354 lines
30 KiB
XML
354 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e83035" ID-Pensoft-Pub="1864-8312-80-439" ID-Pensoft-UUID="3F4E21B524465471831D09738C2304A6" ID-ZooBank="62FFB94CEBF441639F22881435EFC37C" ModsDocID="1864-8312-80-439" checkinTime="1663021027595" checkinUser="pensoft" docAuthor="Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha" docDate="2022" docId="A1D800DFD9DB51439BFCD88AF0C6744B" docLanguage="en" docName="ArthropodSystPhyl 80: 439-494" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-09-12" docSource="http://dx.doi.org/10.3897/asp.80.e83035" docTitle="Amphineurus (Rhamphoneurus) nullus Alexander 1967" docType="treatment" docVersion="1" id="3F4E21B524465471831D09738C2304A6" lastPageNumber="439" masterDocId="3F4E21B524465471831D09738C2304A6" masterDocTitle="Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="494" masterPageNumber="439" pageNumber="439" updateTime="1663021027595" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santos, Daubian</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1220-1267</mods:nameIdentifier>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">daubians@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
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<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
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||
</mods:name>
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||
<mods:name type="personal">
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||
<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>439</mods:start>
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<mods:end>494</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e83035</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e83035</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-439</mods:identifier>
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<mods:identifier type="ZooBank">62FFB94CEBF441639F22881435EFC37C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">3F4E21B524465471831D09738C2304A6</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:A1D800DFD9DB51439BFCD88AF0C6744B" httpUri="http://treatment.plazi.org/id/A1D800DFD9DB51439BFCD88AF0C6744B" lastPageNumber="439" pageId="0" pageNumber="439">
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<subSubSection pageId="0" pageNumber="439" type="nomenclature">
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<paragraph pageId="0" pageNumber="439">
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3.3.19.
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<taxonomicName LSID="A1D800DF-D9DB-5143-9BFC-D88AF0C6744B" authority="Alexander, 1967" authorityName="Alexander" authorityYear="1967" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nullus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nullus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nullus Alexander, 1967</taxonomicName>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="439" type="reference_group">
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<paragraph pageId="0" pageNumber="439">
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<taxonomicName authorityName="Alexander" authorityYear="1967" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nullus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nullus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nullus</taxonomicName>
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Alexander, 1967: 492 (description); Alexander, 1968: fig. 11 (wing), fig. 16 (male terminalia).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="439" type="materials_examined">
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||
<paragraph pageId="0" pageNumber="439">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="439">
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<emphasis bold="true" pageId="0" pageNumber="439">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<specimenCount>♂</specimenCount>
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, Chile,
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<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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Is., Aulen, Duhatao [
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<geoCoordinate degrees="41" direction="south" minutes="58" orientation="latitude" precision="925" value="-41.966667">41°58′S</geoCoordinate>
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<geoCoordinate degrees="74" direction="west" minutes="02" orientation="longitude" precision="925" value="-74.03333">74°02′W</geoCoordinate>
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], 6/
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<date value="1952-02-08">8-II-1952</date>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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,
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<typeStatus>Holotype</typeStatus>
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9620 (USNM) [part of specimen pinned and part in slide prepared by C.P. Alexander].
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<emphasis bold="true" pageId="0" pageNumber="439">
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<typeStatus>Allotype</typeStatus>
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</emphasis>
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:
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<specimenCount type="female">1 ♀</specimenCount>
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, Chile,
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<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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Is., Aulen, Duhatao [
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<geoCoordinate degrees="41" direction="south" minutes="58" orientation="latitude" precision="925" value="-41.966667">41°58′S</geoCoordinate>
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<geoCoordinate degrees="74" direction="west" minutes="02" orientation="longitude" precision="925" value="-74.03333">74°02′W</geoCoordinate>
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], 6/
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<date value="1952-02-08">8-II-1952</date>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM). -
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<emphasis bold="true" pageId="0" pageNumber="439">Additional material</emphasis>
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:
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<materialsCitation collectingDate="1968-04-12" collectorName="Osorno, Pena" country="CHILE" latitude="-40.583332" location="Bahia Mansa" longLatPrecision="1162" longitude="-73.73333" municipality="South of Pucotrihue" specimenCount="1" specimenCount-male="1">
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<emphasis bold="true" pageId="0" pageNumber="439">
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<collectingCountry name="Chile">CHILE</collectingCountry>
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.
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</emphasis>
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<specimenCount type="male">1 ♂</specimenCount>
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and 2 [sex unknown],
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<collectorName>Osorno</collectorName>
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,
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<location LSID="urn:lsid:plazi:treatment:A1D800DFD9DB51439BFCD88AF0C6744B:08A72B9345B0714CED9F1FDE17403870" country="CHILE" latitude="-40.583332" longLatPrecision="1162" longitude="-73.73333" municipality="South of Pucotrihue" name="Bahia Mansa">Bahia Mansa</location>
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[
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<geoCoordinate degrees="40" direction="south" minutes="35" orientation="latitude" precision="925" value="-40.583332">40°35′S</geoCoordinate>
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||
<geoCoordinate degrees="73" direction="west" minutes="44" orientation="longitude" precision="925" value="-73.73333">73°44′W</geoCoordinate>
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],
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<collectingMunicipality>South of Pucotrihue</collectingMunicipality>
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,
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<collectingDate value="1968-04-12">12-IV-1968</collectingDate>
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,
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<normalizedToken originalValue="Peña">
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<collectorName>Pena</collectorName>
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||
</normalizedToken>
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(USNM)
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</materialsCitation>
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;
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<specimenCount type="male">1 ♂</specimenCount>
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, Pucotrihue [
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||
<geoCoordinate degrees="40" direction="south" minutes="32" orientation="latitude" precision="925" value="-40.533333">40°32′S</geoCoordinate>
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||
<geoCoordinate degrees="73" direction="west" minutes="42" orientation="longitude" precision="925" value="-73.7">73°42′W</geoCoordinate>
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||
],
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<quantity metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" unit="m" value="35.0" valueMax="60.0" valueMin="10.0">10-60 m</quantity>
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,
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<date value="1955-03-12">12-III-1955</date>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM); 1 [sex unknown], Osorno, Pucotrihue [
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<geoCoordinate degrees="40" direction="south" minutes="32" orientation="latitude" precision="925" value="-40.533333">40°32′S</geoCoordinate>
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<geoCoordinate degrees="73" direction="west" minutes="42" orientation="longitude" precision="925" value="-73.7">73°42′W</geoCoordinate>
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], 10/
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||
<date value="1968-04-12">12-IV-1968</date>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM);
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<specimenCount type="male">1 ♂</specimenCount>
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||
,
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||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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||
Is., Aucar [
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||
<geoCoordinate degrees="42" direction="south" minutes="9" orientation="latitude" precision="925" value="-42.15">42°9′S</geoCoordinate>
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||
<geoCoordinate degrees="73" direction="west" minutes="30" orientation="longitude" precision="925" value="-73.5">73°30′W</geoCoordinate>
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],
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<date value="1952-01-15">15-I-1952</date>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM)*;
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<specimenCount type="male">1 ♂</specimenCount>
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,
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||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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||
Is., Aulen [
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||
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2′S</geoCoordinate>
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||
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1′W</geoCoordinate>
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],
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||
<date value="1952-02-08">8-II-1952</date>
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, L.P. Guzman (USNM)*;
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<specimenCount type="male">1 ♂</specimenCount>
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, Pucotrihue [
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||
<geoCoordinate degrees="40" direction="south" minutes="35" orientation="latitude" precision="925" value="-40.583332">40°35′S</geoCoordinate>
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||
<geoCoordinate degrees="73" direction="west" minutes="44" orientation="longitude" precision="925" value="-73.73333">73°44′W</geoCoordinate>
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||
],
|
||
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" unit="m" value="35.0" valueMax="60.0" valueMin="10.0">10-60 m</quantity>
|
||
,
|
||
<date value="1955-03-12">12-III-1955</date>
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||
,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
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||
(USNM)*; 1 [sex unknown],
|
||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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||
Is., Chepu [
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||
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2′S</geoCoordinate>
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||
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1′W</geoCoordinate>
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||
],
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||
<date value="1952-02-14">14-II-1952</date>
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||
,
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||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
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||
(USNM)*; 1 [sex unknown],
|
||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
|
||
Is., Chepu [
|
||
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2′S</geoCoordinate>
|
||
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1′W</geoCoordinate>
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||
], 4/
|
||
<date value="1952-04-06">6-IV-1952</date>
|
||
,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
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||
(USNM)*;
|
||
<materialsCitation collectingDate="1968-04-12" collectorName="Osorno, Pena" country="Chile" latitude="-40.583332" location="Bahia Mansa" longLatPrecision="1162" longitude="-73.73333" municipality="South of Pucotrihue" specimenCount="1" specimenCount-male="1">
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<specimenCount type="male">1 ♂</specimenCount>
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||
,
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<collectorName>Osorno</collectorName>
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||
,
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<location LSID="urn:lsid:plazi:treatment:A1D800DFD9DB51439BFCD88AF0C6744B:0D7CDE849D0D10F4438423E20F429312" country="Chile" latitude="-40.583332" longLatPrecision="1162" longitude="-73.73333" municipality="South of Pucotrihue" name="Bahia Mansa">Bahia Mansa</location>
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||
[
|
||
<geoCoordinate degrees="40" direction="south" minutes="35" orientation="latitude" precision="925" value="-40.583332">40°35′S</geoCoordinate>
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||
<geoCoordinate degrees="73" direction="west" minutes="44" orientation="longitude" precision="925" value="-73.73333">73°44′W</geoCoordinate>
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||
],
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||
<collectingMunicipality>South of Pucotrihue</collectingMunicipality>
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||
,
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||
<collectingDate value="1968-04-12">12-IV-1968</collectingDate>
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||
,
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||
<normalizedToken originalValue="Peña">
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||
<collectorName>Pena</collectorName>
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||
</normalizedToken>
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||
(USNM)
|
||
</materialsCitation>
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||
*.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="439" type="diagnosis">
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<paragraph pageId="0" pageNumber="439">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="439">The species is distinguished by absence of markings near R2, cross-vein m-cu attached to M3+4 and length of r-m twice length of basal deflection of R5. Furthermore, the species has distinctive male terminalia with a curved sheath of aedeagus, both mesal lobes with folded tips and branches of clasper of gonostylus of similar sizes. The species further distinguished also by the combination of sheath of aedeagus with small markings and male tergite IX pigmented. Furthermore, females of this species have the thickest cercus of the subgenus.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="439" type="redescription">
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<paragraph pageId="0" pageNumber="439">Redescription.</paragraph>
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<paragraph pageId="0" pageNumber="439">
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Wing length 6.32 mm, width 1.76 mm. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Coloration</emphasis>
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: General coloration brownish-yellow. Rostrum and palpus dark brown. First segment of antenna yellow, remainding segments dark yellow. Head grayish brown. Eyes black. Thorax brownish yellow, pleura brownish-yellow. Scutum without stripes. Halter yellow. Coxae and legs yellow. Abdomen brown. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Head</emphasis>
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(dorsal view Fig.
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Amphineurus (Rhamphoneurus) nullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439">24B</figureCitation>
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, ventral view Fig.
|
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Amphineurus (Rhamphoneurus) nullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439">24A</figureCitation>
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): Head wide, rostrum bifurcated with long appendices, first palpal segment longer than dilated terminal segment; scape as thick as pedicel, terminal flagellomere longer than penultimate. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Thorax</emphasis>
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(lateral view Fig.
|
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Amphineurus (Rhamphoneurus) nullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439">24E</figureCitation>
|
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, dorsal view Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Amphineurus (Rhamphoneurus) nullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439">24F</figureCitation>
|
||
): Anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Amphineurus (Rhamphoneurus) nullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439">24D</figureCitation>
|
||
) clear, without distinct markings; veins near fork of bM (Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Amphineurus (Rhamphoneurus) nullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439">24C</figureCitation>
|
||
) visible; R2 partially faded; crossvein r-m longer than basal deflection of R5; M1+2 about same length of M1+2; R2+3+4 longer than R2+3. -
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Female terminalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Amphineurus (Rhamphoneurus) nullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439">24H</figureCitation>
|
||
): Female tergite X longer than tergite IX, tergite IX wider. Cercus thick, longer than hypoginial valves, slightly curved dorsad. -
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Male terminalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Amphineurus (Rhamphoneurus) nullus Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439">24G</figureCitation>
|
||
): Male tergite IX darkened on posterior margin, with V-shaped median notch. Ventral branch of gonocoxite expanded in distal section. Dorsal branch of gonocoxite half-length of ventral branch. Lobe of gonostylus with round lobule; lobule longer than wide, longer than half length of lobe of gonostylus, wider than stem. Clasper of gonostylus with similarly shaped branches: lateral branch nearly oval: medial branch distally flattened, narrowed at margin. Mesal lobes of gonocoxite asymmetrical, both hook-shaped; left mesal lobe narrower. Sheath of aedeagus angulated, darkened only at tip.
|
||
</paragraph>
|
||
<caption doi="10.3897/asp.80.e83035.figure24" httpUri="https://binary.pensoft.net/fig/740577" pageId="0" pageNumber="439" start="Figure 24" startId="F24">
|
||
<paragraph pageId="0" pageNumber="439">
|
||
<emphasis bold="true" pageId="0" pageNumber="439">Figure 24.</emphasis>
|
||
<taxonomicName authorityName="Alexander" authorityYear="1967" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nullus" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nullus" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nullus</taxonomicName>
|
||
Alexander.
|
||
<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
|
||
head (ventral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
|
||
head (dorsal view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
|
||
detail of wing;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
|
||
wing;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">E</emphasis>
|
||
thorax (lateral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">F</emphasis>
|
||
thorax (dorsal view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">G</emphasis>
|
||
male tergite IX (dorsal view) and remainder of male terminalia (lateral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">H</emphasis>
|
||
female terminalia (dorsal view). - Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
|
||
, sheath of aedeagus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anatg</emphasis>
|
||
, anatergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anepm</emphasis>
|
||
, anepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anepst</emphasis>
|
||
, anepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">aprn</emphasis>
|
||
, anteropronotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cer</emphasis>
|
||
, cercus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
|
||
, clasper of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cx</emphasis>
|
||
, coxa;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
|
||
, gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">hlt</emphasis>
|
||
, halter;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">hyp val</emphasis>
|
||
, hypogynial valve;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">k</emphasis>
|
||
, knob of halter;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">kepm</emphasis>
|
||
, katepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">kepst</emphasis>
|
||
, katepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">ktg</emphasis>
|
||
, katatergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">l ms</emphasis>
|
||
, left mesal lobe of gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
|
||
; lobe of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mr</emphasis>
|
||
, meron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtanepst</emphasis>
|
||
, metanepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtepm</emphasis>
|
||
, metepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtg</emphasis>
|
||
, mediotergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtkepst</emphasis>
|
||
, metakatepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtn</emphasis>
|
||
, metanotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">p</emphasis>
|
||
, posterior basalare;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">patg</emphasis>
|
||
, paratergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">pprn</emphasis>
|
||
, postpronotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">presct</emphasis>
|
||
, prescutum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">r ms</emphasis>
|
||
, right mesal lobe of gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">sct</emphasis>
|
||
, scutum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">sctl</emphasis>
|
||
, scutellum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">st</emphasis>
|
||
, sternite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">tg</emphasis>
|
||
, tergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
|
||
, male tergite IX.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="439" type="remarks">
|
||
<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="439">
|
||
This species resembles
|
||
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) falcatus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="falcatus" subGenus="R.">A. (R.) falcatus</taxonomicName>
|
||
sp. nov. but differs mainly in the attachment of m-cu, shape of male tergite IX, prescutum and left mesal lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |