1339,1364]" pageId="90" pageNumber="91" type="paratype">paratype</typeStatus>
3
</emphasis>
: GSC 141102 (collector number SBA-5928) (
<figureCitation id="13262A510048FFCDFCE0FADEFC2C1740" box="[812,898,1338,1365]" captionStart="FIGURE 57" captionStartId="95.[151,250,1964,1988]" captionTargetBox="[264,1322,182,1920]" captionTargetId="figure-17@95.[251,1337,181,1940]" captionTargetPageId="95" captionText="FIGURE 57. Okanopteryx macabeensis paratype 3, GSC 141102. A, photograph of specimen; B, drawing of upper wing in A, (vertically inverted for anterior margin on top); C, drawing of bottom wing in A. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558940" httpUri="https://zenodo.org/record/4558940/files/figure.png" pageId="90" pageNumber="91">Fig. 57</figureCitation>
), collected at the
<location id="8EC2600F0048FFCDFB98FADEFAF61740" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F0048FFCDFB98FADEFAF61740" box="[1108,1368,1339,1365]" country="Canada" county="Geological Survey" municipality="McAbee Hoodoo Face" name="McAbee Hoodoo Face" pageId="90" pageNumber="91" stateProvince="British Columbia">McAbee Hoodoo Face</location>
beds,
<location id="8EC2600F0048FFCDFF5BFABAFEC6176C" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F0048FFCDFF5BFABAFEC6176C" box="[151,360,1375,1401]" country="Canada" county="Geological Survey" municipality="McAbee Hoodoo Face" name="Geological Survey" pageId="90" pageNumber="91" stateProvince="British Columbia">Geological Survey</location>
of
<collectingCountry id="F30A76440048FFCDFE41FABAFE4F176C" box="[397,481,1375,1401]" name="Canada" pageId="90" pageNumber="91">Canada</collectingCountry>
locality code V-016800, by SBA, date unknown, in the
<location id="8EC2600F0048FFCDFB9EFABAFA8D176C" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F0048FFCDFB9EFABAFA8D176C" box="[1106,1315,1375,1401]" country="Canada" county="Geological Survey" municipality="McAbee Hoodoo Face" name="Geological Survey" pageId="90" pageNumber="91" stateProvince="British Columbia">Geological Survey</location>
of
<collectingCountry id="F30A76440048FFCDFA84FABAFA32176C" box="[1352,1436,1375,1401]" name="Canada" pageId="90" pageNumber="91">Canada</collectingCountry>
collections;
<emphasis id="B969EAC60048FFCDFEEDFA66FE321788" box="[289,412,1411,1437]" italics="true" pageId="90" pageNumber="91">
<typeStatus id="54A688760048FFCDFEEDFA66FE2B1789" box="[289,389,1411,1436]" pageId="90" pageNumber="91" type="paratype">paratype</typeStatus>
4
</emphasis>
: RBCM 3087 (
<figureCitation id="13262A510048FFCDFD83FA66FD0A1788" box="[591,676,1410,1437]" captionStart="FIGURE 53" captionStartId="91.[151,250,1868,1892]" captionTargetBox="[161,1426,181,1835]" captionTargetId="figure-17@91.[151,1436,181,1844]" captionTargetPageId="91" captionText="FIGURE 53. Okanopteryx macabeensis holotype, GSC 141100: A, photograph; B, drawing. Paratype 4: RBCM 3087: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558930" httpUri="https://zenodo.org/record/4558930/files/figure.png" pageId="90" pageNumber="91">Fig. 53</figureCitation>
), collected at the
<location id="8EC2600F0048FFCDFCBDFA66FBDC1788" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F0048FFCDFCBDFA66FBDC1788" box="[881,1138,1411,1437]" country="Canada" county="Geological Survey" municipality="McAbee Hoodoo Face" name="McAbee Hoodoo Face" pageId="90" pageNumber="91" stateProvince="British Columbia">McAbee Hoodoo Face</location>
beds by
<location id="8EC2600F0048FFCDFB10FA66FACF1788" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F0048FFCDFB10FA66FACF1788" box="[1244,1377,1411,1437]" country="Canada" county="Geological Survey" municipality="McAbee Hoodoo Face" name="John Leahy" pageId="90" pageNumber="91" stateProvince="British Columbia">John Leahy</location>
, date unknown in the
<location id="8EC2600F0048FFCDFE80FA42FE2117D4" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F0048FFCDFE80FA42FE2117D4" box="[332,399,1447,1473]" country="Canada" county="Geological Survey" municipality="McAbee Hoodoo Face" name="Royal" pageId="90" pageNumber="91" stateProvince="British Columbia">Royal</location>
<collectingRegion id="49D9F8360048FFCDFE5AFA42FDF617D4" box="[406,600,1447,1473]" country="Canada" name="British Columbia" pageId="90" pageNumber="91">British Columbia</collectingRegion>
<location id="8EC2600F0048FFCDFD93FA42FD6F17D4" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F0048FFCDFD93FA42FD6F17D4" box="[607,705,1447,1473]" country="Canada" county="Geological Survey" municipality="McAbee Hoodoo Face" name="Museum" pageId="90" pageNumber="91" stateProvince="British Columbia">Museum</location>
collections.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C307655F0048FFF7FF0BFA2FFAEC1788" lastPageId="96" lastPageNumber="97" pageId="90" pageNumber="91" type="description">
<paragraph id="8BA236D40048FFCDFF0BFA2FFB9615C8" blockId="90.[151,1437,1014,2013]" pageId="90" pageNumber="91">
<emphasis id="B969EAC60048FFCDFF0BFA2FFEF717F1" bold="true" box="[199,345,1482,1508]" pageId="90" pageNumber="91">Description.</emphasis>
<typeStatus id="54A688760048FFCDFEADFA2EFE6217F0" box="[353,460,1483,1509]" pageId="90" pageNumber="91" type="holotype">Holotype</typeStatus>
, GSC 141100, wing (
<figureCitation id="13262A510048FFCDFD08FA2EFCB717F0" box="[708,793,1482,1509]" captionStart="FIGURE 53" captionStartId="91.[151,250,1868,1892]" captionTargetBox="[161,1426,181,1835]" captionTargetId="figure-17@91.[151,1436,181,1844]" captionTargetPageId="91" captionText="FIGURE 53. Okanopteryx macabeensis holotype, GSC 141100: A, photograph; B, drawing. Paratype 4: RBCM 3087: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558930" httpUri="https://zenodo.org/record/4558930/files/figure.png" pageId="90" pageNumber="91">Fig. 53</figureCitation>
). Almost complete, missing petiole, small portion of the posterior margin in the apical quarter. Oval distal to nodus. Nodus to apex estimated
<quantity id="4CE59B310048FFCDFBA6FA0AFB7E141C" box="[1130,1232,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="90" pageNumber="91" unit="mm" value="18.4">18.4 mm</quantity>
(presumed: small portion of apex missing), nodus to pterostigma
<quantity id="4CE59B310048FFCDFD60F9F6FCBD1438" box="[684,787,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="90" pageNumber="91" unit="mm" value="14.2">14.2 mm</quantity>
; width
<quantity id="4CE59B310048FFCDFCAAF9F7FC121438" box="[870,956,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="90" pageNumber="91" unit="mm" value="6.5">6.5 mm</quantity>
wide. Hyaline except for broad, darkly infuscate fascia from level of nodus to about half distance to apex (similar to other
<taxonomicName id="4C1D4D570048FFCDFBD4F9D2FB071444" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1048,1193,1591,1617]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60048FFCDFBD4F9D2FB071444" box="[1048,1193,1591,1617]" italics="true" pageId="90" pageNumber="91">Okanopteryx</emphasis>
</taxonomicName>
species). Pterostigma broad, 2.5 times longer than wide. Region of possible oblique brace vein missing. Crossveins in postnodal, postsubnodal spaces mostly aligned. Two cells wide distal to pterostigma in region of costal space preserved except one wide at pterostigma. RARP1 space one cells wide (apical-most portion not preserved). RP1IR1 space goes to two cells wide seven cells distal to origin of IR1. IR1: origin two cells distal to origin of RP2; two cells wide close before level of pterostigma. RP2: origin 3.5 cells distal to origin of IR2; two cells wide same level as IR1RP2 space; linear, straight to slight, even curve posteriorly near margin. IR2 origin at subnodus, mostly straight, linear except near margin slightly zigzag with slight, even curve posteriad. RP3-4: origin about half way between anterior-distal corner of quadrangle, nodus; straight, linear to slight, even curve posteriorly near margin. MA straight, linear, begins slight zigzag shortly beyond level of origin of IR1, increasingly so, but missing close to margin. MP linear, slightly curved from quadrangle, missing near terminus. CuA slightly zigzagged distal to level of origin of RP2; CuAA space about six columns of three cells wide; otherwise two distal to level of nodus.
