474 lines
59 KiB
XML
474 lines
59 KiB
XML
<document id="3B18856B46C22863CA860CED423BE626" ID-CLB-Dataset="7439" ID-DOI="10.11646/zootaxa.4934.1.1" ID-GBIF-Dataset="300bac18-55f5-4e34-90e1-7fccd02d68c1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4558796" ID-ZooBank="79895443-4597-42A5-AF8A-023EACB20E10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1614157470936" checkinUser="plazi" docAuthor="Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. & Mathewes, Rolf W." docDate="2021" docId="03B487C20046FFCDFF5BF8EBFDCE112C" docLanguage="en" docName="zootaxa.4934.1.1.pdf" docOrigin="Zootaxa 4934 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Okanopteryx fraseri Archibald & Cannings & Erickson & Bybee & Mathewes 2021, new species" docType="treatment" docVersion="9" lastPageNumber="91" masterDocId="FF8DFFBA0012FF97FFCCFFE5FFAE1215" masterDocTitle="The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America" masterLastPageNumber="133" masterPageNumber="1" pageNumber="85" updateTime="1698931040708" updateUser="ExternalLinkService">
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<mods:title id="3DE8EE202CFE2F49C6B221D40962D07F">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
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<mods:namePart id="20AAC3879C7FE9E0F4B0CDFDF565A48E">Archibald, Bruce</mods:namePart>
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<mods:namePart id="E047E86992ACAB1140C96AC6C8505E49">Cannings, Robert A.</mods:namePart>
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<mods:nameIdentifier id="7FA626CB7A1179CA3DB2679F4592CDCA" type="email">annings@royalbcmuseum.bc.ca</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="B36BD46DBF5ABD17EA951FE661BEE6F1">Erickson, Robert J.</mods:namePart>
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<mods:nameIdentifier id="6F22703838A2353ABF0F6EF48B3AC551" type="email">1990platypus@gmail.com</mods:nameIdentifier>
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<mods:namePart id="DCAEC3B4A79D28B75201CAE68B3E80A6">Bybee, Seth M.</mods:namePart>
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<mods:nameIdentifier id="EDED49BD09FD27321D7E059AD434D949" type="email">seth.bybee@gmail.com</mods:nameIdentifier>
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<mods:namePart id="6713297F64905CA640C993377222C8C0">Mathewes, Rolf W.</mods:namePart>
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<mods:nameIdentifier id="B628C458678807C14AC5818FE4BC0869" type="email">r_mathewes@sfu.ca</mods:nameIdentifier>
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<mods:title id="41EA64E2C64A7D898B762F745459F0DE">Zootaxa</mods:title>
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<mods:part id="FD9DB1041F602DCCC25B2EFC78A49A76">
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<mods:date id="2429170C837C4C1F349FD4924CCA00AE">2021</mods:date>
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<mods:number id="32C8C63768DE7CE971F15A1001093BAC">2021-02-24</mods:number>
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<mods:number id="4662E40F8E970E2212819E917ED1BF2D">4934</mods:number>
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<mods:identifier id="CE18B0ECAEDC22CBB6447B77DD0D6050" type="DOI">10.11646/zootaxa.4934.1.1</mods:identifier>
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<treatment id="03B487C20046FFCDFF5BF8EBFDCE112C" ID-DOI="http://doi.org/10.5281/zenodo.4559805" ID-GBIF-Taxon="180881800" ID-Zenodo-Dep="4559805" LSID="urn:lsid:plazi:treatment:03B487C20046FFCDFF5BF8EBFDCE112C" httpUri="http://treatment.plazi.org/id/03B487C20046FFCDFF5BF8EBFDCE112C" lastPageId="90" lastPageNumber="91" pageId="84" pageNumber="85">
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<subSubSection id="C307655F0046FFC3FF5BF8EBFCE2153D" box="[151,844,1806,1833]" pageId="84" pageNumber="85" type="nomenclature">
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<paragraph id="8BA236D40046FFC3FF5BF8EBFCE2153D" blockId="84.[151,844,1806,1869]" box="[151,844,1806,1833]" pageId="84" pageNumber="85">
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<heading id="D0EA81B80046FFC3FF5BF8EBFCE2153D" bold="true" box="[151,844,1806,1833]" fontSize="11" level="2" pageId="84" pageNumber="85" reason="4">
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<emphasis id="B969EAC60046FFC3FF5BF8EBFCE2153D" bold="true" box="[151,844,1806,1833]" pageId="84" pageNumber="85">
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<taxonomicName id="4C1D4D570046FFC3FF5BF8EBFD01153D" authority="Archibald & Cannings" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[151,687,1806,1833]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="fraseri" status="sp. nov.">
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<emphasis id="B969EAC60046FFC3FF5BF8EBFE25153D" bold="true" box="[151,395,1806,1833]" italics="true" pageId="84" pageNumber="85">Okanopteryx fraseri</emphasis>
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Archibald & Cannings
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25A57BD0046FFC3FD70F8EAFCE2153D" box="[700,844,1806,1833]" pageId="84" pageNumber="85" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C307655F0046FFC3FF5BF8D6FE8D1558" box="[151,291,1842,1869]" pageId="84" pageNumber="85" type="description">
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<paragraph id="8BA236D40046FFC3FF5BF8D6FE8D1558" blockId="84.[151,844,1806,1869]" box="[151,291,1842,1869]" pageId="84" pageNumber="85">
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<figureCitation id="13262A510046FFC3FF5BF8D6FEB11558" box="[151,287,1842,1869]" captionStart-0="FIGURE 49" captionStart-1="FIGURE 50" captionStart-2="FIGURE 51" captionStart-3="FIGURE 52" captionStartId-0="86.[151,250,1943,1967]" captionStartId-1="87.[151,250,2001,2025]" captionStartId-2="88.[151,250,1934,1958]" captionStartId-3="89.[151,250,1954,1978]" captionTargetBox-0="[166,1422,181,1919]" captionTargetBox-1="[243,1343,186,1972]" captionTargetBox-2="[154,1427,181,1898]" captionTargetBox-3="[361,1219,184,1913]" captionTargetId-0="figure-19@86.[166,1422,181,1919]" captionTargetId-1="figure-17@87.[231,1357,181,1977]" captionTargetId-2="figure-19@88.[151,1436,181,1910]" captionTargetId-3="figure-17@89.[353,1235,181,1930]" captionTargetPageId-0="86" captionTargetPageId-1="87" captionTargetPageId-2="88" captionTargetPageId-3="89" captionText-0="FIGURE 49. Okanopteryx fraseri holotype, GSC 141104/F-778. A, photograph of entire specimen (part: GSC 141104); B, drawing of wing. Both to scale, 5 mm." captionText-1="FIGURE 50. Okanopteryx fraseri paratype 1, RBCM P1551. A, photograph; B, drawing. Both to scale, 5 mm." captionText-2="FIGURE 51. Okanopteryx fraseri paratype 3, PB-4157. A, photograph; B, drawing; paratype 2, F-1091: C, photograph; D, drawing.All to scale, 5 mm." captionText-3="FIGURE 52. Okanopteryx fraseri paratype 5, RBCM P1552. A, photograph; B, drawing; paratype 4, F-1045: C, photograph; D, drawing. All to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4558920" figureDoi-1="http://doi.org/10.5281/zenodo.4558923" figureDoi-2="http://doi.org/10.5281/zenodo.4558926" figureDoi-3="http://doi.org/10.5281/zenodo.4558928" httpUri-0="https://zenodo.org/record/4558920/files/figure.png" httpUri-1="https://zenodo.org/record/4558923/files/figure.png" httpUri-2="https://zenodo.org/record/4558926/files/figure.png" httpUri-3="https://zenodo.org/record/4558928/files/figure.png" pageId="84" pageNumber="85">Figs. 49–52</figureCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C307655F0046FFC2FF5BF89FFF4E1308" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="diagnosis">
