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<document ID-DOI="http://doi.org/10.5281/zenodo.6397752" ID-GBIF-Dataset="45351c32-25dd-422c-bdb2-00e73deb4943" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6397752" approvalRequired="37" approvalRequired_for_taxonomicNames="8" approvalRequired_for_treatments="29" checkinTime="1648655544658" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="4C3D87E8FF776AC9FA5F95C21FA5B96B" docLanguage="en" docName="hbmw_9_Vespertilionidae_716.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Kerivoula picta Pallas 1767" docType="treatment" docVersion="10" lastPageNumber="901" masterDocId="B004FF90FFFB6A44FFFC96591E00BB32" masterDocTitle="Vespertilionidae" masterLastPageNumber="981" masterPageNumber="716" pageNumber="900" updateTime="1658412944878" updateUser="diego">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Vespertilionidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Don E. Wilson</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Russell A. Mittermeier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>716</mods:start>
<mods:end>981</mods:end>
</mods:extent>
</mods:part>
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<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.6397752</mods:identifier>
<mods:identifier type="GBIF-Dataset">45351c32-25dd-422c-bdb2-00e73deb4943</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">6397752</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6576767" ID-GBIF-Taxon="195628339" ID-Zenodo-Dep="6576767" LSID="urn:lsid:plazi:treatment:4C3D87E8FF776AC9FA5F95C21FA5B96B" httpUri="http://treatment.plazi.org/id/4C3D87E8FF776AC9FA5F95C21FA5B96B" lastPageId="141" lastPageNumber="901" pageId="140" pageNumber="900">
<subSubSection box="[1443,1525,923,969]" pageId="140" pageNumber="900" type="multiple">
<paragraph blockId="140.[1441,2339,923,1048]" box="[1443,1525,923,969]" pageId="140" pageNumber="900">
<heading box="[1443,1525,923,969]" pageId="140" pageNumber="900">
<figureCitation box="[1443,1525,923,969]" captionStart="Plate 67: Vespertilionidae" captionStartId="135.[114,144,3260,3285]" captionTargetBox="[11,2751,19,3659]" captionTargetPageId="134" captionText="293. Groove-toothed Trumpet-eared Bat (Phoniscus atrox), 294. Peterss Trumpet-eared Bat (Phoniscus jagorii), 295. Dubious Trumpet-eared Bat (Phoniscus aerosus), 296. Golden-tipped Bat (Phonuscus papuensis), 297. Hardwickes Woolly Bat (Kerivoula hardwickii), 298. Flatskulled Woolly Bat (Kerivoula depressa), 299. Sri Lankan Woolly Bat (Kerivoula malpasi), 300. Cryptic Woolly Bat (Kerivoula crypta), 301. Dark Woolly Bat (Kerivoula furva), 302. Kachin Woolly Bat (Kerivoula kachinensis), 303. Indochinese Woolly Bat (Kerivoula dongduongana), 304. Krau Woolly Bat (Kerwoula krauensis), 305. Titanias Woolly Bat (Kerwoula titania), 306. Least Woolly Bat (Kerivoula minuta), 307. Small Woolly Bat (Kerivoula intermedia), 308. Papillose Woolly Bat (Kerivoula papillosa), 309. Lenis Woolly Bat (Kerivoula lenis), 310. Clear-winged Woolly Bat (Kerivoula pellucida), 311. Painted Woolly Bat (Kerwoula picta), 312. Whiteheads Woolly Bat (Kerivoula whiteheadi), 313. Flores Woolly Bat (Kerivoula flora), 314. Fly River Woolly Bat (Kerivoula muscina), 315. Saint Aignans Woolly Bat (Kerivoula agnella), 316. Bismarck Woolly Bat (Kerwoula myrella), 317. Lesser Woolly Bat (Kerivoula lanosa), 318. Ethiopian Woolly Bat (Kerwoula eriophora), 319. Tanzanian Woolly Bat (Kerwwoula africana), 320. Smiths Woolly Bat (Kerivoula smithii), 321. Coppery Woolly Bat (Kerivoula cuprosa), 322. Spurrells Woolly Bat (Kerivoula phalaena), 323. Damara Woolly Bat (Kerwoula argentata)" figureDoi="http://doi.org/10.5281/zenodo.6398545" httpUri="https://zenodo.org/record/6398545/files/figure.png" pageId="140" pageNumber="900">311.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1542,1938,923,969]" pageId="140" pageNumber="900" type="vernacular_names">
<paragraph blockId="140.[1441,2339,923,1048]" box="[1542,1938,923,969]" pageId="140" pageNumber="900">
<heading box="[1542,1938,923,969]" pageId="140" pageNumber="900">
<vernacularName box="[1542,1938,923,969]" pageId="140" pageNumber="900">Painted Woolly Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[2008,2275,923,969]" pageId="140" pageNumber="900" type="nomenclature">
<paragraph blockId="140.[1441,2339,923,1048]" box="[2008,2275,923,969]" pageId="140" pageNumber="900">
<heading box="[2008,2275,923,969]" pageId="140" pageNumber="900">
<taxonomicName authority="(Pallas, 1767)" authorityName="Pallas" authorityYear="1767" box="[2008,2275,923,969]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="140" pageNumber="900" phylum="Chordata" rank="species" species="picta">
<emphasis box="[2008,2275,923,969]" italics="true" pageId="140" pageNumber="900">Kerivoula picta</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="vernacular_names">
<paragraph blockId="140.[1441,2339,923,1048]" box="[1442,2338,986,1007]" pageId="140" pageNumber="900">
<heading box="[1442,2338,986,1007]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[1442,1518,986,1007]" pageId="140" pageNumber="900">French:</emphasis>
<vernacularName box="[1528,1687,986,1007]" pageId="140" pageNumber="900">Kérivoule peinte</vernacularName>
/
<emphasis bold="true" box="[1708,1799,986,1007]" pageId="140" pageNumber="900">German:</emphasis>
<vernacularName box="[1808,2029,986,1007]" pageId="140" pageNumber="900">Bunte Wollfledermaus</vernacularName>
/
<emphasis bold="true" box="[2049,2141,986,1007]" pageId="140" pageNumber="900">Spanish:</emphasis>
<vernacularName box="[2150,2256,986,1007]" pageId="140" pageNumber="900">Querivoula</vernacularName>
pintado
</heading>
</paragraph>
<paragraph blockId="140.[1441,2339,923,1048]" box="[1442,1811,1027,1048]" pageId="140" pageNumber="900">
<heading box="[1442,1811,1027,1048]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[1442,1689,1027,1048]" pageId="140" pageNumber="900">Other common names:</emphasis>
<vernacularName box="[1699,1811,1027,1048]" pageId="140" pageNumber="900">Painted Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="reference_group">
<paragraph blockId="140.[2054,2647,1090,1517]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[2054,2206,1090,1123]" pageId="140" pageNumber="900">Taxonomy.</emphasis>
<taxonomicName authorityName="Pallas" authorityYear="1767" box="[2230,2644,1090,1123]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="140" pageNumber="900" phylum="Chordata" rank="species" species="picta">Vespertilio pictum Pallas, 1767,</taxonomicName>
<materialsCitation ID-GBIF-Occurrence="3783319310" pageId="140" pageNumber="900">
“Ceylona &amp; e Molluccano Archipelago.” Restricted by G. H. H. Tate in 1941 to “Island of
<collectingCountry box="[2165,2273,1217,1242]" name="Indonesia" pageId="140" pageNumber="900">Ternate</collectingCountry>
, near Halmahera,” Molucca Islands,
<collectingCountry box="[2205,2343,1252,1281]" name="Indonesia" pageId="140" pageNumber="900">Indonesia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="discussion">
<paragraph blockId="140.[2054,2647,1090,1517]" pageId="140" pageNumber="900">
<taxonomicName baseAuthorityName="Pallas" baseAuthorityYear="1767" box="[2056,2246,1287,1320]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="140" pageNumber="900" phylum="Chordata" rank="species" species="picta">Kerivoula picta</taxonomicName>
and
<taxonomicName baseAuthorityName="Waterhouse" baseAuthorityYear="1845" box="[2315,2466,1287,1320]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="140" pageNumber="900" phylum="Chordata" rank="species" species="pellucida">K. pellucida</taxonomicName>
appear to be basal to the rest of the Asiatic
<taxonomicName pageId="140" pageNumber="900">Kerivoula species</taxonomicName>
, and both might be close to the African
<taxonomicName authorityName="Gray" authorityYear="1842" box="[2056,2179,1405,1438]" class="Mammalia" family="Vespertilionidae" genus="Kerivoula" kingdom="Animalia" order="Chiroptera" pageId="140" pageNumber="900" phylum="Chordata" rank="genus">Kerivoula</taxonomicName>
. Two subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.6398470" ID-Zenodo-Dep="6398470" httpUri="https://zenodo.org/record/6398470/files/figure.png" inLine="true" pageId="140" pageNumber="900" targetBox="[1440,2031,1101,1513]" targetPageId="140">
<paragraph blockId="140.[2054,2647,1090,1517]" lastBlockId="140.[1443,2657,1529,3492]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[2056,2456,1445,1478]" pageId="140" pageNumber="900">Subspecies and Distribution.</emphasis>
<taxonomicName authority="Pallas, 1767" authorityName="Pallas" authorityYear="1767" box="[2057,2367,1484,1517]" class="Magnoliopsida" family="Nepenthaceae" genus="Kerwoula" kingdom="Plantae" order="Caryophyllales" pageId="140" pageNumber="900" phylum="Tracheophyta" rank="subSpecies" species="picta" subSpecies="picta">K.p.pictaPallas,1767—India(Rajasthan,Gujarat,Maharashtra,Goa,Karnataka,AndhraPradesh,Kerala,TamilNadu,Sikkim,WestBengal,Odisha,andAssam),SriLanka,CNepal,Bangladesh,Myanmar,Thailand,Laos,Vietnam,Cambodia,PeninsularMalaysia,Sumatra,Java,BaliI,LombokI,andMoluccas(Halmahera,Ternate,andAmbonIs).</taxonomicName>
</paragraph>
<paragraph blockId="140.[1443,2657,1529,3492]" pageId="140" pageNumber="900">
<taxonomicName authority="Thomas, 1906" authorityName="Thomas" authorityYear="1906" box="[1446,1866,1683,1716]" class="Magnoliopsida" family="Nepenthaceae" genus="Kerwoula" kingdom="Plantae" order="Caryophyllales" pageId="140" pageNumber="900" phylum="Tracheophyta" rank="subSpecies" species="picta" subSpecies="bellissima">K. p. bellissima Thomas, 1906</taxonomicName>
— SE
<collectingCountry box="[1962,2047,1683,1716]" name="China" pageId="140" pageNumber="900">China</collectingCountry>
(
<collectingRegion box="[2074,2198,1683,1716]" country="China" name="Guizhou" pageId="140" pageNumber="900">Guizhou</collectingRegion>
,
<collectingRegion box="[2221,2342,1683,1716]" country="China" name="Guangxi" pageId="140" pageNumber="900">Guangxi</collectingRegion>
,
<collectingRegion box="[2364,2533,1683,1716]" country="China" name="Guangdong" pageId="140" pageNumber="900">Guangdong</collectingRegion>
, and
<collectingRegion country="China" name="Jiangxi" pageId="140" pageNumber="900">Jiangxi</collectingRegion>
), including
<collectingRegion box="[1698,1802,1725,1758]" country="China" name="Hainan" pageId="140" pageNumber="900">Hainan</collectingRegion>
I.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="description">
<paragraph blockId="140.[1443,2657,1529,3492]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[1445,1698,1770,1795]" pageId="140" pageNumber="900">Descriptive notes.</emphasis>
Head-body
<quantity box="[1890,2043,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="4.8" metricValueMin="4.0" pageId="140" pageNumber="900" unit="mm" value="44.0" valueMax="48.0" valueMin="40.0">40-48 mm</quantity>
, tail
<quantity box="[2121,2274,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="4.8" metricValueMin="3.7" pageId="140" pageNumber="900" unit="mm" value="42.5" valueMax="48.0" valueMin="37.0">37-48 mm</quantity>
, ear
<quantity box="[2354,2506,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="140" pageNumber="900" unit="mm" value="14.5" valueMax="16.0" valueMin="13.0">13-16 mm</quantity>
, hindfoot
<quantity box="[1446,1564,1801,1834]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="140" pageNumber="900" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">4-8 mm</quantity>
, forearm
<quantity box="[1710,1862,1801,1834]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.9" metricValueMin="3.1" pageId="140" pageNumber="900" unit="mm" value="35.0" valueMax="39.0" valueMin="31.0">31-39 mm</quantity>
; weight 4-5-5-
<quantity box="[2070,2119,1801,1834]" metricMagnitude="-3" metricUnit="kg" metricValue="5.0" pageId="140" pageNumber="900" unit="g" value="5.0">5 g</quantity>
. Fur is long, dense, and woolly. Dorsal pelage is bright orange to tawny red (hairs unicolored); venteris buffy, with distinct orange hue on flanks. Older males are apparently brighter than females. Wings are bright orange surrounding bones, with black between digits on membrane, and uropatagium is orangeto scarlet and hairy dorsally; there is well-defined fringe of hairs at posterior margin of uropatagium. Face is very hairy but lightly colored with hair on lips but not nostrils. Ears are relatively large, with smoothly concave anterior borders and distinct concavity just below rounded tips on posterior borders; tragusis tall and narrow, with pointed tip, slightly convex anterior margin, and basal lobe on posterior margin. Wings are attached at base of outertoes, and feet are hairy. Skull is mediumsized, rostrum is delicate and shallow, postorbital constriction is narrow, and braincase is inflated and elevated above rostrum. I? is conspicuously bicuspid.
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="biology_ecology">
<paragraph blockId="140.[1443,2657,1529,3492]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[1448,1559,2275,2308]" pageId="140" pageNumber="900">Habitat.</emphasis>
Primarily dry forests and scrubby areas but also agricultural areas (e.g. banana and plantain plantations, sugarcane, and other crop fields) from sea level up to elevations of c.
<quantity box="[1582,1691,2356,2389]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="140" pageNumber="900" unit="m" value="1500.0">1500 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="food_feeding">
<paragraph blockId="140.[1443,2657,1529,3492]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[1448,1709,2393,2426]" pageId="140" pageNumber="900">Food and Feeding.</emphasis>
Painted Woolly Bats forage low to the ground around and in bushy and densely vegetated areas. Flight is apparently fluttery, with up and down motion (resembling a large moth). They appearto feed mostly on web-building spiders, with insects making up small percentages of diets.
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="breeding">
<paragraph blockId="140.[1443,2657,1529,3492]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[1449,1584,2551,2584]" pageId="140" pageNumber="900">Breeding.</emphasis>
Painted Woolly Bats form monogamous breeding pairs each year; in
<collectingCountry name="Thailand" pageId="140" pageNumber="900">Thailand</collectingCountry>
, they appear to change partners annually. Two apparently mating individuals and a female carrying a young infant were captured in mid-October in
<collectingCountry box="[2380,2515,2630,2663]" name="Sri Lanka" pageId="140" pageNumber="900">Sri Lanka</collectingCountry>
. A female with a suckling infant and a male were captured under dry banana leaves in late August in
<collectingRegion box="[1550,1642,2709,2742]" country="India" name="Kerala" pageId="140" pageNumber="900">Kerala</collectingRegion>
, southern
<collectingCountry box="[1791,1866,2709,2742]" name="India" pageId="140" pageNumber="900">India</collectingCountry>
. The same family group system was noticed in February— October in
<collectingCountry box="[1614,1744,2748,2781]" name="Thailand" pageId="140" pageNumber="900">Thailand</collectingCountry>
. Oldest recorded age is five years for females and four for males in
<collectingCountry box="[1487,1616,2789,2822]" name="Thailand" pageId="140" pageNumber="900">Thailand</collectingCountry>
, but they likely live longer than this.
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="activity">
<paragraph blockId="140.[1443,2657,1529,3492]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[1449,1687,2827,2860]" pageId="140" pageNumber="900">Activity patterns.</emphasis>
Painted Woolly Bats roost in foliage and underlarge leaves, preferring dried, rolled up, and drooping leaves, particularly of plantains and bananas. They are also known to roost in dry grass, flowers, weaver nests, and other large-leaved plants. In
<collectingCountry box="[1592,1722,2945,2978]" name="Thailand" pageId="140" pageNumber="900">Thailand</collectingCountry>
, a pair was recorded roosting in man-made paper shelters. During the day, Painted Woolly Bats enter semi-torpor while roosting; they are very sluggish when disturbed. They emerge in the evening to forage through the night. In
<collectingCountry name="Thailand" pageId="140" pageNumber="900">Thailand</collectingCountry>
, calls of individuals in flight were steep FM sweep, with average start frequency of 140-2 kHz (98-7-156-9 kHz), end frequency of 103-6 kHz (81-134 kHz), peak frequency of 115-8 kHz (66-144-8 kHz), duration of 0-6 milliseconds (0-3-0-9 milliseconds), and interpulse interval of 41-1 milliseconds (10-2-592 milliseconds).
</paragraph>
</subSubSection>
<subSubSection pageId="140" pageNumber="900" type="biology_ecology">
<paragraph blockId="140.[1443,2657,1529,3492]" pageId="140" pageNumber="900">
<emphasis bold="true" box="[1452,2160,3222,3255]" pageId="140" pageNumber="900">Movements, Home range and Social organization.</emphasis>
Painted Woolly Bats often roost in breeding pairs with young during breeding season in
<collectingCountry box="[2243,2373,3261,3294]" name="Thailand" pageId="140" pageNumber="900">Thailand</collectingCountry>
, but they are often alone during non-breeding season (November—-December). In
<collectingCountry box="[2388,2518,3300,3333]" name="Thailand" pageId="140" pageNumber="900">Thailand</collectingCountry>
, roosting pairs were observed 64% of the time in November—-December, but in February, June, and September—October, they were always in family groups of a breeding pair and a single young. They switch roosts every 1-4 days. Foraging areas of breeding pairs did not overlap and were 6-1 ha for males and 5-6 ha for females.
</paragraph>
</subSubSection>
<subSubSection pageId="141" pageNumber="901" type="conservation">
<paragraph blockId="141.[140,1346,298,604]" pageId="141" pageNumber="901">
<emphasis bold="true" box="[143,491,298,327]" pageId="141" pageNumber="901">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. The Painted Woolly Batis widespread and does not face any major threats. Nevertheless,it is not particularly common anywhere, and its distribution is relatively fragmented.
</paragraph>
</subSubSection>
<subSubSection pageId="141" pageNumber="901" type="bibRefCitation_list">
<paragraph blockId="141.[140,1346,298,604]" pageId="141" pageNumber="901">
<emphasis bold="true" box="[143,295,420,445]" pageId="141" pageNumber="901">Bibliography.</emphasis>
Bates &amp; Harrison (1997), Corbet &amp; Hill (1992), Devkar et al. (2017), Flannery (1995a), Francis (2008a), Funakoshi,
<collectingRegion box="[364,418,459,484]" country="Japan" name="Fukui" pageId="141" pageNumber="901">Fukui</collectingRegion>
et al. (2015), Hawkeswood &amp; Sommung (2017), Hill (1965b), Hutson, Francis, Molur &amp; Srinivasulu (2008), Kruskop (2013a), Madhavan et al. (2014), Molur et al. (2002), Patel et al. (2017), Sarker &amp; Sarker (2005), Smith &amp; Xie Yan (2008), Srinivasulu &amp; Srinivasulu (2005, 2012), Sripathi et al. (2006), Tate (1941e), Tu Vuong Tan et al. (2018).
</paragraph>
</subSubSection>
</treatment>
</document>