</paragraph>
<caption id="DF62665C0049FFCCFF5BF8A9FD9F159D" ID-DOI="http://doi.org/10.5281/zenodo.4558930" ID-Zenodo-Dep="4558930" httpUri="https://zenodo.org/record/4558930/files/figure.png" pageId="91" pageNumber="92" startId="91.[151,250,1868,1892]" targetBox="[161,1426,181,1835]" targetPageId="91">
<paragraph id="8BA236D40049FFCCFF5BF8A9FD9F159D" blockId="91.[151,1436,1868,1928]" pageId="91" pageNumber="92">
<emphasis id="B969EAC60049FFCCFF5BF8A9FE8E1571" bold="true" box="[151,288,1868,1892]" pageId="91" pageNumber="92">FIGURE 53.</emphasis>
<taxonomicName id="4C1D4D570049FFCCFEEAF8A9FD9E1571" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,560,1868,1892]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC60049FFCCFEEAF8A9FD9E1571" box="[294,560,1868,1892]" italics="true" pageId="91" pageNumber="92">Okanopteryx macabeensis</emphasis>
</taxonomicName>
holotype, GSC 141100: A, photograph; B, drawing. Paratype 4: RBCM 3087: C, photograph; D, drawing. All to scale, 5 mm.
</paragraph>
</caption>
<caption id="DF62665C004EFFCBFF5BFA36FD081426" ID-DOI="http://doi.org/10.5281/zenodo.4558932" ID-Zenodo-Dep="4558932" httpUri="https://zenodo.org/record/4558932/files/figure.png" pageId="92" pageNumber="93" startId="92.[151,250,1491,1515]" targetBox="[157,1436,181,1465]" targetPageId="92">
<paragraph id="8BA236D4004EFFCBFF5BFA36FD081426" blockId="92.[151,1437,1491,1587]" pageId="92" pageNumber="93">
<emphasis id="B969EAC6004EFFCBFF5BFA36FE8E17FE" bold="true" box="[151,288,1491,1515]" pageId="92" pageNumber="93">FIGURE 54.</emphasis>
<taxonomicName id="4C1D4D57004EFFCBFEEBFA36FD9F17FE" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[295,561,1491,1515]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6004EFFCBFEEBFA36FD9F17FE" box="[295,561,1491,1515]" italics="true" pageId="92" pageNumber="93">Okanopteryx macabeensis</emphasis>
</taxonomicName>
paratype 1, GSC 141101. A, photograph of specimen; B close-up photograph of male anal appendages; C, drawing of male anal appendages labelling abdominal segments 810 and indicating paraprocts (red) and cerci (green). A, to 5mm scale, B, C, to 3 mm scale.
</paragraph>
</caption>
<paragraph id="8BA236D4004EFFCAFF0BF987FA2510E4" blockId="92.[151,1436,1633,2020]" lastBlockId="93.[151,1437,151,753]" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">
<materialsCitation id="3B753C89004EFFCAFF0BF987FA2910E4" ID-GBIF-Occurrence="3069296331" collectionCode="MA, MP, CuA" lastPageId="93" lastPageNumber="94" location="Same region" pageId="92" pageNumber="93" specimenCode="RP1, RP2, RP3-4, RP3- 4" specimenCount="3" stateProvince="Para" typeStatus="paratype">
<typeStatus id="54A68876004EFFCBFF0BF987FE871469" box="[199,297,1634,1660]" pageId="92" pageNumber="93" type="paratype">Paratype</typeStatus>
1, GSC 141101, male (
<figureCitation id="13262A51004EFFCBFDE2F984FD251469" box="[558,651,1633,1660]" captionStart="FIGURE 54" captionStartId="92.[151,250,1491,1515]" captionTargetBox="[157,1436,181,1465]" captionTargetId="figure-19@92.[151,1436,181,1466]" captionTargetPageId="92" captionText="FIGURE 54. Okanopteryx macabeensis paratype 1, GSC 141101. A, photograph of specimen; B close-up photograph of male anal appendages; C, drawing of male anal appendages labelling abdominal segments 810 and indicating paraprocts (red) and cerci (green). A, to 5mm scale, B, C, to 3 mm scale." figureDoi="http://doi.org/10.5281/zenodo.4558932" httpUri="https://zenodo.org/record/4558932/files/figure.png" pageId="92" pageNumber="93">Figs. 54</figureCitation>
,
<figureCitation id="13262A51004EFFCBFD5BF984FD5F1469" box="[663,753,1633,1660]" captionStart="FIGURE 55" captionStartId="93.[151,250,1835,1859]" captionTargetBox="[151,1423,782,1802]" captionTargetId="figure-330@93.[151,1436,777,1812]" captionTargetPageId="93" captionText="FIGURE 55. Okanopteryx macabeensis and O. fraseri. A, O. macabeensis paratype 1, drawing of body; B, holotype O. fraseri drawing of body; abdominal segments labelled; CF, wings of O. macabeensis paratype 1. A, B and CF to scales, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558936" httpUri="https://zenodo.org/record/4558936/files/figure.png" pageId="92" pageNumber="93">55A, 55</figureCitation>
C55F). Complete body, total length
<quantity id="4CE59B31004EFFCBFB5CF984FB591469" box="[1168,1271,1633,1660]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="92" pageNumber="93" unit="mm" value="37.7">37.7 mm</quantity>
; abdomen preserved in ventral aspect, at least distal to segment 4; thorax and head in dorsal aspect; portions of four legs, portions of all four wings. Head (