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<paragraph id="8BA236D40046FFC2FF5BF89FFF4E1308" blockId="84.[151,1437,1914,2013]" lastBlockId="85.[151,1437,151,2013]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
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<emphasis id="B969EAC60046FFC3FF5BF89FFEBC1581" bold="true" box="[151,274,1914,1940]" pageId="84" pageNumber="85">Diagnosis.</emphasis>
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Separated from other
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<taxonomicName id="4C1D4D570046FFC3FDC5F89EFD341580" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[521,666,1915,1941]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="84" pageNumber="85" phylum="Arthropoda" rank="genus">
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<emphasis id="B969EAC60046FFC3FDC5F89EFD341580" box="[521,666,1915,1941]" italics="true" pageId="84" pageNumber="85">Okanopteryx</emphasis>
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</taxonomicName>
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species by: abdomen longer,
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<quantity id="4CE59B310046FFC3FC2DF89FFBEB1580" box="[993,1093,1914,1941]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" pageId="84" pageNumber="85" unit="mm" value="29.5">29.5 mm</quantity>
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from proximal end of abdominal segment 3 to distal end of segment 7 [
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<taxonomicName id="4C1D4D570046FFC3FDAEF87AFCBD15AC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[610,787,1951,1977]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="macabeensis">
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<emphasis id="B969EAC60046FFC3FDAEF87AFCBD15AC" box="[610,787,1951,1977]" italics="true" pageId="84" pageNumber="85">O. macabeensis</emphasis>
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</taxonomicName>
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: 20.0 mm]. We have compared the abdomen lengths using segments 3 to 7 only, as the
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<taxonomicName id="4C1D4D570046FFC3FE14F826FDE915C8" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[472,583,1987,2013]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="fraseri">
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<emphasis id="B969EAC60046FFC3FE14F826FDE915C8" box="[472,583,1987,2013]" italics="true" pageId="84" pageNumber="85">O. fraseri</emphasis>
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</taxonomicName>
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abdomen is missing from the middle of segment 8 distally and the margins of the two proximal segments are difficult to interpret (
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<taxonomicName id="4C1D4D570047FFC2FCC6FF72FC6412A5" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[778,970,151,177]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="jeppesenorum">
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<emphasis id="B969EAC60047FFC2FCC6FF72FC6412A5" box="[778,970,151,177]" italics="true" pageId="85" pageNumber="86">O. jeppesenorum</emphasis>
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</taxonomicName>
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only known from wings). Wings distinct from
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<taxonomicName id="4C1D4D570047FFC2FF19FF5EFE3A12C1" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[213,404,187,213]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="jeppesenorum">
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<emphasis id="B969EAC60047FFC2FF19FF5EFE3A12C1" box="[213,404,187,213]" italics="true" pageId="85" pageNumber="86">O. jeppesenorum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C1D4D570047FFC2FE6FFF5EFDFA12C0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[419,596,187,213]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="macabeensis">
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<emphasis id="B969EAC60047FFC2FE6FFF5EFDFA12C0" box="[419,596,187,213]" italics="true" pageId="85" pageNumber="86">O. macabeensis</emphasis>
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</taxonomicName>
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by CuA–A space narrower, usually two cell rows at widest between levels of origins of IR1, RP2, sometimes a single column of three [
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<taxonomicName id="4C1D4D570047FFC2FCFFFF3AFC5C12ED" box="[819,1010,223,249]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="jeppesneorum">
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<emphasis id="B969EAC60047FFC2FCFFFF3AFC5C12ED" box="[819,1010,223,249]" italics="true" pageId="85" pageNumber="86">O. jeppesneorum</emphasis>
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</taxonomicName>
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: always four;
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<taxonomicName id="4C1D4D570047FFC2FB5CFF3AFAEE12EC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[1168,1344,223,249]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="macabeensis">
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<emphasis id="B969EAC60047FFC2FB5CFF3AFAEE12EC" box="[1168,1344,223,249]" italics="true" pageId="85" pageNumber="86">O. macabeensis</emphasis>
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</taxonomicName>
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: always three].
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C307655F0047FFC2FF0BFEC3FC761090" pageId="85" pageNumber="86" type="materials_examined">
