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<document ID-DOI="http://doi.org/10.5281/zenodo.6397752" ID-GBIF-Dataset="45351c32-25dd-422c-bdb2-00e73deb4943" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6397752" approvalRequired="37" approvalRequired_for_taxonomicNames="8" approvalRequired_for_treatments="29" checkinTime="1648655544658" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="4C3D87E8FFC06A7FFF89910519B9B276" docLanguage="en" docName="hbmw_9_Vespertilionidae_716.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Mimetillus moloneyi Thomas 1891" docType="treatment" docVersion="9" lastPageNumber="817" masterDocId="B004FF90FFFB6A44FFFC96591E00BB32" masterDocTitle="Vespertilionidae" masterLastPageNumber="981" masterPageNumber="716" pageNumber="817" updateTime="1658412944878" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Vespertilionidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>716</mods:start>
<mods:end>981</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6397752</mods:identifier>
<mods:identifier type="GBIF-Dataset">45351c32-25dd-422c-bdb2-00e73deb4943</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">6397752</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6403502" ID-GBIF-Taxon="195628476" ID-Zenodo-Dep="6403502" LSID="urn:lsid:plazi:treatment:4C3D87E8FFC06A7FFF89910519B9B276" httpUri="http://treatment.plazi.org/id/4C3D87E8FFC06A7FFF89910519B9B276" lastPageNumber="817" pageId="59" pageNumber="817">
<subSubSection box="[117,196,1884,1934]" pageId="59" pageNumber="817" type="multiple">
<paragraph blockId="59.[112,1209,1886,2018]" box="[117,196,1884,1934]" pageId="59" pageNumber="817">
<heading box="[117,196,1884,1934]" pageId="59" pageNumber="817">
<figureCitation box="[117,196,1884,1934]" captionStart="Plate 59" captionStartId="51.[133,163,3283,3308]" captionTargetBox="[12,2757,17,3656]" captionTargetPageId="50" captionText="94. Fairy Pipistrelle (Hypsugo ariel), 95. Lanzas Pipistrelle (Hypsugo lanzai), 96. Arabian Pipistrelle (Hypsugo arabicus), 97. Savis Pipistrelle (Hypsugo savii), 98. Alashanian Pipistrelle (Hypsugo alaschanicus), 99. Cadornas Pipistrelle (Hypsugo cadornae), 100. Joftres Pipistrelle (Hypsugo joffrei), 101. Chinese Pipistrelle (Hypsugo pulveratus), 102. Chocolate Pipistrelle (Hypsugo affinis), 103. Pungent Pipistrelle (Hypsugo mordax), 104. Peterss Pipistrelle (Hypsugo petersi), 105. Long-toothed Pipistrelle (Hypsugo dolichodon), 106. Burmese Pipistrelle (Hypsugo lophurus), 107. Red-brown Pipistrelle (Hypsugo kitcheneri), 108. Brown Pipistrelle (Hypsugo imbricatus), 109. Big-eared Pipistrelle (Hypsugo macrotis), 110. Vordermanns Pipistrelle (Hypsugo vordermanni), 111. Anchietas Pipistrelle (Hypsugo anchieta), 112. Kirindy Pipistrelle (Hypsugo bemainty), 113. Mouse-like Pipistrelle (Hypsugo musciculus), 114. Broad-headed Pipistrelle (Hypsugo crassulus), 115. Eisentrauts Pipistrelle (Hypsugo eisentrauti), 116. Schlieffens Bat (Nycticeinops schlieffenii), 117. Moloneys Mimic Bat (Mimetillus moloneyi)" figureDoi="http://doi.org/10.5281/zenodo.6398514" httpUri="https://zenodo.org/record/6398514/files/figure.png" pageId="59" pageNumber="817">117.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[214,654,1884,1934]" pageId="59" pageNumber="817" type="vernacular_names">
<paragraph blockId="59.[112,1209,1886,2018]" box="[214,654,1884,1934]" pageId="59" pageNumber="817">
<heading box="[214,654,1884,1934]" pageId="59" pageNumber="817">
<vernacularName box="[214,654,1884,1934]" pageId="59" pageNumber="817">Moloneys Mimic Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[704,1082,1890,1936]" pageId="59" pageNumber="817" type="nomenclature">
<paragraph blockId="59.[112,1209,1886,2018]" box="[704,1082,1890,1936]" pageId="59" pageNumber="817">