<figureCitation id="13262A51004EFFCBFE67F94CFDBB14D1" box="[427,533,1705,1732]" captionStart="FIGURE 10" captionStartId="22.[151,250,1598,1622]" captionTargetBox="[327,1259,184,1571]" captionTargetId="figure-19@22.[321,1266,181,1574]" captionTargetPageId="22" captionText="FIGURE 10. Heads of Zygoptera (A, B) and comparative Anisoptera, Anisozygoptera, Tarsophlebiidae, Stenophlebiidae (C H); and Cephalozygoptera (IQ).A, Zygoptera, Heteropodagrion superbum Ris redrawn from Fig. 318 of Garrison et al. (2010); B, Zygoptera, Minagrion ribeiroi (Santos) redrawn from Fig. 1842 of Garrison et al. (2010); C, Anisoptera, Gomphidae, Cyanogomphus waltheri Selys redrawn from Fig. 383 of Garrison et al. (2006); D, Anisozygoptera, Epiophlebia laidlawi Tillyard redrawn from Fig. 2 of Büsse (2016); E, Anisoptera, Aeshnidae, Limnetron antarcticum Förster redrawn from Fig. 16 of Garrison et al. (2006); F, Stenophlebia sp. redrawn from Tillyard &amp; Fraser 1940, Fig, 4C; G, Tarsophlebiidae, Tarsophlebia eximia Hagen redrawn from Fleck et al. (2004) Fig. 1C; H, photograph of Tarsophlebia eximia in MCZ collections; I, Sieblosia jucunda redrawn from Hagen (1858, Plate 24, Fig. 1); J, Cephalozygoptera incertae sedis SR 14-003-004; K, Phenacolestes parallelus UCM-4545; L, Okanopteryx macabeensis, GSC 141101; M, Okanagrion beardi paratype 1, RBCM P1546; N, Okanopteryx fraseri GSC 141104; O, Okanopteryx macabeensis F-790; P, Dysagrion fredericii redrawn from Scudder 1890, plate 6, Fig. 10; Q, Dysagrion fredericii redrawn from Scudder 1890, plate 6, Fig. 2. G and H, I, J to O to scales, all 5 mm; others, originals without scales." figureDoi="http://doi.org/10.5281/zenodo.4558820" httpUri="https://zenodo.org/record/4558820/files/figure.png" pageId="92" pageNumber="93">Fig. 10 L</figureCitation>
,
<tableCitation id="C69F036F004EFFCBFDEDF94CFDDD14D6" box="[545,627,1705,1731]" captionStart="TABLE 1" captionStartId="23.[151,239,485,511]" captionTargetPageId="23" captionText="TABLE 1. Head measurements of Cephalozygoptera and measurement ratios: WH, width of head; WBE, width between eyes; WE, width of one eye; LE, length of one eye; LH, length of head. See Fig. 10 for head images and text for measurement details." httpUri="http://table.plazi.org/id/DF62665C0005FF80FF5BFE00FC7F1036" pageId="92" pageNumber="93" tableUuid="DF62665C0005FF80FF5BFE00FC7F1036">Table 1</tableCitation>
). In dorsal view about
<quantity id="4CE59B31004EFFCBFCB5F94CFC6114D1" box="[889,975,1705,1732]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="92" pageNumber="93" unit="mm" value="3.7">3.7 mm</quantity>
across eyes at widest point; approximate length from anterior edge of antefrons to posterior of occiput about
<quantity id="4CE59B31004EFFCBFC4BF928FC4E14FD" box="[903,992,1741,1768]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="92" pageNumber="93" unit="mm" value="1.5">1.5 mm</quantity>
; at level of hind ocelli distance between medial margins of eyes about 1.0 mm, width of eye about
<quantity id="4CE59B31004EFFCBFCFEF914FC221519" box="[818,908,1777,1804]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="92" pageNumber="93" unit="mm" value="1.2">1.2 mm</quantity>
, length
<quantity id="4CE59B31004EFFCBFC2BF914FBEF1519" box="[999,1089,1777,1804]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="92" pageNumber="93" unit="mm" value="1.8">1.8 mm</quantity>
. Thorax. Dorsum of prothorax oval,
<quantity id="4CE59B31004EFFCBFF19F8F0FE851525" box="[213,299,1813,1840]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="92" pageNumber="93" unit="mm" value="2.1">2.1 mm</quantity>
wide,
<quantity id="4CE59B31004EFFCBFEBBF8F0FE631525" box="[375,461,1813,1840]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="92" pageNumber="93" unit="mm" value="1.1">1.1 mm</quantity>
long; pterothorax about 6.0 mm long. Legs. Lengths of leg segments unclear, not measured. Wings. Left fore(?)wing. Proximal portion to presumed (based on
<typeStatus id="54A68876004EFFCBFC12F8DCFBEC1541" box="[990,1090,1849,1876]" pageId="92" pageNumber="93" type="holotype">holotype</typeStatus>
) half distance between nodus, apex. Darkly infuscate from nodus to distal portion preserved, hyaline proximal to nodus. Maximum width
<quantity id="4CE59B31004EFFCBFAB5F8B8FF681589" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="92" pageNumber="93" unit="mm" value="6.9">6.9 mm</quantity>
. Crossveins aligned in postnodal, postsubnodal spaces (portion preserved).
<specimenCode id="DBBB9EAF004EFFCBFBE8F864FBFD158E" box="[1060,1107,1921,1947]" collectionCode="RP" pageId="92" pageNumber="93">RP1</specimenCode>
IR1 space goes to two cells wide three cells distal to origin of IR1. IR1: origin two cells distal to origin of
<specimenCode id="DBBB9EAF004EFFCBFBCFF840FB9915AA" box="[1027,1079,1957,1983]" collectionCode="RP" pageId="92" pageNumber="93">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF004EFFCBFB8DF840FBDB15AA" box="[1089,1141,1957,1983]" collectionCode="RP" pageId="92" pageNumber="93">RP2</specimenCode>
: origin three cells distal to origin of IR2. IR2 origin very close to subnodus; straight, linear.
<specimenCode id="DBBB9EAF004EFFCBFCA7F82CFC1915F6" box="[875,951,1993,2019]" collectionCode="RP" pageId="92" pageNumber="93">RP3-4</specimenCode>
: origin not preserved; linear, straight where reserved.
<collectionCode id="ED0CAE11004FFFCAFEC8FF72FE9E12A4" box="[260,304,151,177]" pageId="93" pageNumber="94">MA</collectionCode>
linear, slightly curved from quadrangle to zigzag increasingly from about level of origin of IR1.
<collectionCode id="ED0CAE11004FFFCAFAB9FF72FA3212A4" box="[1397,1436,151,177]" country="0" name="Mohonk Preserve, Inc." pageId="93" pageNumber="94">MP</collectionCode>
linear, slightly curved from quadrangle.
<collectionCode id="ED0CAE11004FFFCAFD91FF5EFD3E12C0" box="[605,656,187,213]" pageId="93" pageNumber="94">CuA</collectionCode>
with slight zigzag beyond level of nodus, increasing beyond level of origin of
<specimenCode id="DBBB9EAF004FFFCAFF32FF3AFE9F12EC" box="[254,305,223,249]" collectionCode="RP" pageId="93" pageNumber="94">RP2</specimenCode>
, CuAA space about 10 columns with 3 cells wide, the rest two distal to level of origin of
<collectingRegion id="49D9F836004FFFCAFAD5FF3AFAEE12EC" box="[1305,1344,223,249]" country="Brazil" name="Para" pageId="93" pageNumber="94">PR</collectingRegion>
3-4. Left hind(?) wing.
<location id="8EC2600F004FFFCAFEFBFEE6FE681308" LSID="urn:lsid:plazi:treatment:03B487C20048FFF7FF5BFC6FFC8717F0:8EC2600F004FFFCAFEFBFEE6FE681308" box="[311,454,259,285]" name="Same region" pageId="93" pageNumber="94" stateProvince="Para">Same region</location>
preserved as left fore(?)wing, but more complete in antenodal region. Petiole partially preserved. Shape as for left fore(?)wing. Width
<quantity id="4CE59B31004FFFCAFD66FEC2FCAD1354" box="[682,771,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="93" pageNumber="94" unit="mm" value="6.9">6.9 mm</quantity>
. Hyaline, infuscate in same regions, but infuscation notably lighter. Crossveins in postnodal, postsubnodal spaces aligned.
<specimenCode id="DBBB9EAF004FFFCAFC84FEAEFCD91370" box="[840,887,331,357]" collectionCode="RP" pageId="93" pageNumber="94">RP1</specimenCode>
IR1 space goes to two cells wide four cells distal to origin of IR1. IR1: origin three cells distal to origin of
<specimenCode id="DBBB9EAF004FFFCAFCDCFE8AFCEA139C" box="[784,836,367,393]" collectionCode="RP" pageId="93" pageNumber="94">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF004FFFCAFC82FE8AFC2C139C" box="[846,898,367,393]" collectionCode="RP" pageId="93" pageNumber="94">RP2</specimenCode>
: origin four cells distal to origin of IR1; straight, linear. IR2 origin very close to subnodus; straight, linear.
<specimenCode id="DBBB9EAF004FFFCAFCD0FE76FCC913B8" box="[796,871,403,429]" collectionCode="RP" pageId="93" pageNumber="94">RP3-4</specimenCode>
: origin at level of Ax2, half way between anteriordistal corner of quadrangle, nodus, in middle third between arculus, nodus.
<collectionCode id="ED0CAE11004FFFCAFC36FE52FB8813C4" box="[1018,1062,439,465]" pageId="93" pageNumber="94">MA</collectionCode>
slightly zigzagged from level of origin of IR1, increasingly so distal to it.
<collectionCode id="ED0CAE11004FFFCAFD93FE3EFD2813E0" box="[607,646,475,501]" country="0" name="Mohonk Preserve, Inc." pageId="93" pageNumber="94">MP</collectionCode>
linear, slightly curved from quadrangle through preserved portion near terminus.
<collectionCode id="ED0CAE11004FFFCAFECBFE1AFE94100C" box="[263,314,511,537]" pageId="93" pageNumber="94">CuA</collectionCode>
linear from level of quadrangle until slight zigzag from nodus level increasingly so after level of origin of
<specimenCode id="DBBB9EAF004FFFCAFF12FDC6FEBF1028" box="[222,273,547,573]" collectionCode="RP" pageId="93" pageNumber="94">RP2</specimenCode>
; CuAA space with about ten columns of three cells at widest, otherwise two from level of origin of
<specimenCode id="DBBB9EAF004FFFCAFAAFFDC6FF061074" collectionCode="RP" pageId="93" pageNumber="94">RP3- 4</specimenCode>
. Right wings fragmentary, preserved portions do not disagree with left wings; presumed hind(?) wing with same lighter infuscation as left hind(?) wing. Presumed fore(?)wing colouration as in
<typeStatus id="54A68876004FFFCAFBDAFD8EFBD21090" box="[1046,1148,619,645]" pageId="93" pageNumber="94" type="holotype">holotype</typeStatus>
. Abdomen approximately 20.0 mm long. Anal appendages: cerci
<quantity id="4CE59B31004FFFCAFD90FD6AFD1B10BC" box="[604,693,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="93" pageNumber="94" unit="mm" value="1.3">1.3 mm</quantity>
long,
<quantity id="4CE59B31004FFFCAFCCDFD6AFCF710BC" box="[769,857,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="93" pageNumber="94" unit="mm" value="0.4">0.4 mm</quantity>
wide, evenly curved medially, the rounded apices nearly touching; paraprocts about
<quantity id="4CE59B31004FFFCAFDD5FD56FDDE10D8" box="[537,624,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="93" pageNumber="94" unit="mm" value="0.8">0.8 mm</quantity>
long,
<quantity id="4CE59B31004FFFCAFD7BFD56FCA010D8" box="[695,782,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="93" pageNumber="94" unit="mm" value="0.4">0.4 mm</quantity>
wide at base tapering to
<quantity id="4CE59B31004FFFCAFBE6FD56FB2F10D8" box="[1066,1153,691,717]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="93" pageNumber="94" unit="mm" value="0.2">0.2 mm</quantity>
at apex, extending about 0.7 the length of cerci; medial side straight, lateral side curved medially; apex slightly hooked medially, rounded
</materialsCitation>
.
</paragraph>
<caption id="DF62665C004FFFCAFF5BF8CEFAC61572" ID-DOI="http://doi.org/10.5281/zenodo.4558936" ID-Zenodo-Dep="4558936" httpUri="https://zenodo.org/record/4558936/files/figure.png" pageId="93" pageNumber="94" startId="93.[151,250,1835,1859]" targetBox="[151,1423,782,1802]" targetPageId="93">
<paragraph id="8BA236D4004FFFCAFF5BF8CEFAC61572" blockId="93.[151,1437,1835,1896]" pageId="93" pageNumber="94">
<emphasis id="B969EAC6004FFFCAFF5BF8CEFE8E1556" bold="true" box="[151,288,1835,1859]" pageId="93" pageNumber="94">FIGURE 55.</emphasis>
<taxonomicName id="4C1D4D57004FFFCAFEEAF8CEFD9E1556" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,560,1835,1859]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6004FFFCAFEEAF8CEFD9E1556" box="[294,560,1835,1859]" italics="true" pageId="93" pageNumber="94">Okanopteryx macabeensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D57004FFFCAFDADF8C9FD6A1556" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[609,708,1835,1859]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B969EAC6004FFFCAFDADF8C9FD6A1556" box="[609,708,1835,1859]" italics="true" pageId="93" pageNumber="94">O. fraseri</emphasis>
</taxonomicName>
. A,
<taxonomicName id="4C1D4D57004FFFCAFD21F8C9FC231556" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[749,909,1835,1859]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6004FFFCAFD21F8C9FC231556" box="[749,909,1835,1859]" italics="true" pageId="93" pageNumber="94">O. macabeensis</emphasis>
</taxonomicName>
paratype 1, drawing of body; B, holotype
<taxonomicName id="4C1D4D57004FFFCAFAF6F8C9FA331556" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1338,1437,1835,1859]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="fraseri">
<emphasis id="B969EAC6004FFFCAFAF6F8C9FA331556" box="[1338,1437,1835,1859]" italics="true" pageId="93" pageNumber="94">O. fraseri</emphasis>
</taxonomicName>
drawing of body; abdominal segments labelled; CF, wings of
<taxonomicName id="4C1D4D57004FFFCAFCD8F8B5FC1A1572" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[788,948,1871,1895]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6004FFFCAFCD8F8B5FC1A1572" box="[788,948,1871,1895]" italics="true" pageId="93" pageNumber="94">O. macabeensis</emphasis>
</taxonomicName>
paratype 1. A, B and CF to scales, 5 mm.