|
||
<paragraph id="8BA236D40047FFC2FF0BFEC3FC761090" blockId="85.[151,1437,151,2013]" pageId="85" pageNumber="86">
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||
<emphasis id="B969EAC60047FFC2FF0BFEC3FE981355" bold="true" box="[199,310,294,320]" pageId="85" pageNumber="86">Material.</emphasis>
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||
<materialsCitation id="3B753C890047FFC2FE8CFEC2FC761090" ID-GBIF-Occurrence="3069296325" collectorName="John Fraser" country="Canada" county="McAbee Hoodoo Face" location="Royal" municipality="John Leahy" pageId="85" pageNumber="86" specimenCount="6" stateProvince="British Columbia" typeStatus="holotype">
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||
<emphasis id="B969EAC60047FFC2FE8CFEC2FE091354" box="[320,423,295,321]" italics="true" pageId="85" pageNumber="86">
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<typeStatus id="54A688760047FFC2FE8CFEC2FE091354" box="[320,423,295,321]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
: GSC 141104, part, housed in the Geological Survey of
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<collectingCountry id="F30A76440047FFC2FBF5FEC2FB231354" box="[1081,1165,295,321]" name="Canada" pageId="85" pageNumber="86">Canada</collectingCountry>
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collections, and F-778, counterpart, in the Thompson Rivers University collections (
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<figureCitation id="13262A510047FFC2FC8EFEAEFC381370" box="[834,918,331,357]" captionStart="FIGURE 49" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-19@86.[166,1422,181,1919]" captionTargetPageId="86" captionText="FIGURE 49. Okanopteryx fraseri holotype, GSC 141104/F-778. A, photograph of entire specimen (part: GSC 141104); B, drawing of wing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558920" httpUri="https://zenodo.org/record/4558920/files/figure.png" pageId="85" pageNumber="86">Fig. 49</figureCitation>
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), collected at the McAbee Hoodoo Face beds, Geological Survey of
|
||
<collectingCountry id="F30A76440047FFC2FE41FE8AFE4F139C" box="[397,481,367,393]" name="Canada" pageId="85" pageNumber="86">Canada</collectingCountry>
|
||
locality code V-016800, collected by
|
||
<collectorName id="26E853020047FFC2FC46FE8AFBA3139C" box="[906,1037,367,393]" pageId="85" pageNumber="86">John Fraser</collectorName>
|
||
, unknown date;
|
||
<emphasis id="B969EAC60047FFC2FB08FE8AFA93139C" box="[1220,1341,367,393]" italics="true" pageId="85" pageNumber="86">
|
||
<typeStatus id="54A688760047FFC2FB08FE8AFA86139D" box="[1220,1320,367,392]" pageId="85" pageNumber="86" type="paratype">paratype</typeStatus>
|
||
1
|
||
</emphasis>
|
||
: RBCM P1551 (
|
||
<figureCitation id="13262A510047FFC2FF3CFE76FEEB13B8" box="[240,325,402,429]" captionStart="FIGURE 50" captionStartId="87.[151,250,2001,2025]" captionTargetBox="[243,1343,186,1972]" captionTargetId="figure-17@87.[231,1357,181,1977]" captionTargetPageId="87" captionText="FIGURE 50. Okanopteryx fraseri paratype 1, RBCM P1551. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558923" httpUri="https://zenodo.org/record/4558923/files/figure.png" pageId="85" pageNumber="86">Fig. 50</figureCitation>
|
||
), collected at the McAbee Hoodoo Face beds by John Leahy, date unknown, in the Royal
|
||
<collectingRegion id="49D9F8360047FFC2FA9CFE76FEAB13C4" country="Canada" name="British Columbia" pageId="85" pageNumber="86">British Columbia</collectingRegion>
|
||
Museum collections;
|
||
<emphasis id="B969EAC60047FFC2FE31FE52FDDB13C4" box="[509,629,439,465]" italics="true" pageId="85" pageNumber="86">
|
||
<typeStatus id="54A688760047FFC2FE31FE52FDCF13C5" box="[509,609,439,464]" pageId="85" pageNumber="86" type="paratype">paratype</typeStatus>
|
||
2
|
||
</emphasis>
|
||
: F-1091 (
|
||
<figureCitation id="13262A510047FFC2FD2FFE52FC9B13C4" box="[739,821,438,465]" captionStart="FIGURE 51" captionStartId="88.[151,250,1934,1958]" captionTargetBox="[154,1427,181,1898]" captionTargetId="figure-19@88.[151,1436,181,1910]" captionTargetPageId="88" captionText="FIGURE 51. Okanopteryx fraseri paratype 3, PB-4157. A, photograph; B, drawing; paratype 2, F-1091: C, photograph; D, drawing.All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558926" httpUri="https://zenodo.org/record/4558926/files/figure.png" pageId="85" pageNumber="86">Fig. 51</figureCitation>
|
||
), collected at the McAbee Hoodoo Face beds, date and collector unknown, in the Thompson Rivers University collections;
|
||
<emphasis id="B969EAC60047FFC2FC5CFE3EFBA713E0" box="[912,1033,475,501]" italics="true" pageId="85" pageNumber="86">
|
||
<typeStatus id="54A688760047FFC2FC5CFE3EFC5A13E1" box="[912,1012,475,500]" pageId="85" pageNumber="86" type="paratype">paratype</typeStatus>
|
||
3
|
||
</emphasis>
|
||
: PB-4157 (
|
||
<figureCitation id="13262A510047FFC2FB47FE3EFB7313E0" box="[1163,1245,474,501]" captionStart="FIGURE 51" captionStartId="88.[151,250,1934,1958]" captionTargetBox="[154,1427,181,1898]" captionTargetId="figure-19@88.[151,1436,181,1910]" captionTargetPageId="88" captionText="FIGURE 51. Okanopteryx fraseri paratype 3, PB-4157. A, photograph; B, drawing; paratype 2, F-1091: C, photograph; D, drawing.All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558926" httpUri="https://zenodo.org/record/4558926/files/figure.png" pageId="85" pageNumber="86">Fig. 51</figureCitation>
|
||
), collected at the McAbee Hoodoo Face beds, date and collector unknown, in the Thompson Rivers University collections;
|
||
<emphasis id="B969EAC60047FFC2FAF5FE1AFF0B1028" italics="true" pageId="85" pageNumber="86">
|
||
<typeStatus id="54A688760047FFC2FAF5FE1AFA33100D" box="[1337,1437,511,536]" pageId="85" pageNumber="86" type="paratype">paratype</typeStatus>
|
||
4
|
||
</emphasis>
|
||
: F-1045 (
|
||
<figureCitation id="13262A510047FFC2FEDEFDC6FECB1028" box="[274,357,546,573]" captionStart="FIGURE 52" captionStartId="89.[151,250,1954,1978]" captionTargetBox="[361,1219,184,1913]" captionTargetId="figure-17@89.[353,1235,181,1930]" captionTargetPageId="89" captionText="FIGURE 52. Okanopteryx fraseri paratype 5, RBCM P1552. A, photograph; B, drawing; paratype 4, F-1045: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558928" httpUri="https://zenodo.org/record/4558928/files/figure.png" pageId="85" pageNumber="86">Fig. 52</figureCitation>
|
||
), collected at the McAbee Hoodoo Face beds, date and collector unknown, in the Thompson Rivers University collections;
|
||
<emphasis id="B969EAC60047FFC2FE0BFDA2FDEE1074" box="[455,576,583,609]" italics="true" pageId="85" pageNumber="86">
|
||