<heading box="[704,1082,1890,1936]" pageId="59" pageNumber="817">
<taxonomicName authorityName="Thomas" authorityYear="1891" box="[704,1082,1890,1936]" class="Mammalia" family="Vespertilionidae" genus="Mimetillus" kingdom="Animalia" order="Chiroptera" pageId="59" pageNumber="817" phylum="Chordata" rank="species" species="moloneyi">
<emphasis box="[704,1082,1890,1936]" italics="true" pageId="59" pageNumber="817">Mimetillus moloneyi</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="vernacular_names">
<paragraph blockId="59.[112,1209,1886,2018]" box="[114,1209,1953,1974]" pageId="59" pageNumber="817">
<heading box="[114,1209,1953,1974]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[114,189,1953,1974]" pageId="59" pageNumber="817">French:</emphasis>
<vernacularName box="[199,418,1953,1974]" pageId="59" pageNumber="817">Mimétille de Moloney</vernacularName>
/
<emphasis bold="true" box="[438,529,1953,1974]" pageId="59" pageNumber="817">German:</emphasis>
<vernacularName box="[539,873,1953,1974]" pageId="59" pageNumber="817">Moloney-Schmalfligelfledermaus</vernacularName>
/
<emphasis bold="true" box="[893,984,1953,1974]" pageId="59" pageNumber="817">Spanish:</emphasis>
<vernacularName box="[995,1209,1953,1974]" pageId="59" pageNumber="817">Mimétilo de Moloney</vernacularName>
</heading>
</paragraph>
<paragraph blockId="59.[112,1209,1886,2018]" box="[113,991,1993,2014]" pageId="59" pageNumber="817">
<heading box="[113,991,1993,2014]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[113,360,1993,2014]" pageId="59" pageNumber="817">Other common names:</emphasis>
<vernacularName box="[370,638,1993,2014]" pageId="59" pageNumber="817">Moloney's Flat-headed Bat</vernacularName>
,
<vernacularName box="[652,991,1993,2014]" pageId="59" pageNumber="817">Moloney's Flat-headed Vesper Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[112,861,2060,2093]" pageId="59" pageNumber="817" type="reference_group">
<paragraph blockId="59.[111,1320,2060,2376]" box="[112,861,2060,2093]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[112,266,2060,2093]" pageId="59" pageNumber="817">Taxonomy.</emphasis>
<taxonomicName authority="Thomas, 1891" authorityName="Thomas" authorityYear="1891" box="[280,855,2060,2093]" class="Mammalia" family="Vespertilionidae" genus="Vesperugo" kingdom="Animalia" order="Chiroptera" pageId="59" pageNumber="817" phylum="Chordata" rank="species" species="moloneyi" subGenus="Vesperus">Vesperugo (Vesperus) moloneyi Thomas, 1891</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection box="[871,1316,2060,2093]" pageId="59" pageNumber="817" type="materials_examined">
<paragraph blockId="59.[111,1320,2060,2376]" box="[871,1316,2060,2093]" pageId="59" pageNumber="817">
<materialsCitation ID-GBIF-Occurrence="3783319740" box="[871,1316,2060,2093]" pageId="59" pageNumber="817">
<collectingRegion box="[885,966,2060,2093]" country="Nigeria" name="Lagos" pageId="59" pageNumber="817">Lagos</collectingRegion>
, West Africa [=
<collectingCountry box="[1180,1290,2060,2093]" name="Nigeria" pageId="59" pageNumber="817">Nigeria</collectingCountry>
].”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="discussion">
<paragraph blockId="59.[111,1320,2060,2376]" pageId="59" pageNumber="817">
Phylogenetic relationship of
<taxonomicName authorityName="Thomas" authorityYear="1904" box="[519,655,2099,2132]" class="Mammalia" family="Vespertilionidae" genus="Mimetillus" kingdom="Animalia" order="Chiroptera" pageId="59" pageNumber="817" phylum="Chordata" rank="genus">Mimetillus</taxonomicName>
to other genera of Vespertilioniniis still uncertain. As currently recognized, there may be multiple species included within
<taxonomicName authorityName="Thomas" authorityYear="1891" class="Mammalia" family="Vespertilionidae" genus="Mimetillus" kingdom="Animalia" order="Chiroptera" pageId="59" pageNumber="817" phylum="Chordata" rank="species" species="moloneyi">M. moloneyi</taxonomicName>