</paragraph>
</caption>
<caption id="DF62665C004CFFC9FF5BF97DFCBA14C1" ID-DOI="http://doi.org/10.5281/zenodo.4558938" ID-Zenodo-Dep="4558938" httpUri="https://zenodo.org/record/4558938/files/figure.png" pageId="94" pageNumber="95" startId="94.[151,250,1688,1712]" targetBox="[160,1436,181,1656]" targetPageId="94">
<paragraph id="8BA236D4004CFFC9FF5BF97DFCBA14C1" blockId="94.[151,1436,1688,1748]" pageId="94" pageNumber="95">
<emphasis id="B969EAC6004CFFC9FF5BF97DFE8C14A5" bold="true" box="[151,290,1688,1712]" pageId="94" pageNumber="95">FIGURE 56.</emphasis>
<taxonomicName id="4C1D4D57004CFFC9FEE5F97DFD9B14A5" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[297,565,1688,1712]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6004CFFC9FEE5F97DFD9B14A5" box="[297,565,1688,1712]" italics="true" pageId="94" pageNumber="95">Okanopteryx macabeensis</emphasis>
</taxonomicName>
paratype 2, F-790. A, labelled drawing of wing; B, drawing of specimen; C, photograph of specimen. A to scale, B, C to scale, both scales 5 mm.
</paragraph>
</caption>
<paragraph id="8BA236D4004CFFF7FF0BF8E6FEF81194" blockId="94.[151,1437,1794,2037]" lastBlockId="96.[151,1437,150,1509]" lastPageId="96" lastPageNumber="97" pageId="94" pageNumber="95">
<materialsCitation id="3B753C89004CFFF7FF0BF8E6FEFC1194" ID-GBIF-Occurrence="3069296334" collectionCode="MA, MP, CuA" lastPageId="96" lastPageNumber="97" pageId="94" pageNumber="95" specimenCode="RP1, RP2, RP3-4" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A68876004CFFC9FF0BF8E6FE871508" box="[199,297,1795,1821]" pageId="94" pageNumber="95" type="paratype">Paratype</typeStatus>
2, F-790 (
<figureCitation id="13262A51004CFFC9FE68F8E7FE551508" box="[420,507,1794,1821]" captionStart="FIGURE 55" captionStartId="93.[151,250,1835,1859]" captionTargetBox="[151,1423,782,1802]" captionTargetId="figure-330@93.[151,1436,777,1812]" captionTargetPageId="93" captionText="FIGURE 55. Okanopteryx macabeensis and O. fraseri. A, O. macabeensis paratype 1, drawing of body; B, holotype O. fraseri drawing of body; abdominal segments labelled; CF, wings of O. macabeensis paratype 1. A, B and CF to scales, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558936" httpUri="https://zenodo.org/record/4558936/files/figure.png" pageId="94" pageNumber="95">Fig. 55</figureCitation>
). Head; thorax, parts of five legs, both forewings missing the apical quarters, the proximal portions of both hind wings; the abdomen to part of segment 3; preserved in dorsal aspect. Head (
<figureCitation id="13262A51004CFFC9FAA3F8C3FF7C1570" captionStart="FIGURE 10" captionStartId="22.[151,250,1598,1622]" captionTargetBox="[327,1259,184,1571]" captionTargetId="figure-19@22.[321,1266,181,1574]" captionTargetPageId="22" captionText="FIGURE 10. Heads of Zygoptera (A, B) and comparative Anisoptera, Anisozygoptera, Tarsophlebiidae, Stenophlebiidae (C H); and Cephalozygoptera (IQ).A, Zygoptera, Heteropodagrion superbum Ris redrawn from Fig. 318 of Garrison et al. (2010); B, Zygoptera, Minagrion ribeiroi (Santos) redrawn from Fig. 1842 of Garrison et al. (2010); C, Anisoptera, Gomphidae, Cyanogomphus waltheri Selys redrawn from Fig. 383 of Garrison et al. (2006); D, Anisozygoptera, Epiophlebia laidlawi Tillyard redrawn from Fig. 2 of Büsse (2016); E, Anisoptera, Aeshnidae, Limnetron antarcticum Förster redrawn from Fig. 16 of Garrison et al. (2006); F, Stenophlebia sp. redrawn from Tillyard &amp; Fraser 1940, Fig, 4C; G, Tarsophlebiidae, Tarsophlebia eximia Hagen redrawn from Fleck et al. (2004) Fig. 1C; H, photograph of Tarsophlebia eximia in MCZ collections; I, Sieblosia jucunda redrawn from Hagen (1858, Plate 24, Fig. 1); J, Cephalozygoptera incertae sedis SR 14-003-004; K, Phenacolestes parallelus UCM-4545; L, Okanopteryx macabeensis, GSC 141101; M, Okanagrion beardi paratype 1, RBCM P1546; N, Okanopteryx fraseri GSC 141104; O, Okanopteryx macabeensis F-790; P, Dysagrion fredericii redrawn from Scudder 1890, plate 6, Fig. 10; Q, Dysagrion fredericii redrawn from Scudder 1890, plate 6, Fig. 2. G and H, I, J to O to scales, all 5 mm; others, originals without scales." figureDoi="http://doi.org/10.5281/zenodo.4558820" httpUri="https://zenodo.org/record/4558820/files/figure.png" pageId="94" pageNumber="95">Fig. 10 O</figureCitation>
,
<tableCitation id="C69F036F004CFFC9FF13F8AFFE9C1570" box="[223,306,1866,1893]" captionStart="TABLE 1" captionStartId="23.[151,239,485,511]" captionTargetPageId="23" captionText="TABLE 1. Head measurements of Cephalozygoptera and measurement ratios: WH, width of head; WBE, width between eyes; WE, width of one eye; LE, length of one eye; LH, length of head. See Fig. 10 for head images and text for measurement details." httpUri="http://table.plazi.org/id/DF62665C0005FF80FF5BFE00FC7F1036" pageId="94" pageNumber="95" tableUuid="DF62665C0005FF80FF5BFE00FC7F1036">Table 1</tableCitation>
). Relatively well preserved; in dorsal view about
<quantity id="4CE59B31004CFFC9FCA3F8AEFC691570" box="[879,967,1867,1893]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="94" pageNumber="95" unit="mm" value="3.8">3.8 mm</quantity>
across eyes at widest point; approximate length from anterior edge of antefrons to posterior of occiput about
<quantity id="4CE59B31004CFFC9FC4BF88AFC4E159C" box="[903,992,1903,1929]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="94" pageNumber="95" unit="mm" value="1.7">1.7 mm</quantity>
; at level of hind ocelli distance between medial margins of eyes about
<quantity id="4CE59B31004CFFC9FE2AF876FD9015B8" box="[486,574,1939,1965]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="94" pageNumber="95" unit="mm" value="1.1">1.1 mm</quantity>
, width of eye about
<quantity id="4CE59B31004CFFC9FCEDF876FCD615B8" box="[801,888,1939,1965]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="94" pageNumber="95" unit="mm" value="1.3">1.3 mm</quantity>
, length
<quantity id="4CE59B31004CFFC9FC1DF876FB8715B8" box="[977,1065,1939,1965]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="94" pageNumber="95" unit="mm" value="2.2">2.2 mm</quantity>
. Thorax. Prothorax roughly oval, about
<quantity id="4CE59B31004CFFC9FF11F853FE9D15C4" box="[221,307,1974,2001]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="94" pageNumber="95" unit="mm" value="1.5">1.5 mm</quantity>
wide, 1.0 mm long; pterothorax slightly deformed, especially posterior part, about
<quantity id="4CE59B31004CFFC9FB10F853FA9C15C4" box="[1244,1330,1974,2001]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="94" pageNumber="95" unit="mm" value="5.9">5.9 mm</quantity>
long,
<quantity id="4CE59B31004CFFC9FAB5F852FF6D15E0" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="94" pageNumber="95" unit="mm" value="3.2">3.2 mm</quantity>
wide. Legs. Left profemur 3.0 mm long, tibia
<quantity id="4CE59B31004CFFC9FD1EF83EFC8715E0" box="[722,809,2011,2037]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="94" pageNumber="95" unit="mm" value="3.8">3.8 mm</quantity>
(setae
<quantity id="4CE59B31004CFFC9FCB5F83FFC6115E0" box="[889,975,2010,2037]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="94" pageNumber="95" unit="mm" value="0.6">0.6 mm</quantity>
long, their bases separated by about half their length), tarsus
<quantity id="4CE59B310072FFF7FEB3FF73FE7512A4" box="[383,475,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="96" pageNumber="97" unit="mm" value="1.5">1.5 mm</quantity>
. Right forewing. Petiolate. no accessory antenodal crossveins. Maximum width
<quantity id="4CE59B310072FFF7FAB5FF73FF6812C0" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="96" pageNumber="97" unit="mm" value="6.5">6.5 mm</quantity>
. Proximal portion hyaline, darkly infuscate from level of nodus to near level where
<specimenCode id="DBBB9EAF0072FFF7FB4FFF5EFB1D12C0" box="[1155,1203,187,213]" collectionCode="RP" pageId="96" pageNumber="97">RP1</specimenCode>
IR1 space becomes two cells wide, hyaline in portion preserved distal to this. Crossveins in postnodal, postsubnodal spaces almost all aligned.
<specimenCode id="DBBB9EAF0072FFF7FF35FEE6FE871308" box="[249,297,259,285]" collectionCode="RP" pageId="96" pageNumber="97">RP1</specimenCode>
space goes to two cells wide five cells distal to origin of IR1. IR1: origin three cells distal to origin of
<specimenCode id="DBBB9EAF0072FFF7FF7AFEC2FF441354" box="[182,234,295,321]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF0072FFF7FF39FEC2FE871354" box="[245,297,295,321]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
: origin four cells distal to origin of IR2. IR2: origin at subnodus; linear, straight in portion preserved.
<specimenCode id="DBBB9EAF0072FFF7FF5BFEAEFF4D1370" box="[151,227,331,357]" collectionCode="RP" pageId="96" pageNumber="97">RP3-4</specimenCode>
: origin about 2/3 from arculus to nodus; linear, straight in portion preserved.
<collectionCode id="ED0CAE110072FFF7FBA0FEAEFB361370" box="[1132,1176,331,357]" pageId="96" pageNumber="97">MA</collectionCode>
linear, slightly curved from quadrangle to about level of origin of IR1, increasingly zigzags distal to this.
<collectionCode id="ED0CAE110072FFF7FBF7FE8AFBCC139C" box="[1083,1122,367,393]" country="0" name="Mohonk Preserve, Inc." pageId="96" pageNumber="97">MP</collectionCode>
linear, slightly curved from quadrangle, small portion near terminus not preserved.
<collectionCode id="ED0CAE110072FFF7FCCAFE76FC9713B8" box="[774,825,403,429]" pageId="96" pageNumber="97">CuA</collectionCode>
increasingly zigzagged from level of origin of
<specimenCode id="DBBB9EAF0072FFF7FA81FE76FA3613B8" box="[1357,1432,403,429]" collectionCode="RP" pageId="96" pageNumber="97">RP3-4</specimenCode>
; CuAA space about seven columns of three cells at widest, otherwise two beyond level of origin of
<specimenCode id="DBBB9EAF0072FFF7FADAFE52FACF13C4" box="[1302,1377,439,465]" collectionCode="RP" pageId="96" pageNumber="97">RP3-4</specimenCode>
. Left forewing. Petiolate. no accessory antenodal crossveins in areas preserved. Width
<quantity id="4CE59B310072FFF7FBD1FE3EFBDB13E0" box="[1053,1141,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="96" pageNumber="97" unit="mm" value="6.6">6.6 mm</quantity>
. Proximal portion hyaline, darkly infuscate from level of nodus to near level where
<specimenCode id="DBBB9EAF0072FFF7FCC0FE1AFC92100C" box="[780,828,511,537]" collectionCode="RP" pageId="96" pageNumber="97">RP1</specimenCode>
IR1 space becomes two cells wide, hyaline in portion preserved distal to this. Crossveins in postnodal, postsubnodal spaces almost all aligned.
<specimenCode id="DBBB9EAF0072FFF7FB48FDC6FB1A1028" box="[1156,1204,547,573]" collectionCode="RP" pageId="96" pageNumber="97">RP1</specimenCode>
IR1 space becomes two cells wide four cells distal to origin of IR1. IR1: origin three cells distal to origin of
<specimenCode id="DBBB9EAF0072FFF7FBBBFDA2FB051074" box="[1143,1195,583,609]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF0072FFF7FB7AFDA2FB441074" box="[1206,1258,583,609]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
: origin 3.5 cells distal to origin of IR2. IR2: origin at subnodus; linear, straight in portion preserved.
<specimenCode id="DBBB9EAF0072FFF7FB86FD8EFB381090" box="[1098,1174,619,645]" collectionCode="RP" pageId="96" pageNumber="97">RP3-4</specimenCode>
: origin similar distance from nodus as right forewing; linear, straight in portion preserved.
<collectionCode id="ED0CAE110072FFF7FC5DFD6AFC1310BC" box="[913,957,655,681]" pageId="96" pageNumber="97">MA</collectionCode>
linear, slightly curved from quadrangle to about level of origin of IR1, zigzagged distal to this.
<collectionCode id="ED0CAE110072FFF7FD2FFD56FCA410D8" box="[739,778,691,717]" country="0" name="Mohonk Preserve, Inc." pageId="96" pageNumber="97">MP</collectionCode>
linear, slightly curved from quadrangle, small portion near terminus not preserved.
<collectionCode id="ED0CAE110072FFF7FE65FD32FE7210E4" box="[425,476,727,753]" pageId="96" pageNumber="97">CuA</collectionCode>
increasingly zigzagged from level of origin of
<specimenCode id="DBBB9EAF0072FFF7FC3BFD32FBEC10E4" box="[1015,1090,727,753]" collectionCode="RP" pageId="96" pageNumber="97">RP3-4</specimenCode>
; CuAA space number of columns of three cells wide difficult to determine as preserved, two evident, likely more, otherwise two beyond level of origin of
<specimenCode id="DBBB9EAF0072FFF7FEEDFCFAFEC2112C" box="[289,364,799,825]" collectionCode="RP" pageId="96" pageNumber="97">RP3-4</specimenCode>
. Right hind wing. Proximal part of wing from articulation to distal side of quadrangle present. Left hind wing. Only base of petiolate portion present. Abdomen mostly missing; only segments, 1, 2 and proximal part of 3 present
</materialsCitation>
.
</paragraph>
<caption id="DF62665C004DFFC8FF5BF849FB2315F2" ID-DOI="http://doi.org/10.5281/zenodo.4558940" ID-Zenodo-Dep="4558940" httpUri="https://zenodo.org/record/4558940/files/figure.png" pageId="95" pageNumber="96" startId="95.[151,250,1964,1988]" targetBox="[264,1322,182,1920]" targetPageId="95">
<paragraph id="8BA236D4004DFFC8FF5BF849FB2315F2" blockId="95.[151,1436,1964,2024]" pageId="95" pageNumber="96">
<emphasis id="B969EAC6004DFFC8FF5BF849FE8E15D1" bold="true" box="[151,288,1964,1988]" pageId="95" pageNumber="96">FIGURE 57.</emphasis>
<taxonomicName id="4C1D4D57004DFFC8FEEAF849FD9F15D1" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[294,561,1964,1988]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="macabeensis">
<emphasis id="B969EAC6004DFFC8FEEAF849FD9F15D1" box="[294,561,1964,1988]" italics="true" pageId="95" pageNumber="96">Okanopteryx macabeensis</emphasis>
</taxonomicName>
paratype 3, GSC 141102. A, photograph of specimen; B, drawing of upper wing in A, (vertically inverted for anterior margin on top); C, drawing of bottom wing in A. All to scale, 5 mm.
</paragraph>
</caption>
<paragraph id="8BA236D40072FFF7FF0BFC6EFB2116D0" blockId="96.[151,1437,150,1509]" pageId="96" pageNumber="97">
<materialsCitation id="3B753C890072FFF7FF0BFC6EFB2516D0" ID-GBIF-Occurrence="3069296315" collectionCode="MP, MA, CuA" pageId="96" pageNumber="97" specimenCode="RP1, RP2, RP3-4" specimenCount="2" typeStatus="paratype">
<typeStatus id="54A688760072FFF7FF0BFC6EFE8711B0" box="[199,297,907,933]" pageId="96" pageNumber="97" type="paratype">Paratype</typeStatus>
3, GSC 141102, two wings (
<figureCitation id="13262A510072FFF7FDBAFC6EFD6211B0" box="[630,716,906,933]" captionStart="FIGURE 57" captionStartId="95.[151,250,1964,1988]" captionTargetBox="[264,1322,182,1920]" captionTargetId="figure-17@95.[251,1337,181,1940]" captionTargetPageId="95" captionText="FIGURE 57. Okanopteryx macabeensis paratype 3, GSC 141102. A, photograph of specimen; B, drawing of upper wing in A, (vertically inverted for anterior margin on top); C, drawing of bottom wing in A. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558940" httpUri="https://zenodo.org/record/4558940/files/figure.png" pageId="96" pageNumber="97">Fig. 57</figureCitation>
). Middle portions. Wing 1 (
<figureCitation id="13262A510072FFF7FBC5FC6EFBC611B0" box="[1033,1128,906,933]" captionStart="FIGURE 57" captionStartId="95.[151,250,1964,1988]" captionTargetBox="[264,1322,182,1920]" captionTargetId="figure-17@95.[251,1337,181,1940]" captionTargetPageId="95" captionText="FIGURE 57. Okanopteryx macabeensis paratype 3, GSC 141102. A, photograph of specimen; B, drawing of upper wing in A, (vertically inverted for anterior margin on top); C, drawing of bottom wing in A. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558940" httpUri="https://zenodo.org/record/4558940/files/figure.png" pageId="96" pageNumber="97">Fig 57B</figureCitation>
) missing posterior portion. Darkly infuscate consistent with the
<typeStatus id="54A688760072FFF7FDF5FC4AFD3111DC" box="[569,671,943,969]" pageId="96" pageNumber="97" type="holotype">holotype</typeStatus>
. Crossveins in preserved portion of postnodal, postsubnodal spaces aligned.
<specimenCode id="DBBB9EAF0072FFF7FF3AFC36FE8811F8" box="[246,294,979,1005]" collectionCode="RP" pageId="96" pageNumber="97">RP1</specimenCode>
IR1 space becomes two cells wide four cells distal to origin of IR1. IR1: origin three cells distal to that of
<specimenCode id="DBBB9EAF0072FFF7FF7AFC12FF441604" box="[182,234,1015,1041]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF0072FFF7FF39FC12FE841604" box="[245,298,1015,1041]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
: origin not preserved. Preserved portions of IR1,
<specimenCode id="DBBB9EAF0072FFF7FC95FC12FC221604" box="[857,908,1015,1041]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
, IR2,
<specimenCode id="DBBB9EAF0072FFF7FC1DFC12FBB51604" box="[977,1051,1015,1041]" collectionCode="RP" pageId="96" pageNumber="97">RP3-4</specimenCode>
,
<collectionCode id="ED0CAE110072FFF7FBEBFC12FBE01604" box="[1063,1102,1015,1041]" country="0" name="Mohonk Preserve, Inc." pageId="96" pageNumber="97">MP</collectionCode>
linear, mostly rather straight.
<collectionCode id="ED0CAE110072FFF7FF5BFBFEFF6D1620" box="[151,195,1051,1077]" pageId="96" pageNumber="97">MA</collectionCode>
becoming, increasingly zigzagged distal to level of origin of IR1.
<collectionCode id="ED0CAE110072FFF7FC53FBFEFC681620" box="[927,966,1051,1077]" country="0" name="Mohonk Preserve, Inc." pageId="96" pageNumber="97">MP</collectionCode>
linear, slightly curved. Wing 2 (
<figureCitation id="13262A510072FFF7FAE6FBFEFA3E1620" box="[1322,1424,1050,1077]" captionStart="FIGURE 50" captionStartId="87.[151,250,2001,2025]" captionTargetBox="[243,1343,186,1972]" captionTargetId="figure-17@87.[231,1357,181,1977]" captionTargetPageId="87" captionText="FIGURE 50. Okanopteryx fraseri paratype 1, RBCM P1551. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558923" httpUri="https://zenodo.org/record/4558923/files/figure.png" pageId="96" pageNumber="97">Fig. 50C</figureCitation>
). As for wing 1, except: maximum width
<quantity id="4CE59B310072FFF7FD99FBDAFD03164C" box="[597,685,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="96" pageNumber="97" unit="mm" value="6.8">6.8 mm</quantity>
. Crossveins in preserved portion of postnodal, postsubnodal spaces aligned, at most slightly not.
<specimenCode id="DBBB9EAF0072FFF7FE12FB86FDA31668" box="[478,525,1123,1149]" collectionCode="RP" pageId="96" pageNumber="97">RP1</specimenCode>
IR1 space becomes two cells wide three cells distal to origin of IR1. IR1: origin two cells distal to that of
<specimenCode id="DBBB9EAF0072FFF7FE63FB62FE4D16B4" box="[431,483,1159,1185]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
.
<specimenCode id="DBBB9EAF0072FFF7FE21FB62FD8F16B4" box="[493,545,1159,1185]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
: origin 2.5 cells distal to that of IR2.
<collectionCode id="ED0CAE110072FFF7FC71FB62FC5E16B4" box="[957,1008,1159,1185]" pageId="96" pageNumber="97">CuA</collectionCode>
increasingly zigzagged about distal to level of origin of
<specimenCode id="DBBB9EAF0072FFF7FE97FB4EFE2116D0" box="[347,399,1195,1221]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
;&gt;7 columns of cells three cells deep, the rest two, then one apically
</materialsCitation>
.
</paragraph>
<paragraph id="8BA236D40072FFF7FF0BFB2AFAEC1788" blockId="96.[151,1437,150,1509]" pageId="96" pageNumber="97">
<materialsCitation id="3B753C890072FFF7FF0BFB2AFA901788" ID-GBIF-Occurrence="3069296320" collectionCode="MP, MA, CuA" pageId="96" pageNumber="97" specimenCode="RP1, RP2, RP3-4" specimenCount="2" typeStatus="paratype">
<typeStatus id="54A688760072FFF7FF0BFB2AFE8716FC" box="[199,297,1231,1257]" pageId="96" pageNumber="97" type="paratype">Paratype</typeStatus>
4, RBCM 3087, wing (
<figureCitation id="13262A510072FFF7FDF4FB2AFD2216FC" box="[568,652,1230,1257]" captionStart="FIGURE 53" captionStartId="91.[151,250,1868,1892]" captionTargetBox="[161,1426,181,1835]" captionTargetId="figure-17@91.[151,1436,181,1844]" captionTargetPageId="91" captionText="FIGURE 53. Okanopteryx macabeensis holotype, GSC 141100: A, photograph; B, drawing. Paratype 4: RBCM 3087: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558930" httpUri="https://zenodo.org/record/4558930/files/figure.png" pageId="96" pageNumber="97">Fig. 53</figureCitation>
). Parts of apical half, missing much of apical-most posterior portion. Maximum width
<quantity id="4CE59B310072FFF7FE95FB16FE1F1718" box="[345,433,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="96" pageNumber="97" unit="mm" value="6.9">6.9 mm</quantity>
. Proximally-preserved portion infuscate, hyaline apical to this as in
<typeStatus id="54A688760072FFF7FB6BFB16FAA31718" box="[1191,1293,1267,1293]" pageId="96" pageNumber="97" type="holotype">holotype</typeStatus>
. Pterostigma broad, 2.5 times longer than wide. Preserved crossveins in postnodal, postsubnodal spaces aligned or almost so. Two (possibly more?) cells wide distal to pterostigma in costal space (fragmentarily preserved).
<specimenCode id="DBBB9EAF0072FFF7FB44FADEFB161740" box="[1160,1208,1339,1365]" collectionCode="RP" pageId="96" pageNumber="97">RP1</specimenCode>
IR1 space becomes two cells wide four cells distal to origin of IR1. Preserved portions of
<specimenCode id="DBBB9EAF0072FFF7FC63FABAFC4C176C" box="[943,994,1375,1401]" collectionCode="RP" pageId="96" pageNumber="97">RP2</specimenCode>
, IR2,
<specimenCode id="DBBB9EAF0072FFF7FBEAFABAFBDE176C" box="[1062,1136,1375,1401]" collectionCode="RP" pageId="96" pageNumber="97">RP3-4</specimenCode>
,
<collectionCode id="ED0CAE110072FFF7FBB0FABAFB0D176C" box="[1148,1187,1375,1401]" country="0" name="Mohonk Preserve, Inc." pageId="96" pageNumber="97">MP</collectionCode>
linear, rather straight.
<collectionCode id="ED0CAE110072FFF7FF5BFA66FF6D1788" box="[151,195,1411,1437]" pageId="96" pageNumber="97">MA</collectionCode>
indistinctly preserved.
<collectionCode id="ED0CAE110072FFF7FE06FA66FE531788" box="[458,509,1411,1437]" pageId="96" pageNumber="97">CuA</collectionCode>
zigzagged in portion preserved; at least a few columns of three cells wide
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C307655F0072FFF7FF0BFA42FC8717F0" pageId="96" pageNumber="97" type="etymology">
<paragraph id="8BA236D40072FFF7FF0BFA42FBF017D4" blockId="96.[151,1437,150,1509]" box="[199,1118,1447,1473]" pageId="96" pageNumber="97">
<emphasis id="B969EAC60072FFF7FF0BFA42FEE617D4" bold="true" box="[199,328,1447,1473]" pageId="96" pageNumber="97">Etymology</emphasis>
. The specific epithet is a toponym referring to its only known locality.
</paragraph>
<paragraph id="8BA236D40072FFF7FF0BFA2EFC8717F0" blockId="96.[151,1437,150,1509]" box="[199,809,1483,1509]" pageId="96" pageNumber="97">
<emphasis id="B969EAC60072FFF7FF0BFA2EFED917F0" bold="true" box="[199,375,1483,1509]" pageId="96" pageNumber="97">Range and age</emphasis>
. McAbee, BC,
<collectingCountry id="F30A76440072FFF7FDEBFA2EFDD117F0" box="[551,639,1483,1509]" name="Canada" pageId="96" pageNumber="97">Canada</collectingCountry>
; mid-Ypresian.
</paragraph>
</subSubSection>
</treatment>
</document>