<typeStatus id="54A688760047FFC2FE0BFDA2FD851075" box="[455,555,583,608]" pageId="85" pageNumber="86" type="paratype">paratype</typeStatus>
|
||
5
|
||
</emphasis>
|
||
: RBCM P1552 (
|
||
<figureCitation id="13262A510047FFC2FCCDFDA2FCFB1074" box="[769,853,582,609]" captionStart="FIGURE 52" captionStartId="89.[151,250,1954,1978]" captionTargetBox="[361,1219,184,1913]" captionTargetId="figure-17@89.[353,1235,181,1930]" captionTargetPageId="89" captionText="FIGURE 52. Okanopteryx fraseri paratype 5, RBCM P1552. A, photograph; B, drawing; paratype 4, F-1045: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558928" httpUri="https://zenodo.org/record/4558928/files/figure.png" pageId="85" pageNumber="86">Fig. 52</figureCitation>
|
||
), collected at the
|
||
<collectingCounty id="62C34E580047FFC2FBD2FDA2FAB31074" box="[1054,1309,583,609]" pageId="85" pageNumber="86">McAbee Hoodoo Face</collectingCounty>
|
||
beds,
|
||
<collectingMunicipality id="6BC6ACAE0047FFC2FAABFDA2FF701090" pageId="85" pageNumber="86">John Leahy</collectingMunicipality>
|
||
, date unknown, in the
|
||
<location id="8EC2600F0047FFC2FE17FD8EFDB01090" LSID="urn:lsid:plazi:treatment:03B487C20046FFCDFF5BF8EBFDCE112C:8EC2600F0047FFC2FE17FD8EFDB01090" box="[475,542,619,645]" country="Canada" county="McAbee Hoodoo Face" municipality="John Leahy" name="Royal" pageId="85" pageNumber="86" stateProvince="British Columbia">Royal</location>
|
||
<collectingRegion id="49D9F8360047FFC2FDE9FD8EFD481090" box="[549,742,619,645]" country="Canada" name="British Columbia" pageId="85" pageNumber="86">British Columbia</collectingRegion>
|
||
<location id="8EC2600F0047FFC2FD22FD8EFCFE1090" LSID="urn:lsid:plazi:treatment:03B487C20046FFCDFF5BF8EBFDCE112C:8EC2600F0047FFC2FD22FD8EFCFE1090" box="[750,848,619,645]" country="Canada" county="McAbee Hoodoo Face" municipality="John Leahy" name="Museum" pageId="85" pageNumber="86" stateProvince="British Columbia">Museum</location>
|
||
collections.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F0047FFCDFF0BFD6BFE091354" lastPageId="90" lastPageNumber="91" pageId="85" pageNumber="86" type="description">
|
||
<paragraph id="8BA236D40047FFC2FF0BFD6BFB4E1620" blockId="85.[151,1437,151,2013]" pageId="85" pageNumber="86">
|
||
<emphasis id="B969EAC60047FFC2FF0BFD6BFEF710BD" bold="true" box="[199,345,654,680]" pageId="85" pageNumber="86">Description.</emphasis>
|
||
<typeStatus id="54A688760047FFC2FEADFD6AFE6210BC" box="[353,460,655,681]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
|
||
, GSC 141104, F-778 (
|
||
<figureCitation id="13262A510047FFC2FD01FD6AFC8C10BC" box="[717,802,655,681]" captionStart="FIGURE 49" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[166,1422,181,1919]" captionTargetId="figure-19@86.[166,1422,181,1919]" captionTargetPageId="86" captionText="FIGURE 49. Okanopteryx fraseri holotype, GSC 141104/F-778. A, photograph of entire specimen (part: GSC 141104); B, drawing of wing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558920" httpUri="https://zenodo.org/record/4558920/files/figure.png" pageId="85" pageNumber="86">Fig. 49</figureCitation>
|
||
). A mostly complete specimen in dorsal aspect but with the bases of some of the legs possibly displaced to a lateral or dorsal position; one leg, at least, with part lying above the thorax. Abdomen distal to middle of segment 8 missing; two wings lacking the apical portions and only partially, mostly poorly, preserved, the other two wings fragmented; all legs mostly complete but poorly preserved. As in diagnosis and: Head (
|
||
<figureCitation id="13262A510047FFC2FEBCFCFAFE72112C" box="[368,476,799,825]" captionStart="FIGURE 10" captionStartId="22.[151,250,1598,1622]" captionTargetBox="[327,1259,184,1571]" captionTargetId="figure-19@22.[321,1266,181,1574]" captionTargetPageId="22" captionText="FIGURE 10. Heads of Zygoptera (A, B) and comparative Anisoptera, Anisozygoptera, Tarsophlebiidae, Stenophlebiidae (C– H); and Cephalozygoptera (I–Q).A, Zygoptera, Heteropodagrion superbum Ris redrawn from Fig. 318 of Garrison et al. (2010); B, Zygoptera, Minagrion ribeiroi (Santos) redrawn from Fig. 1842 of Garrison et al. (2010); C, Anisoptera, Gomphidae, Cyanogomphus waltheri Selys redrawn from Fig. 383 of Garrison et al. (2006); D, Anisozygoptera, Epiophlebia laidlawi Tillyard redrawn from Fig. 2 of Büsse (2016); E, Anisoptera, Aeshnidae, Limnetron antarcticum Förster redrawn from Fig. 16 of Garrison et al. (2006); F, Stenophlebia sp. redrawn from Tillyard & Fraser 1940, Fig, 4C; G, Tarsophlebiidae, Tarsophlebia eximia Hagen redrawn from Fleck et al. (2004) Fig. 1C; H, photograph of Tarsophlebia eximia in MCZ collections; I, Sieblosia jucunda redrawn from Hagen (1858, Plate 24, Fig. 1); J, Cephalozygoptera incertae sedis SR 14-003-004; K, Phenacolestes parallelus UCM-4545; L, Okanopteryx macabeensis, GSC 141101; M, Okanagrion beardi paratype 1, RBCM P1546; N, Okanopteryx fraseri GSC 141104; O, Okanopteryx macabeensis F-790; P, Dysagrion fredericii redrawn from Scudder 1890, plate 6, Fig. 10; Q, Dysagrion fredericii redrawn from Scudder 1890, plate 6, Fig. 2. G and H, I, J to O to scales, all 5 mm; others, originals without scales." figureDoi="http://doi.org/10.5281/zenodo.4558820" httpUri="https://zenodo.org/record/4558820/files/figure.png" pageId="85" pageNumber="86">Fig. 10 N</figureCitation>
|
||
,
|
||
<tableCitation id="C69F036F0047FFC2FE29FCFAFD98112C" box="[485,566,799,825]" captionStart="TABLE 1" captionStartId="23.[151,239,485,511]" captionTargetPageId="23" captionText="TABLE 1. Head measurements of Cephalozygoptera and measurement ratios: WH, width of head; WBE, width between eyes; WE, width of one eye; LE, length of one eye; LH, length of head. See Fig. 10 for head images and text for measurement details." httpUri="http://table.plazi.org/id/DF62665C0005FF80FF5BFE00FC7F1036" pageId="85" pageNumber="86" tableUuid="DF62665C0005FF80FF5BFE00FC7F1036">Table 1</tableCitation>
|
||
): in dorsal view, about
|
||
<quantity id="4CE59B310047FFC2FCF4FCFAFC23112C" box="[824,909,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="85" pageNumber="86" unit="mm" value="3.6">3.6 mm</quantity>
|
||
across eyes at widest point; approximate length from anterior edge of antefrons to posterior of occiput
|
||
<quantity id="4CE59B310047FFC2FD3EFCA6FCE41148" box="[754,842,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="85" pageNumber="86" unit="mm" value="1.8">1.8 mm</quantity>
|
||
; at level of hind ocelli distance between eyes
|
||
<quantity id="4CE59B310047FFC2FA8DFCA6FA361148" box="[1345,1432,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="85" pageNumber="86" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, width of eye about
|
||