and subspecies thomasi has been recognized as a distinct species on morphological grounds. Race berneri has been treated also as a synonym of thomas. However, a full revision of the validity of the recognized subspecies is pending and only a single species is recognized tentatively. Three subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6398018" ID-Zenodo-Dep="6398018" httpUri="https://zenodo.org/record/6398018/files/figure.png" inLine="true" pageId="59" pageNumber="817" targetBox="[1390,1982,299,713]" targetPageId="59">
<paragraph blockId="59.[111,1320,2060,2376]" box="[112,513,2335,2368]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[112,513,2335,2368]" pageId="59" pageNumber="817">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph blockId="59.[2004,2598,287,715]" pageId="59" pageNumber="817">
<taxonomicName authority=": Thomas, 1891" authorityName=": Thomas" authorityYear="1891" box="[2006,2424,287,320]" class="Mammalia" family="Vespertilionidae" genus="Mimetillus" kingdom="Animalia" order="Chiroptera" pageId="59" pageNumber="817" phylum="Chordata" rank="subSpecies" species="moloneyi" subSpecies="moloney">M.m.moloney:Thomas,1891—scatteredfromSEGuineaandSierraLeoneEtoSWEthiopia,Kenya,andNETanzania(includingBiokoI),andStoNWAngolaandextremeWDRCongo.</taxonomicName>
</paragraph>
<paragraph blockId="59.[2004,2598,287,715]" pageId="59" pageNumber="817">
<taxonomicName authority="Monard, 1933" authorityName="Monard" authorityYear="1933" box="[2006,2394,493,518]" class="Mammalia" family="Vespertilionidae" genus="Mimetillus" kingdom="Animalia" order="Chiroptera" pageId="59" pageNumber="817" phylum="Chordata" rank="subSpecies" species="moloneyi" subSpecies="berneri">M.m.berneriMonard,1933—endemictotheAngolanPlateau.</taxonomicName>
</paragraph>
<paragraph blockId="59.[2004,2598,287,715]" pageId="59" pageNumber="817">
<taxonomicName authority=": Hinton, 1920" authorityName=": Hinton" authorityYear="1920" box="[2006,2391,568,597]" class="Mammalia" family="Vespertilionidae" genus="Mimetillus" kingdom="Animalia" order="Chiroptera" pageId="59" pageNumber="817" phylum="Chordata" rank="subSpecies" species="moloneyi" subSpecies="thomas">M. m. thomas: Hinton, 1920</taxonomicName>
— SE
<collectingCountry name="Democratic Republic of the Congo" pageId="59" pageNumber="817">DR Congo</collectingCountry>
, S
<collectingCountry box="[2098,2226,607,636]" name="Tanzania" pageId="59" pageNumber="817">Tanzania</collectingCountry>
,
<collectingCountry box="[2251,2358,607,636]" name="Zambia" pageId="59" pageNumber="817">Zambia</collectingCountry>
, and S
<collectingCountry name="Mozambique" pageId="59" pageNumber="817">Mozambique</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="description">
<paragraph blockId="59.[2004,2598,287,715]" lastBlockId="59.[1387,2600,725,2372]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[2004,2283,682,715]" pageId="59" pageNumber="817">Descriptive notes.</emphasis>
Head-body c¢.51-63 mm, tail 23-32 mm, ear 9-14 mm, hindfoot 5-9 mm, forearm 27-31 mm; weight 5:5-10 g. Fur of Moloneys Mimic Bat is velvety; dorsally and ventrally dark chocolatebrown to almost black, with unicolored hairs. Ears are blackish, naked, short (compared to other vespertilionids), and pointed, with short, mushroom-shaped tragus. Muzzle is broad and has large glands in sexually active individuals. Wings are dark brown to black, extremely short, and narrow. Finger-wing is translucent. Phalanges of third, fourth, and fifth fingers are relatively short. Legs are very short and tail is long. Penis is long, without evidence of a baculum. Braincase is flattened, rostrum shortened and broad, with well-developed supraorbital tubercles, and pronounced postorbital constriction; sagittal crest is very low. Wing loading is higher than in any other vespertilionid, except Leislers Noctule (
<taxonomicName baseAuthorityName="Kuhl" baseAuthorityYear="1817" box="[1966,2175,1115,1148]" class="Mammalia" family="Vespertilionidae" genus="Nyctalus" kingdom="Animalia" order="Chiroptera" pageId="59" pageNumber="817" phylum="Chordata" rank="species" species="leisleri">Nyctalus leisleri</taxonomicName>
). 1° is comparatively long and lateral to I*; C' has no secondary cusp; and P* exceeds height of molars. Dental formula 1512/3, C1/1, P1722, M 3/3 (#2) = 32.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="biology_ecology">
<paragraph blockId="59.[1387,2600,725,2372]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[1391,1502,1234,1267]" pageId="59" pageNumber="817">Habitat.</emphasis>
Nominate moloney: inhabits forest-savanna mosaic and savanna habitats; it has been recorded in lowland rainforest, coastal forest, montane forests, mangroves, forest-savanna mosaics,
<taxonomicName box="[1733,1819,1317,1346]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="genus">Acacia</taxonomicName>
(
<taxonomicName box="[1844,1962,1317,1346]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
)—
<taxonomicName box="[2002,2170,1317,1346]" class="Magnoliopsida" family="Burseraceae" genus="Commiphora" kingdom="Plantae" order="Sapindales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="genus">Commiphora</taxonomicName>
(
<taxonomicName box="[2196,2370,1317,1346]" class="Magnoliopsida" family="Burseraceae" kingdom="Plantae" order="Sapindales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="family">Burseraceae</taxonomicName>
) bushland and thicket, and
<taxonomicName box="[1578,1709,1352,1385]" class="Magnoliopsida" family="Fabaceae" genus="Isoberlinia" kingdom="Plantae" order="Fabales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="genus">Isoberlinia</taxonomicName>
(
<taxonomicName box="[1735,1866,1352,1385]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
) woodland. It apparently prefers edge of rainforest from which it penetrates into forest-savanna mosaics and woodlands. Reported to elevations of up to
<quantity box="[1676,1784,1431,1464]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="59" pageNumber="817" unit="m" value="2300.0">2300 m</quantity>
. Race thomas: mostly inhabits savanna, but has been recorded from mopane and miombo woodlands,
<taxonomicName authorityName="Bentham" authorityYear="1865" box="[2084,2269,1470,1503]" class="Magnoliopsida" family="Fabaceae" genus="Cryptosepalum" kingdom="Plantae" order="Fabales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="genus">Cryptosepalum</taxonomicName>
(
<taxonomicName box="[2293,2425,1470,1503]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
) dry forest, and coastal forests in southern
<collectingCountry box="[1835,1963,1514,1543]" name="Tanzania" pageId="59" pageNumber="817">Tanzania</collectingCountry>
. Race berneri also occurs in savanna habitats, with records from miombo woodland and montane forest-grassland mosaic on the Angolan Plateau.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="food_feeding">
<paragraph blockId="59.[1387,2600,725,2372]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[1390,1651,1628,1661]" pageId="59" pageNumber="817">Food and Feeding.</emphasis>
Moloneys Mimic Bat feeds on small winged termites, and perhaps other small to medium-sized flying insects.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="breeding">
<paragraph blockId="59.[1387,2600,725,2372]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[1389,1524,1707,1740]" pageId="59" pageNumber="817">Breeding.</emphasis>
The little information available is indicative of a seasonal bimodal polyestry, but asynchronous breeding; more than two births per year cannot be discarded. Litter size 1s one.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="activity">
<paragraph blockId="59.[1387,2600,725,2372]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[1387,1625,1825,1858]" pageId="59" pageNumber="817">Activity patterns.</emphasis>
Moloneys Mimic Bat leaves the roost to forage at dusk, with some bats returning to the day roost after 10-15 minutes. In
<collectingCountry box="[2147,2249,1865,1898]" name="Liberia" pageId="59" pageNumber="817">Liberia</collectingCountry>
, one bat with a full stomach was captured at 19:15 h. This speciesflies fast, without sudden turns, and with very fast wingbeats; it is not able to take off from ground, but climbs well. It roosts under bark of dead trees, in houses and roofs, and one was found roosting in a hollow in a baobab tree (
<taxonomicName box="[1579,1829,2027,2056]" class="Magnoliopsida" family="Malvaceae" genus="Adansonia" kingdom="Plantae" order="Malvales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="species" species="digitata">Adansonia digitata</taxonomicName>
,
<taxonomicName box="[1845,1997,2027,2056]" class="Magnoliopsida" family="Malvaceae" kingdom="Plantae" order="Malvales" pageId="59" pageNumber="817" phylum="Tracheophyta" rank="family">Malvaceae</taxonomicName>
). Predators include bat hawks (Macheiramphus alcinus).
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="biology_ecology">
<paragraph blockId="59.[1387,2600,725,2372]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[1389,2094,2101,2134]" pageId="59" pageNumber="817">Movements, Home range and Social organization.</emphasis>
In
<collectingCountry box="[2147,2242,2101,2134]" name="Gabon" pageId="59" pageNumber="817">Gabon</collectingCountry>
, two colonies numbered 9-12 individuals. Reported ectoparasites include the mites
<taxonomicName authorityName="Latreille" authorityYear="1796" box="[2226,2476,2141,2174]" class="Arachnida" family="Argasidae" genus="Carios" kingdom="Animalia" order="Ixodida" pageId="59" pageNumber="817" phylum="Arthropoda" rank="species" species="vespertilionis">Carios vespertilionis</taxonomicName>
(
<taxonomicName class="Arachnida" family="Argasidae" kingdom="Animalia" order="Ixodida" pageId="59" pageNumber="817" phylum="Arthropoda" rank="family">Argasidae</taxonomicName>
), Acanthophthirius mimetilli, Pteracarus mimetillus (both
<taxonomicName authorityName="Megnin" authorityYear="1877" box="[2225,2378,2184,2213]" class="Arachnida" family="Myobiidae" kingdom="Animalia" order="Trombidiformes" pageId="59" pageNumber="817" phylum="Arthropoda" rank="family">Myobiidae</taxonomicName>
), and Notoedres mimetilli (
<taxonomicName authorityName="A.Murray" authorityYear="1877" box="[1520,1695,2223,2252]" class="Arachnida" family="Sarcoptidae" kingdom="Animalia" order="Sarcoptiformes" pageId="59" pageNumber="817" phylum="Arthropoda" rank="family">Sarcoptidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection box="[1388,2439,2259,2292]" pageId="59" pageNumber="817" type="conservation">
<paragraph blockId="59.[1387,2600,725,2372]" box="[1388,2439,2259,2292]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[1388,1738,2259,2292]" pageId="59" pageNumber="817">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List.
</paragraph>
</subSubSection>
<subSubSection pageId="59" pageNumber="817" type="bibRefCitation_list">
<paragraph blockId="59.[1387,2600,725,2372]" pageId="59" pageNumber="817">
<emphasis bold="true" box="[1389,1542,2307,2332]" pageId="59" pageNumber="817">Bibliography.</emphasis>
Brosset (1966b), Cotterill (2001¢), Fahr (2013x), Gallagher &amp; Harrison (1977), Hill &amp; Harrison (1987), Kingdon (1974), Lang &amp; Chapin (1917a, 1917b).
</paragraph>
</subSubSection>
</treatment>
</document>