<quantity id="4CE59B310047FFC2FEA2FC82FE681194" box="[366,454,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="85" pageNumber="86" unit="mm" value="1.1">1.1 mm</quantity>
|
||
, length
|
||
<quantity id="4CE59B310047FFC2FDD1FC82FDD81194" box="[541,630,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="85" pageNumber="86" unit="mm" value="2.3">2.3 mm</quantity>
|
||
; eyes rather compressed laterally, with acute but rounded postero-lateral corners; protuberance of clypeus and labrum not evident. Thorax: pterothorax about
|
||
<quantity id="4CE59B310047FFC2FB82FC6FFB0A11B0" box="[1102,1188,906,933]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="85" pageNumber="86" unit="mm" value="7.5">7.5 mm</quantity>
|
||
long,
|
||
<quantity id="4CE59B310047FFC2FB27FC6EFAEC11B0" box="[1259,1346,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="85" pageNumber="86" unit="mm" value="3.8">3.8 mm</quantity>
|
||
wide in dorsal view, prothorax
|
||
<quantity id="4CE59B310047FFC2FE5BFC4AFE4311DC" box="[407,493,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="85" pageNumber="86" unit="mm" value="1.3">1.3 mm</quantity>
|
||
long. Wing (
|
||
<figureCitation id="13262A510047FFC2FDB2FC4AFD4B11DC" box="[638,741,943,969]" captionStart="FIGURE 43" captionStartId="77.[151,250,1943,1967]" captionTargetBox="[204,1385,181,1914]" captionTargetId="figure-17@77.[203,1385,181,1919]" captionTargetPageId="77" captionText="FIGURE 43. Okanagrion incertae sedis. SR 94-05-30: A, photograph; B, drawing; SR 11-51-07: C, photograph; D, drawing; RBCM P1550: E, photograph of top wing in G; F, drawing of wing in E; G, all three wings. A–D to scale; E, F to scale; G to scale; all scales 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558904" httpUri="https://zenodo.org/record/4558904/files/figure.png" pageId="85" pageNumber="86">Fig. 43B</figureCitation>
|
||
): CuA–A space two cells wide between level of origin of RP2, IR1. Legs. No spines or other setae visible; some coxae evident; metafemur
|
||
<quantity id="4CE59B310047FFC2FBCDFC37FBF511F8" box="[1025,1115,978,1005]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="85" pageNumber="86" unit="mm" value="5.6">5.6 mm</quantity>
|
||
, metatibia 5.0 mm, twice as long as abdominal segment 2. Wing 1. Mid-portion well preserved. Abdomen: colouration pale dorsally, extrapolated total length about
|
||
<quantity id="4CE59B310047FFC2FE6CFBFFFE5C1620" box="[416,498,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="85" pageNumber="86" unit="mm" value="45.0">45 mm</quantity>
|
||
; segment 2,
|
||
<quantity id="4CE59B310047FFC2FDB1FBFFFD7D1620" box="[637,723,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="85" pageNumber="86" unit="mm" value="2.5">2.5 mm</quantity>
|
||
long; segment 3,
|
||
<quantity id="4CE59B310047FFC2FC57FBFEFC5F1620" box="[923,1009,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="85" pageNumber="86" unit="mm" value="6.3">6.3 mm</quantity>
|
||
long x
|
||
<quantity id="4CE59B310047FFC2FB89FBFEFB351620" box="[1093,1179,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="85" pageNumber="86" unit="mm" value="1.3">1.3 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40047FFC2FF0BFBDAFEBE17F0" blockId="85.[151,1437,151,2013]" pageId="85" pageNumber="86">
|
||
<materialsCitation id="3B753C890047FFC2FF0BFBDAFCA616B4" ID-GBIF-Occurrence="3069296369" collectionCode="CuA" pageId="85" pageNumber="86" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54A688760047FFC2FF0BFBDAFE87164C" box="[199,297,1087,1113]" pageId="85" pageNumber="86" type="paratype">Paratype</typeStatus>
|
||
1, RBCM P1551, wing (
|
||
<figureCitation id="13262A510047FFC2FD8BFBDAFD33164C" box="[583,669,1086,1113]" captionStart="FIGURE 50" captionStartId="87.[151,250,2001,2025]" captionTargetBox="[243,1343,186,1972]" captionTargetId="figure-17@87.[231,1357,181,1977]" captionTargetPageId="87" captionText="FIGURE 50. Okanopteryx fraseri paratype 1, RBCM P1551. A, photograph; B, drawing. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558923" httpUri="https://zenodo.org/record/4558923/files/figure.png" pageId="85" pageNumber="86">Fig. 50</figureCitation>
|
||
). Missing the apical quarter and parts proximal to the nodus. Maximum width
|
||
<quantity id="4CE59B310047FFC2FED2FB86FED91668" box="[286,375,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="85" pageNumber="86" unit="mm" value="6.8">6.8 mm</quantity>
|
||
. Hyaline with broad dark fascia between level of nodus, level of terminus of
|
||
<collectionCode id="ED0CAE110047FFC2FB1AFB86FAA71668" box="[1238,1289,1123,1149]" pageId="85" pageNumber="86">CuA</collectionCode>
|
||
(presumably to half of wing distal to nodus, as in
|
||
<taxonomicName id="4C1D4D570047FFC2FDF0FB62FD5216B5" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[572,764,1159,1185]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="jeppesenorum">
|
||
<emphasis id="B969EAC60047FFC2FDF0FB62FDFE16B4" box="[572,592,1159,1185]" italics="true" pageId="85" pageNumber="86">O</emphasis>
|
||
.
|
||
<emphasis id="B969EAC60047FFC2FD93FB62FD5216B5" box="[607,764,1159,1184]" italics="true" pageId="85" pageNumber="86">jeppesenorum</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B753C890047FFC2FCD9FB62FBE216D0" ID-GBIF-Occurrence="3069296374" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP1" specimenCount="1" typeStatus="paratype">
|
||
Pterostigma not preserved. Most crossveins in postnodal, postsubnodal spaces aligned in region preserved (proximal half of these spaces).
|
||
<specimenCode id="DBBB9EAF0047FFC2FBD0FB4EFBE216D0" box="[1052,1100,1195,1221]" collectionCode="RP" pageId="85" pageNumber="86">RP1</specimenCode>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="3B753C890047FFC2FB90FB4EFBEB16FC" ID-GBIF-Occurrence="3069296310" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
|
||
IR1 space becomes two cells wide three cells distal to origin of IR1. IR1: origin four cells distal to origin of
|
||
<specimenCode id="DBBB9EAF0047FFC2FBDDFB2AFBEB16FC" box="[1041,1093,1231,1257]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B753C890047FFC2FB9DFB2AFE971718" ID-GBIF-Occurrence="3069296378" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
|
||
two cells wide ten cells distal to origin.
|
||
<specimenCode id="DBBB9EAF0047FFC2FEC9FB16FE971718" box="[261,313,1267,1293]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="3B753C890047FFC2FE88FB16FA361724" ID-GBIF-Occurrence="3069296340" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP3-4" specimenCount="1" typeStatus="paratype">
|
||
origin three cells distal to origin of IR2; linear, rather straight (distal quarter missing). IR2 origin at subnodus; straight, linear but slight zigzag near end of preserved portion (small portion missing near end).
|
||
<specimenCode id="DBBB9EAF0047FFC2FA81FAF2FA361724" box="[1357,1432,1303,1329]" collectionCode="RP" pageId="85" pageNumber="86">RP3-4</specimenCode>
|
||
</materialsCitation>
|
||
: origin half way between anterior-distal corner of quadrangle, nodus; almost complete: straight, linear. MA
|
||
<materialsCitation id="3B753C890047FFC2FA90FADEFC3B176C" ID-GBIF-Occurrence="3069296357" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
|
||
linear proximally, slight zigzag beginning between levels of origins of
|
||
<specimenCode id="DBBB9EAF0047FFC2FCAEFABAFC3B176C" box="[866,917,1375,1401]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
</materialsCitation>
|
||
, IR1, increasingly zigzagged to extreme apical to this. MP linear, rather straight from quadrangle to terminus. CuA
|
||
<materialsCitation id="3B753C890047FFC2FC68FA66FD3F17D4" ID-GBIF-Occurrence="3069296381" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
|
||
slightly zigzagged from level of quadrangle, increasing from about level of origin of
|
||
<specimenCode id="DBBB9EAF0047FFC2FD91FA42FD3F17D4" box="[605,657,1447,1473]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B753C890047FFC2FD51FA42FEA217F0" ID-GBIF-Occurrence="3069296370" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
|
||
CuA–A space two cells wide at widest (between levels of origins of IR1,
|
||
<specimenCode id="DBBB9EAF0047FFC2FF03FA2EFEAA17F0" box="[207,260,1483,1509]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40047FFC2FF0BFA0AFEFE1444" blockId="85.[151,1437,151,2013]" pageId="85" pageNumber="86">
|
||
<materialsCitation id="3B753C890047FFC2FF0BFA0AFEE31444" ID-GBIF-Occurrence="3069296326" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP2" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54A688760047FFC2FF0BFA0AFE87141C" box="[199,297,1519,1545]" pageId="85" pageNumber="86" type="paratype">Paratype</typeStatus>
|
||
2, F-1091, wing (
|
||
<figureCitation id="13262A510047FFC2FE39FA0AFDE7141C" box="[501,585,1518,1545]" captionStart="FIGURE 51" captionStartId="88.[151,250,1934,1958]" captionTargetBox="[154,1427,181,1898]" captionTargetId="figure-19@88.[151,1436,181,1910]" captionTargetPageId="88" captionText="FIGURE 51. Okanopteryx fraseri paratype 3, PB-4157. A, photograph; B, drawing; paratype 2, F-1091: C, photograph; D, drawing.All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558926" httpUri="https://zenodo.org/record/4558926/files/figure.png" pageId="85" pageNumber="86">Fig. 51</figureCitation>
|
||
). As RBCM P1551, except: antenodal space preserved, no accessory crossveins. Width
|
||
<quantity id="4CE59B310047FFC2FEE1F9F6FE281438" box="[301,390,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="85" pageNumber="86" unit="mm" value="6.6">6.6 mm</quantity>
|
||
. IR1: origin six cells distal to that of
|
||
<specimenCode id="DBBB9EAF0047FFC2FCE6F9F6FCF01438" box="[810,862,1555,1581]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
. CuA–A space two cells wide at widest except one column of three
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40047FFC2FF0BF9BEFA9A14F4" blockId="85.[151,1437,151,2013]" pageId="85" pageNumber="86">
|
||
<materialsCitation id="3B753C890047FFC2FF0BF9BEFA9F14F4" ID-GBIF-Occurrence="3069296304" collectionCode="RP" pageId="85" pageNumber="86" specimenCode="RP3-4, RP2, RP1" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54A688760047FFC2FF0BF9BEFE871460" box="[199,297,1627,1653]" pageId="85" pageNumber="86" type="paratype">Paratype</typeStatus>
|
||
3, PB-4157, wing (
|
||
<figureCitation id="13262A510047FFC2FDC0F9BEFDCF1460" box="[524,609,1626,1653]" captionStart="FIGURE 51" captionStartId="88.[151,250,1934,1958]" captionTargetBox="[154,1427,181,1898]" captionTargetId="figure-19@88.[151,1436,181,1910]" captionTargetPageId="88" captionText="FIGURE 51. Okanopteryx fraseri paratype 3, PB-4157. A, photograph; B, drawing; paratype 2, F-1091: C, photograph; D, drawing.All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558926" httpUri="https://zenodo.org/record/4558926/files/figure.png" pageId="85" pageNumber="86">Fig. 51</figureCitation>
|
||
). As RBCM P1551, except the following. Much of antenodal space preserved, no accessory crossveins.
|
||
<specimenCode id="DBBB9EAF0047FFC2FDDEF99AFDF0148C" box="[530,606,1663,1689]" collectionCode="RP" pageId="85" pageNumber="86">RP3-4</specimenCode>
|
||
: origin two thirds distance between anterior-distal corner of quadrangle, nodus. IR1: origin two cells distal to origin of
|
||
<specimenCode id="DBBB9EAF0047FFC2FD65F946FD7314A8" box="[681,733,1699,1725]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
.
|
||
<specimenCode id="DBBB9EAF0047FFC2FD24F946FCB614A8" box="[744,792,1699,1725]" collectionCode="RP" pageId="85" pageNumber="86">RP1</specimenCode>
|
||
–IR1 space two cells wide six cells distal to origin.
|
||
<specimenCode id="DBBB9EAF0047FFC2FAA8F946FA3614A8" box="[1380,1432,1699,1725]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
: origin 3.5 cells distal to origin of IR2. CuA–A space two cells wide at widest except one column of three
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40047FFC2FF0BF90EFD8E1580" blockId="85.[151,1437,151,2013]" pageId="85" pageNumber="86">
|
||
<materialsCitation id="3B753C890047FFC2FF0BF90EFDB21580" ID-GBIF-Occurrence="3069296329" collectionCode="MP, CuA" pageId="85" pageNumber="86" specimenCode="RP1, RP2, RP3-4" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54A688760047FFC2FF0BF90EFE871510" box="[199,297,1771,1797]" pageId="85" pageNumber="86" type="paratype">Paratype</typeStatus>
|
||
4, F-1045, wing (
|
||
<figureCitation id="13262A510047FFC2FE34F90EFDE01510" box="[504,590,1770,1797]" captionStart="FIGURE 52" captionStartId="89.[151,250,1954,1978]" captionTargetBox="[361,1219,184,1913]" captionTargetId="figure-17@89.[353,1235,181,1930]" captionTargetPageId="89" captionText="FIGURE 52. Okanopteryx fraseri paratype 5, RBCM P1552. A, photograph; B, drawing; paratype 4, F-1045: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558928" httpUri="https://zenodo.org/record/4558928/files/figure.png" pageId="85" pageNumber="86">Fig. 52</figureCitation>
|
||
). As RBCM P1551, except: almost all antenodal space missing. Width
|
||
<quantity id="4CE59B310047FFC2FAB5F90EFF68153C" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="85" pageNumber="86" unit="mm" value="6.4">6.4 mm</quantity>
|
||
.
|
||
<specimenCode id="DBBB9EAF0047FFC2FF1EF8EAFEAF153C" box="[210,257,1807,1833]" collectionCode="RP" pageId="85" pageNumber="86">RP1</specimenCode>
|
||
–IR1 space goes to two cells wide six cells distal to origin of IR1. IR1: origin three cells distal to origin of
|
||
<specimenCode id="DBBB9EAF0047FFC2FF7AF8D6FF441558" box="[182,234,1843,1869]" collectionCode="RP" pageId="85" pageNumber="86">RP2</specimenCode>
|
||
.
|
||
<specimenCode id="DBBB9EAF0047FFC2FF39F8D6FE921558" box="[245,316,1843,1869]" collectionCode="RP" pageId="85" pageNumber="86">RP3-4</specimenCode>
|
||
origin: quadrangle not preserved, but similar distance from nodus.
|
||
<collectionCode id="ED0CAE110047FFC2FBFFF8D6FBF41558" box="[1075,1114,1843,1869]" country="0" name="Mohonk Preserve, Inc." pageId="85" pageNumber="86">MP</collectionCode>
|
||
missing near terminus.
|
||
<collectionCode id="ED0CAE110047FFC2FAA5F8D6FA321558" box="[1385,1436,1843,1869]" pageId="85" pageNumber="86">CuA</collectionCode>
|
||
slightly zigzagged from level of origin of
|
||
<specimenCode id="DBBB9EAF0047FFC2FDB5F8B2FD6A1564" box="[633,708,1879,1905]" collectionCode="RP" pageId="85" pageNumber="86">RP3-4</specimenCode>
|
||
, increasing from about level of origin of IR1; two cells wide at widest except one column of three
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40047FFCDFF0BF87AFE091354" blockId="85.[151,1437,151,2013]" lastBlockId="90.[151,1437,151,825]" lastPageId="90" lastPageNumber="91" pageId="85" pageNumber="86">
|
||
<materialsCitation id="3B753C890047FFCDFF0BF87AFE0D1354" ID-GBIF-Occurrence="3069296364" collectionCode="MA, CuA" lastPageId="90" lastPageNumber="91" pageId="85" pageNumber="86" specimenCode="RP1, RP2" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54A688760047FFC2FF0BF87AFE8715AC" box="[199,297,1951,1977]" pageId="85" pageNumber="86" type="paratype">Paratype</typeStatus>
|
||
5, RBCM P1552, wing (
|
||
<figureCitation id="13262A510047FFC2FD82F87AFD0B15AC" box="[590,677,1950,1977]" captionStart="FIGURE 52" captionStartId="89.[151,250,1954,1978]" captionTargetBox="[361,1219,184,1913]" captionTargetId="figure-17@89.[353,1235,181,1930]" captionTargetPageId="89" captionText="FIGURE 52. Okanopteryx fraseri paratype 5, RBCM P1552. A, photograph; B, drawing; paratype 4, F-1045: C, photograph; D, drawing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558928" httpUri="https://zenodo.org/record/4558928/files/figure.png" pageId="85" pageNumber="86">Fig. 52</figureCitation>
|
||
). Smallest portion, from about base of IR1 to
|
||
<specimenCode id="DBBB9EAF0047FFC2FB73F87AFB4015AC" box="[1215,1262,1951,1977]" collectionCode="RP" pageId="85" pageNumber="86">RP1</specimenCode>
|
||
–IR1 space becoming two cells wide. Width
|
||
<quantity id="4CE59B310047FFC2FDC9F826FDCD15C8" box="[517,611,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="85" pageNumber="86" unit="mm" value="6.8">6.8 mm</quantity>
|
||
. Darkly infuscate throughout. Preserved portion as for RBCM P1551, except:
|
||
<specimenCode id="DBBB9EAF0048FFCDFF3DFF72FE8E12A4" box="[241,288,151,177]" collectionCode="RP" pageId="90" pageNumber="91">RP1</specimenCode>
|
||
–IR1 space becomes two cells wide five cells distal to origin of IR1. IR1: origin three cells distal to origin of
|
||
<specimenCode id="DBBB9EAF0048FFCDFEC9FF5EFE9712C0" box="[261,313,187,213]" collectionCode="RP" pageId="90" pageNumber="91">RP2</specimenCode>
|
||
.
|
||
<specimenCode id="DBBB9EAF0048FFCDFE8BFF5EFED512C0" box="[327,379,187,213]" collectionCode="RP" pageId="90" pageNumber="91">RP2</specimenCode>
|
||
: origin three cells distal to origin of IR2 by preservation, but probably four.
|
||
<collectionCode id="ED0CAE110048FFCDFB34FF5EFA8A12C0" box="[1272,1316,187,213]" pageId="90" pageNumber="91">MA</collectionCode>
|
||
becoming zigzagged about level of origin of IR1.
|
||
<collectionCode id="ED0CAE110048FFCDFD9FFF3AFD2812EC" box="[595,646,223,249]" pageId="90" pageNumber="91">CuA</collectionCode>
|
||
probably becoming zigzagged about level of origin of IR2, increasing distal to it. CuA–A space apparently two cells wide at widest, unsure by preservation if a third cell wide as in F- 1091, PB-4157, F-1045
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF62665C0044FFC1FF5BF872FDB015C6" ID-DOI="http://doi.org/10.5281/zenodo.4558920" ID-Zenodo-Dep="4558920" httpUri="https://zenodo.org/record/4558920/files/figure.png" pageId="86" pageNumber="87" startId="86.[151,250,1943,1967]" targetBox="[166,1422,181,1919]" targetPageId="86">
|
||
<paragraph id="8BA236D40044FFC1FF5BF872FDB015C6" blockId="86.[151,1437,1943,2004]" pageId="86" pageNumber="87">
|
||
<emphasis id="B969EAC60044FFC1FF5BF872FE8C15BA" bold="true" box="[151,290,1943,1967]" pageId="86" pageNumber="87">FIGURE 49.</emphasis>
|
||
<taxonomicName id="4C1D4D570044FFC1FEE7F87DFE5515A5" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[299,507,1944,1968]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B969EAC60044FFC1FEE7F87DFE5515A5" box="[299,507,1944,1968]" italics="true" pageId="86" pageNumber="87">Okanopteryx fraseri</emphasis>
|
||
</taxonomicName>
|
||
holotype, GSC 141104/F-778. A, photograph of entire specimen (part: GSC 141104); B, drawing of wing. Both to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF62665C0045FFC0FF5BF834FB5515FC" ID-DOI="http://doi.org/10.5281/zenodo.4558923" ID-Zenodo-Dep="4558923" box="[151,1275,2001,2026]" httpUri="https://zenodo.org/record/4558923/files/figure.png" pageId="87" pageNumber="88" startId="87.[151,250,2001,2025]" targetBox="[243,1343,186,1972]" targetPageId="87">
|
||
<paragraph id="8BA236D40045FFC0FF5BF834FB5515FC" blockId="87.[151,1275,2001,2026]" box="[151,1275,2001,2026]" pageId="87" pageNumber="88">
|
||
<emphasis id="B969EAC60045FFC0FF5BF834FE8E15FC" bold="true" box="[151,288,2001,2025]" pageId="87" pageNumber="88">FIGURE 50.</emphasis>
|
||
<taxonomicName id="4C1D4D570045FFC0FEEAF834FE5D15FC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[294,499,2001,2025]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B969EAC60045FFC0FEEAF834FE5D15FC" box="[294,499,2001,2025]" italics="true" pageId="87" pageNumber="88">Okanopteryx fraseri</emphasis>
|
||
</taxonomicName>
|
||
paratype 1, RBCM P1551. A, photograph; B, drawing. Both to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF62665C004AFFCFFF5BF86BFE1915DF" ID-DOI="http://doi.org/10.5281/zenodo.4558926" ID-Zenodo-Dep="4558926" httpUri="https://zenodo.org/record/4558926/files/figure.png" pageId="88" pageNumber="89" startId="88.[151,250,1934,1958]" targetBox="[154,1427,181,1898]" targetPageId="88">
|
||
<paragraph id="8BA236D4004AFFCFFF5BF86BFE1915DF" blockId="88.[151,1436,1934,1995]" pageId="88" pageNumber="89">
|
||
<emphasis id="B969EAC6004AFFCFFF5BF86BFE8C15B3" bold="true" box="[151,290,1934,1958]" pageId="88" pageNumber="89">FIGURE 51.</emphasis>
|
||
<taxonomicName id="4C1D4D57004AFFCFFEE7F86AFE5415B2" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[299,506,1935,1959]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B969EAC6004AFFCFFEE7F86AFE5415B2" box="[299,506,1935,1959]" italics="true" pageId="88" pageNumber="89">Okanopteryx fraseri</emphasis>
|
||
</taxonomicName>
|
||
paratype 3, PB-4157. A, photograph; B, drawing; paratype 2, F-1091: C, photograph; D, drawing. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF62665C004BFFCEFF5BF847FE7815C8" ID-DOI="http://doi.org/10.5281/zenodo.4558928" ID-Zenodo-Dep="4558928" httpUri="https://zenodo.org/record/4558928/files/figure.png" pageId="89" pageNumber="90" startId="89.[151,250,1954,1978]" targetBox="[361,1219,184,1913]" targetPageId="89">
|
||
<paragraph id="8BA236D4004BFFCEFF5BF847FE7815C8" blockId="89.[151,1436,1954,2014]" pageId="89" pageNumber="90">
|
||
<emphasis id="B969EAC6004BFFCEFF5BF847FE8F15AF" bold="true" box="[151,289,1954,1978]" pageId="89" pageNumber="90">FIGURE 52.</emphasis>
|
||
<taxonomicName id="4C1D4D57004BFFCEFEE4F847FE5815AF" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[296,502,1954,1978]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B969EAC6004BFFCEFEE4F847FE5815AF" box="[296,502,1954,1978]" italics="true" pageId="89" pageNumber="90">Okanopteryx fraseri</emphasis>
|
||
</taxonomicName>
|
||
paratype 5, RBCM P1552. A, photograph; B, drawing; paratype 4, F-1045: C, photograph; D, drawing. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C307655F0048FFCDFF0BFEAEFC8713B8" pageId="90" pageNumber="91" type="etymology">
|
||
<paragraph id="8BA236D40048FFCDFF0BFEAEFD77139C" blockId="90.[151,1437,151,825]" pageId="90" pageNumber="91">
|
||
<emphasis id="B969EAC60048FFCDFF0BFEAEFEE61370" bold="true" box="[199,328,331,357]" pageId="90" pageNumber="91">Etymology</emphasis>
|
||
. The specific epithet is a patronymic formed from the surname of John Fraser, the collector and donor of the
|
||
<typeStatus id="54A688760048FFCDFEEBFE8AFE22139C" box="[295,396,367,393]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
|
||
, recognising his contribution.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40048FFCDFF0BFE76FC8713B8" blockId="90.[151,1437,151,825]" box="[199,809,403,429]" pageId="90" pageNumber="91">
|
||
<emphasis id="B969EAC60048FFCDFF0BFE76FED913B8" bold="true" box="[199,375,403,429]" pageId="90" pageNumber="91">Range and age</emphasis>
|
||
. McAbee, BC,
|
||
<collectingCountry id="F30A76440048FFCDFDEBFE76FDD113B8" box="[551,639,403,429]" name="Canada" pageId="90" pageNumber="91">Canada</collectingCountry>
|
||
; mid-Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C307655F0048FFCDFF0BFE53FDCE112C" pageId="90" pageNumber="91" type="discussion">
|
||
<paragraph id="8BA236D40048FFCDFF0BFE53FE501028" blockId="90.[151,1437,151,825]" pageId="90" pageNumber="91">
|
||
<emphasis id="B969EAC60048FFCDFF0BFE53FEE213C5" bold="true" box="[199,332,438,464]" pageId="90" pageNumber="91">Discussion.</emphasis>
|
||
The abdomen of the
|
||
<typeStatus id="54A688760048FFCDFDF5FE52FD3513C4" box="[569,667,439,465]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
|
||
is much longer than that of
|
||
<taxonomicName id="4C1D4D570048FFCDFC18FE52FB2B13C4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[980,1157,439,465]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="macabeensis">
|
||
<emphasis id="B969EAC60048FFCDFC18FE52FB2B13C4" box="[980,1157,439,465]" italics="true" pageId="90" pageNumber="91">O. macabeensis</emphasis>
|
||
</taxonomicName>
|
||
; indeed, the specimen as a whole is likely about 1.7 times longer than
|
||
<taxonomicName id="4C1D4D570048FFCDFD43FE3EFCEE13E0" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[655,832,475,501]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="macabeensis">
|
||
<emphasis id="B969EAC60048FFCDFD43FE3EFCEE13E0" box="[655,832,475,501]" italics="true" pageId="90" pageNumber="91">O. macabeensis</emphasis>
|
||
</taxonomicName>
|
||
, confidently setting these two species apart (see diagnosis). Based on known variation in extant species, this size difference is unlikely to be the result of intraspecific variation or sexual dimorphism.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40048FFCDFF0BFDA2FE2010E4" blockId="90.[151,1437,151,825]" pageId="90" pageNumber="91">
|
||
Although the wing of the
|
||
<typeStatus id="54A688760048FFCDFE27FDA2FDE31074" box="[491,589,583,609]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1D4D570048FFCDFDB8FDA2FD4C1074" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[628,738,583,609]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B969EAC60048FFCDFDB8FDA2FD4C1074" box="[628,738,583,609]" italics="true" pageId="90" pageNumber="91">O. fraseri</emphasis>
|
||
</taxonomicName>
|
||
is only partially preserved, its CuA–A space is only two cells wide at this level, with an occasional single column of three. By this, we associate wings which share the colouration and other venational character states with either
|
||
<taxonomicName id="4C1D4D570048FFCDFD77FD6AFC8710BC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[699,809,655,681]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B969EAC60048FFCDFD77FD6AFC8710BC" box="[699,809,655,681]" italics="true" pageId="90" pageNumber="91">O. fraseri</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C1D4D570048FFCDFC83FD6AFBAF10BC" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[847,1025,655,681]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="macabeensis">
|
||
<emphasis id="B969EAC60048FFCDFC83FD6AFBAF10BC" box="[847,1025,655,681]" italics="true" pageId="90" pageNumber="91">O. macabeensis</emphasis>
|
||
</taxonomicName>
|
||
based on the number of cells across the CuA–A space between the levels of IR1 and RP2: three in
|
||
<taxonomicName id="4C1D4D570048FFCDFC81FD56FC5010D8" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[845,1022,691,717]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="macabeensis">
|
||
<emphasis id="B969EAC60048FFCDFC81FD56FC5010D8" box="[845,1022,691,717]" italics="true" pageId="90" pageNumber="91">O. macabeensis</emphasis>
|
||
</taxonomicName>
|
||
and two (sometimes a single column of three) in
|
||
<taxonomicName id="4C1D4D570048FFCDFED7FD32FE2610E4" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[283,392,727,753]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B969EAC60048FFCDFED7FD32FE2610E4" box="[283,392,727,753]" italics="true" pageId="90" pageNumber="91">O. fraseri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA236D40048FFCDFF0BFD1EFDCE112C" blockId="90.[151,1437,151,825]" pageId="90" pageNumber="91">
|
||
The thorax of
|
||
<taxonomicName id="4C1D4D570048FFCDFEA5FD1EFE761100" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[361,472,763,789]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="fraseri">
|
||
<emphasis id="B969EAC60048FFCDFEA5FD1EFE761100" box="[361,472,763,789]" italics="true" pageId="90" pageNumber="91">O. fraseri</emphasis>
|
||
</taxonomicName>
|
||
may be more robust than that of
|
||
<taxonomicName id="4C1D4D570048FFCDFC9EFD1EFBAB1100" authorityName="Archibald & Cannings & Erickson & Bybee & Mathewes" authorityYear="2021" box="[850,1029,763,789]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="macabeensis">
|
||
<emphasis id="B969EAC60048FFCDFC9EFD1EFBAB1100" box="[850,1029,763,789]" italics="true" pageId="90" pageNumber="91">O. macabeensis</emphasis>
|
||
</taxonomicName>
|
||
, however, it might also be that they are preserved at slightly different angles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |