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<document id="28F31E60B72BF7FDEA8A597FDCADEE3F" ID-DOI="10.5281/zenodo.186233" ID-GBIF-Dataset="893ef34a-8765-4d78-821b-e196b9c0ce73" ID-ISSN="1175-5326" ID-Zenodo-Dep="186233" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461048143348" checkinUser="plazi" docAuthor="Dunz, Andreas R. &amp; Schliewen, Ulrich K." docDate="2009" docId="03EC8F79426AFFDD0397B840433B092A" docLanguage="en" docName="zt02028p019.pdf" docOrigin="Zootaxa 2028" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nannocharax zebra Dunz &amp; Schliewen, 2009, new species" docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="FFD5F701426EFFD40300BF7446390A36" masterDocTitle="Description of two new species of Nannocharax Günther, 1867 (Teleostei: Characiformes: Distichodontidae) from the Cross River, Cameroon" masterLastPageNumber="19" masterPageNumber="1" pageNumber="5" updateTime="1698577743192" updateUser="plazi">
<mods:mods id="FD5AD3466229B00E1FB84D958163740E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7926B616D90003C5F8E9EDF51B3E3C6E">
<mods:title id="BAD25850403AC705011F370DAC202F0F">Description of two new species of Nannocharax Günther, 1867 (Teleostei: Characiformes: Distichodontidae) from the Cross River, Cameroon</mods:title>
</mods:titleInfo>
<mods:name id="AE598FFBF0D73F5A89FAECF23128E7C0" type="personal">
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<mods:roleTerm id="870BAA0F87359E1725373E7CEB9C4A0B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D9D08A625C7F2F7439B2EF1ED392F2BD">Dunz, Andreas R.</mods:namePart>
</mods:name>
<mods:name id="5FD5FA264C9A0167974F9CEA522E99B8" type="personal">
<mods:role id="B9E00EBDD5D0EECE1D386B35BD264B4B">
<mods:roleTerm id="4B1AF89A94BBC78E955C05338D8FF195">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="99BCB18F9ABB08A52A16677D114C95B5">Schliewen, Ulrich K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="AEA66BD64ACBDFF5D145125F44AC326C">text</mods:typeOfResource>
<mods:relatedItem id="1CBDE96F28B21140BA6064AEC5DE52BE" type="host">
<mods:titleInfo id="CC4875C9CBD5CDFF345A5519AD13B75F">
<mods:title id="41B2D5B85406560C8F45BE4ED55E0608">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DA552440543343583524F865CBB71602">
<mods:date id="65423CB223C6B70C945388562ABDC572">2009</mods:date>
<mods:detail id="9129051EED113AD87D654481A2FE6345" type="volume">
<mods:number id="E3C8A61F9C2C0D5B45786B8450FBA1C0">2028</mods:number>
</mods:detail>
<mods:extent id="52997F7D0706BCD8554FD0E24B2AB613" unit="page">
<mods:start id="1F98732D44A567A7CCB371488399C17D">1</mods:start>
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<mods:classification id="F0C314C97666A5B62C72F061FCD8472F">journal article</mods:classification>
<mods:identifier id="5649DC2688DE8A23BCCC525AFF3B9A42" type="DOI">10.5281/zenodo.186233</mods:identifier>
<mods:identifier id="C346C15AC0973BF00BB30B129ED85BC5" type="GBIF-Dataset">893ef34a-8765-4d78-821b-e196b9c0ce73</mods:identifier>
<mods:identifier id="1DDC8543018E4A3A66B66C6AD9269005" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E8B64ADFEF1727E6F8C76FFF5482F647" type="Zenodo-Dep">186233</mods:identifier>
</mods:mods>
<treatment id="03EC8F79426AFFDD0397B840433B092A" ID-DOI="http://doi.org/10.5281/zenodo.5691649" ID-GBIF-Taxon="119611242" ID-Zenodo-Dep="5691649" LSID="urn:lsid:plazi:treatment:03EC8F79426AFFDD0397B840433B092A" httpUri="http://treatment.plazi.org/id/03EC8F79426AFFDD0397B840433B092A" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5">
<subSubSection id="C35F6DE4426AFFD00397B840478D0D47" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BFA3E6F426AFFD00397B840441C0D7B" blockId="4.[151,549,1843,1905]" box="[151,549,1843,1870]" pageId="4" pageNumber="5">
<heading id="D0B28903426AFFD00397B840441C0D7B" bold="true" box="[151,549,1843,1870]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B931E27D426AFFD00397B840441C0D7B" bold="true" box="[151,549,1843,1870]" pageId="4" pageNumber="5">
<taxonomicName id="4C4545EC426AFFD00397B84047BE0D78" ID-CoL="45MCR" box="[151,391,1844,1870]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="zebra" status="sp. nov.">
<emphasis id="B931E27D426AFFD00397B84047BE0D78" bold="true" box="[151,391,1844,1870]" italics="true" pageId="4" pageNumber="5">Nannocharax zebra</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2025F06426AFFD00295B847441C0D7B" box="[405,549,1843,1869]" pageId="4" pageNumber="5" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA3E6F426AFFD00397B823478D0D47" blockId="4.[151,549,1843,1905]" box="[151,436,1879,1905]" pageId="4" pageNumber="5">
(
<figureCitation id="137E22EA426AFFD0039FB82347380D47" box="[159,257,1879,1905]" captionStart="FIGURE 3" captionStartId="7.[151,260,1242,1266]" captionTargetBox="[151,1435,341,1241]" captionTargetId="figure@7.[151,1436,341,1286]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Nannocharax zebra (ZSM 35730), 29.1 mm SL, Cameroon, Cross River at Mamfé, a. Photograph of holotype showing preserved coloration in alcohol, b. Illustration of holotype, drawing by R. Kühbandner." httpUri="https://zenodo.org/record/186236/files/figure.png" pageId="4" pageNumber="5">Figure 3</figureCitation>
a &amp; b;
<tableCitation id="C6C70BD4426AFFD00253B82347930D47" box="[339,426,1879,1905]" captionStart="TABLE 3" captionStartId="8.[151,239,155,179]" captionText="TABLE 3. Measurements and counts for types of N. zebra sp. nov and N. zebra. MRAC 87 - 24 - P- 87." pageId="4" pageNumber="5">Table 3</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F6DE4426AFFD10397B8D745660B7A" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BFA3E6F426AFFD00397B8D747730DD2" blockId="4.[151,1437,1955,2020]" pageId="4" pageNumber="5">
<emphasis id="B931E27D426AFFD00397B8D747200D8B" bold="true" box="[151,281,1955,1981]" pageId="4" pageNumber="5">
<typeStatus id="54FE80CD426AFFD00397B8D7472D0D8B" box="[151,276,1955,1981]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED54A6AA426AFFD00222B8D047580D88" box="[290,353,1956,1982]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="5">ZSM</collectionCode>
35730 (
<quantity id="4CBD938A426AFFD002CBB8D044040D88" box="[459,573,1956,1982]" metricMagnitude="-2" metricUnit="m" metricValue="2.91" pageId="4" pageNumber="5" unit="mm" value="29.1">29.1 mm</quantity>
SL);
<collectingCountry id="F3527EFF426AFFD00187B8D045340D88" box="[647,781,1956,1982]" name="Cameroon" pageId="4" pageNumber="5">Cameroon</collectingCountry>
, Cross River at Mamfé, collected by U. Schliewen,
<date id="FFFB18AF426AFFD00397B8BE477F0DD2" box="[151,326,1994,2020]" pageId="4" pageNumber="5" value="2002-02">February 2002</date>
.
</paragraph>
<paragraph id="8BFA3E6F426BFFD103C5BFE345F20AC9" blockId="5.[151,1436,151,332]" pageId="5" pageNumber="6">
<emphasis id="B931E27D426BFFD103C5BFE347730A87" bold="true" box="[197,330,151,177]" pageId="5" pageNumber="6">
<typeStatus id="54FE80CD426BFFD103C5BFE3477C0A87" box="[197,325,151,177]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED54A6AA426BFFD10252BFEC47B70A84" box="[338,398,152,178]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="6">ZSM</collectionCode>
35731 (4,
<quantity id="4CBD938A426BFFD1010FBFEC44820A84" box="[527,699,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.25" metricValueMin="2.85" pageId="5" pageNumber="6" unit="mm" value="30.5" valueMax="32.5" valueMin="28.5">28.5–32.5 mm</quantity>
SL); collection data as for
<typeStatus id="54FE80CD426BFFD100FEBFEC42500A84" box="[1022,1129,152,178]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED54A6AA426BFFD10774BFEC42F20A84" box="[1140,1227,152,178]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="5" pageNumber="6">MRAC</collectionCode>
88-53-
<collectionCode id="ED54A6AA426BFFD1061EBFEC43030A84" box="[1310,1338,152,178]" pageId="5" pageNumber="6">P-</collectionCode>
129-153 (25, 25.0–
<quantity id="4CBD938A426BFFD10210BFCA47410AEE" box="[272,376,190,216]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" pageId="5" pageNumber="6" unit="mm" value="39.3">39.3 mm</quantity>
SL); Cross River basin, riffles in outlet of Badi River into Cross River and Cross River at Mamfé, (
<geoCoordinate id="EE7158A8426BFFD10203BF9147630AC9" box="[259,346,229,255]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="5.7666664">5°46’N</geoCoordinate>
<geoCoordinate id="EE7158A8426BFFD10262BF91478E0AC9" box="[354,439,229,255]" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="9.283334">9°17’E</geoCoordinate>
), collected by T.R. Roberts,
<date id="FFFB18AF426BFFD10006BF9145FF0AC9" box="[774,966,229,255]" pageId="5" pageNumber="6" value="1980-03-01" valueMax="1980-03-05" valueMin="1980-03-01">1-5 March 1980</date>
.
</paragraph>
<paragraph id="8BFA3E6F426BFFD103C5BE7F45660B7A" blockId="5.[151,1436,151,332]" pageId="5" pageNumber="6">
<emphasis id="B931E27D426BFFD103C5BE7F44080B13" bold="true" box="[197,561,267,293]" pageId="5" pageNumber="6">Additional non-type material</emphasis>
:
<collectionCode id="ED54A6AA426BFFD10140BE7844AF0B10" box="[576,662,268,294]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="5" pageNumber="6">MRAC</collectionCode>
87-24-
<collectionCode id="ED54A6AA426BFFD101EABE78453F0B10" box="[746,774,268,294]" pageId="5" pageNumber="6">P-</collectionCode>
87 (1,
<quantity id="4CBD938A426BFFD10050BE7845800B10" box="[848,953,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="4.07" pageId="5" pageNumber="6" unit="mm" value="40.7">40.7 mm</quantity>
SL);
<collectingCountry id="F3527EFF426BFFD100FABE78426C0B10" box="[1018,1109,268,294]" name="Nigeria" pageId="5" pageNumber="6">Nigeria</collectingCountry>
, Sombreiro River at Odieke (
<geoCoordinate id="EE7158A8426BFFD103A1BE4646CB0B7A" box="[161,242,306,332]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="5.133333">5°8’ N</geoCoordinate>
,
<geoCoordinate id="EE7158A8426BFFD103FEBE4647650B7A" box="[254,348,306,332]" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="6.6833334">6°41’ E</geoCoordinate>
), collected by C.B. Powell,
<date id="FFFB18AF426BFFD101A4BE4645620B7A" box="[676,859,306,332]" pageId="5" pageNumber="6" value="1986-01-07">7 January 1986</date>
.
</paragraph>
</subSubSection>
<paragraph id="8BFA3E6F426BFFD10397BE0A436B0C2B" pageId="5" pageNumber="6">
<table id="F945CCCF426B002B0397BE0A436B0C2B" box="[151,1362,382,1565]" gridcols="4" gridrows="28" pageId="5" pageNumber="6">
<tr id="35753C2D426B002B0397BE0A436B0BA0" box="[151,1362,382,406]" gridrow="0" pageId="5" pageNumber="6" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76A45551426B002B0397BE0A44FE0BA0" box="[151,711,382,406]" gridcol="0" gridrow="0" pageId="5" pageNumber="6">
<emphasis id="B931E27D426BFFD10397BE0A47310BA0" bold="true" box="[151,264,382,406]" pageId="5" pageNumber="6">TABLE 2.</emphasis>
Factor Loadings of PC I–III for Figure 1.
</th>
</tr>
<tr id="35753C2D426B002B0397BEC5436B0BFE" box="[151,1362,433,456]" gridrow="1" pageId="5" pageNumber="6" rowspan-0="1">
<td id="76A45551426B002B008EBEC545DC0BFE" box="[910,997,433,456]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">PC I</td>
<td id="76A45551426B002B073EBEC542AC0BFE" box="[1086,1173,433,456]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">PC II</td>
<td id="76A45551426B002B07EEBEC5436B0BFE" box="[1262,1362,433,456]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">PC III</td>
</tr>
<tr id="35753C2D426B002B0397BEA8436B0BC5" box="[151,1362,476,499]" gridrow="2" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BEA844FE0BC5" box="[151,711,476,499]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">Standard length (SL)</th>
<td id="76A45551426B002B008EBEA845DC0BC5" box="[910,997,476,499]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">0.1761</td>
<td id="76A45551426B002B073EBEA842AC0BC5" box="[1086,1173,476,499]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">0.06275</td>
<td id="76A45551426B002B07EEBEA8436B0BC5" box="[1262,1362,476,499]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">0.04536</td>
</tr>
<tr id="35753C2D426B002B0397BD72436B082B" box="[151,1362,518,541]" gridrow="3" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BD7244FE082B" box="[151,711,518,541]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">Head length</th>
<td id="76A45551426B002B008EBD7245DC082B" box="[910,997,518,541]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">0.144</td>
<td id="76A45551426B002B073EBD7242AC082B" box="[1086,1173,518,541]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">0.0622</td>
<td id="76A45551426B002B07EEBD72436B082B" box="[1262,1362,518,541]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">0.1991</td>
</tr>
<tr id="35753C2D426B002B0397BD45436B087E" box="[151,1362,561,584]" gridrow="4" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BD4544FE087E" box="[151,711,561,584]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">Head width</th>
<td id="76A45551426B002B008EBD4545DC087E" box="[910,997,561,584]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">0.1654</td>
<td id="76A45551426B002B073EBD4542AC087E" box="[1086,1173,561,584]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">0.1152</td>
<td id="76A45551426B002B07EEBD45436B087E" box="[1262,1362,561,584]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">-0.08629</td>
</tr>
<tr id="35753C2D426B002B0397BD28436B0845" box="[151,1362,604,627]" gridrow="5" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BD2844FE0845" box="[151,711,604,627]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">Interorbital width</th>
<td id="76A45551426B002B008EBD2845DC0845" box="[910,997,604,627]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">0.07548</td>
<td id="76A45551426B002B073EBD2842AC0845" box="[1086,1173,604,627]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">0.2325</td>
<td id="76A45551426B002B07EEBD28436B0845" box="[1262,1362,604,627]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">0.09509</td>
</tr>
<tr id="35753C2D426B002B0397BDF2436B08AB" box="[151,1362,646,669]" gridrow="6" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BDF244FE08AB" box="[151,711,646,669]" gridcol="0" gridrow="6" pageId="5" pageNumber="6">Eye diameter</th>
<td id="76A45551426B002B008EBDF245DC08AB" box="[910,997,646,669]" gridcol="1" gridrow="6" pageId="5" pageNumber="6">0.1127</td>
<td id="76A45551426B002B073EBDF242AC08AB" box="[1086,1173,646,669]" gridcol="2" gridrow="6" pageId="5" pageNumber="6">0.02721</td>
<td id="76A45551426B002B07EEBDF2436B08AB" box="[1262,1362,646,669]" gridcol="3" gridrow="6" pageId="5" pageNumber="6">0.3094</td>
</tr>
<tr id="35753C2D426B002B0397BDC5436B08FE" box="[151,1362,689,712]" gridrow="7" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BDC544FE08FE" box="[151,711,689,712]" gridcol="0" gridrow="7" pageId="5" pageNumber="6">Internostril distance</th>
<td id="76A45551426B002B008EBDC545DC08FE" box="[910,997,689,712]" gridcol="1" gridrow="7" pageId="5" pageNumber="6">0.09182</td>
<td id="76A45551426B002B073EBDC542AC08FE" box="[1086,1173,689,712]" gridcol="2" gridrow="7" pageId="5" pageNumber="6">0.2604</td>
<td id="76A45551426B002B07EEBDC5436B08FE" box="[1262,1362,689,712]" gridcol="3" gridrow="7" pageId="5" pageNumber="6">0.04636</td>
</tr>
<tr id="35753C2D426B002B0397BDA8436B08C5" box="[151,1362,732,755]" gridrow="8" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BDA844FE08C5" box="[151,711,732,755]" gridcol="0" gridrow="8" pageId="5" pageNumber="6">Minimal distance between nostril – eye</th>
<td id="76A45551426B002B008EBDA845DC08C5" box="[910,997,732,755]" gridcol="1" gridrow="8" pageId="5" pageNumber="6">0.164</td>
<td id="76A45551426B002B073EBDA842AC08C5" box="[1086,1173,732,755]" gridcol="2" gridrow="8" pageId="5" pageNumber="6">0.1313</td>
<td id="76A45551426B002B07EEBDA8436B08C5" box="[1262,1362,732,755]" gridcol="3" gridrow="8" pageId="5" pageNumber="6">0.03838</td>
</tr>
<tr id="35753C2D426B002B0397BC72436B092B" box="[151,1362,774,797]" gridrow="9" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BC7244FE092B" box="[151,711,774,797]" gridcol="0" gridrow="9" pageId="5" pageNumber="6">Upper jaw length</th>
<td id="76A45551426B002B008EBC7245DC092B" box="[910,997,774,797]" gridcol="1" gridrow="9" pageId="5" pageNumber="6">0.09636</td>
<td id="76A45551426B002B073EBC7242AC092B" box="[1086,1173,774,797]" gridcol="2" gridrow="9" pageId="5" pageNumber="6">0.1205</td>
<td id="76A45551426B002B07EEBC72436B092B" box="[1262,1362,774,797]" gridcol="3" gridrow="9" pageId="5" pageNumber="6">0.2139</td>
</tr>
<tr id="35753C2D426B002B0397BC45436B097E" box="[151,1362,817,840]" gridrow="10" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BC4544FE097E" box="[151,711,817,840]" gridcol="0" gridrow="10" pageId="5" pageNumber="6">Lower jaw width</th>
<td id="76A45551426B002B008EBC4545DC097E" box="[910,997,817,840]" gridcol="1" gridrow="10" pageId="5" pageNumber="6">0.1494</td>
<td id="76A45551426B002B073EBC4542AC097E" box="[1086,1173,817,840]" gridcol="2" gridrow="10" pageId="5" pageNumber="6">0.1007</td>
<td id="76A45551426B002B07EEBC45436B097E" box="[1262,1362,817,840]" gridcol="3" gridrow="10" pageId="5" pageNumber="6">0.2013</td>
</tr>
<tr id="35753C2D426B002B0397BC28436B0945" box="[151,1362,860,883]" gridrow="11" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BC2844FE0945" box="[151,711,860,883]" gridcol="0" gridrow="11" pageId="5" pageNumber="6">Snout length</th>
<td id="76A45551426B002B008EBC2845DC0945" box="[910,997,860,883]" gridcol="1" gridrow="11" pageId="5" pageNumber="6">0.1785</td>
<td id="76A45551426B002B073EBC2842AC0945" box="[1086,1173,860,883]" gridcol="2" gridrow="11" pageId="5" pageNumber="6">0.1138</td>
<td id="76A45551426B002B07EEBC28436B0945" box="[1262,1362,860,883]" gridcol="3" gridrow="11" pageId="5" pageNumber="6">0.008832</td>
</tr>
<tr id="35753C2D426B002B0397BCF2436B09AB" box="[151,1362,902,925]" gridrow="12" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BCF244FE09AB" box="[151,711,902,925]" gridcol="0" gridrow="12" pageId="5" pageNumber="6">Dorsal-fin length</th>
<td id="76A45551426B002B008EBCF245DC09AB" box="[910,997,902,925]" gridcol="1" gridrow="12" pageId="5" pageNumber="6">0.1533</td>
<td id="76A45551426B002B073EBCF242AC09AB" box="[1086,1173,902,925]" gridcol="2" gridrow="12" pageId="5" pageNumber="6">0.09759</td>
<td id="76A45551426B002B07EEBCF2436B09AB" box="[1262,1362,902,925]" gridcol="3" gridrow="12" pageId="5" pageNumber="6">-0.06089</td>
</tr>
<tr id="35753C2D426B002B0397BCC5436B09FE" box="[151,1362,945,968]" gridrow="13" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BCC544FE09FE" box="[151,711,945,968]" gridcol="0" gridrow="13" pageId="5" pageNumber="6">Length of dorsal-fin base</th>
<td id="76A45551426B002B008EBCC545DC09FE" box="[910,997,945,968]" gridcol="1" gridrow="13" pageId="5" pageNumber="6">0.1959</td>
<td id="76A45551426B002B073EBCC542AC09FE" box="[1086,1173,945,968]" gridcol="2" gridrow="13" pageId="5" pageNumber="6">0.04191</td>
<td id="76A45551426B002B07EEBCC5436B09FE" box="[1262,1362,945,968]" gridcol="3" gridrow="13" pageId="5" pageNumber="6">0.1553</td>
</tr>
<tr id="35753C2D426B002B0397BCA8436B09C5" box="[151,1362,988,1011]" gridrow="14" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BCA844FE09C5" box="[151,711,988,1011]" gridcol="0" gridrow="14" pageId="5" pageNumber="6">Length of anal-fin base</th>
<td id="76A45551426B002B008EBCA845DC09C5" box="[910,997,988,1011]" gridcol="1" gridrow="14" pageId="5" pageNumber="6">0.1328</td>
<td id="76A45551426B002B073EBCA842AC09C5" box="[1086,1173,988,1011]" gridcol="2" gridrow="14" pageId="5" pageNumber="6">0.1289</td>
<td id="76A45551426B002B07EEBCA8436B09C5" box="[1262,1362,988,1011]" gridcol="3" gridrow="14" pageId="5" pageNumber="6">-0.4077</td>
</tr>
<tr id="35753C2D426B002B0397BB72436B0E2B" box="[151,1362,1030,1053]" gridrow="15" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BB7244FE0E2B" box="[151,711,1030,1053]" gridcol="0" gridrow="15" pageId="5" pageNumber="6">Length of adipose-fin base</th>
<td id="76A45551426B002B008EBB7245DC0E2B" box="[910,997,1030,1053]" gridcol="1" gridrow="15" pageId="5" pageNumber="6">0.1047</td>
<td id="76A45551426B002B073EBB7242AC0E2B" box="[1086,1173,1030,1053]" gridcol="2" gridrow="15" pageId="5" pageNumber="6">0.1832</td>
<td id="76A45551426B002B07EEBB72436B0E2B" box="[1262,1362,1030,1053]" gridcol="3" gridrow="15" pageId="5" pageNumber="6">0.3006</td>
</tr>
<tr id="35753C2D426B002B0397BB45436B0E7E" box="[151,1362,1073,1096]" gridrow="16" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BB4544FE0E7E" box="[151,711,1073,1096]" gridcol="0" gridrow="16" pageId="5" pageNumber="6">Pelvic-fin length</th>
<td id="76A45551426B002B008EBB4545DC0E7E" box="[910,997,1073,1096]" gridcol="1" gridrow="16" pageId="5" pageNumber="6">0.1977</td>
<td id="76A45551426B002B073EBB4542AC0E7E" box="[1086,1173,1073,1096]" gridcol="2" gridrow="16" pageId="5" pageNumber="6">0.03908</td>
<td id="76A45551426B002B07EEBB45436B0E7E" box="[1262,1362,1073,1096]" gridcol="3" gridrow="16" pageId="5" pageNumber="6">-0.1809</td>
</tr>
<tr id="35753C2D426B002B0397BB28436B0E45" box="[151,1362,1116,1139]" gridrow="17" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BB2844FE0E45" box="[151,711,1116,1139]" gridcol="0" gridrow="17" pageId="5" pageNumber="6">Width of pelvic-fin base</th>
<td id="76A45551426B002B008EBB2845DC0E45" box="[910,997,1116,1139]" gridcol="1" gridrow="17" pageId="5" pageNumber="6">0.2461</td>
<td id="76A45551426B002B073EBB2842AC0E45" box="[1086,1173,1116,1139]" gridcol="2" gridrow="17" pageId="5" pageNumber="6">0.1174</td>
<td id="76A45551426B002B07EEBB28436B0E45" box="[1262,1362,1116,1139]" gridcol="3" gridrow="17" pageId="5" pageNumber="6">-0.2811</td>
</tr>
<tr id="35753C2D426B002B0397BBF2436B0EAB" box="[151,1362,1158,1181]" gridrow="18" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BBF244FE0EAB" box="[151,711,1158,1181]" gridcol="0" gridrow="18" pageId="5" pageNumber="6">Pectoral-fin length</th>
<td id="76A45551426B002B008EBBF245DC0EAB" box="[910,997,1158,1181]" gridcol="1" gridrow="18" pageId="5" pageNumber="6">0.1773</td>
<td id="76A45551426B002B073EBBF242AC0EAB" box="[1086,1173,1158,1181]" gridcol="2" gridrow="18" pageId="5" pageNumber="6">0.1364</td>
<td id="76A45551426B002B07EEBBF2436B0EAB" box="[1262,1362,1158,1181]" gridcol="3" gridrow="18" pageId="5" pageNumber="6">-0.2665</td>
</tr>
<tr id="35753C2D426B002B0397BBC5436B0EFE" box="[151,1362,1201,1224]" gridrow="19" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BBC544FE0EFE" box="[151,711,1201,1224]" gridcol="0" gridrow="19" pageId="5" pageNumber="6">Body depth</th>
<td id="76A45551426B002B008EBBC545DC0EFE" box="[910,997,1201,1224]" gridcol="1" gridrow="19" pageId="5" pageNumber="6">0.1588</td>
<td id="76A45551426B002B073EBBC542AC0EFE" box="[1086,1173,1201,1224]" gridcol="2" gridrow="19" pageId="5" pageNumber="6">0.1568</td>
<td id="76A45551426B002B07EEBBC5436B0EFE" box="[1262,1362,1201,1224]" gridcol="3" gridrow="19" pageId="5" pageNumber="6">0.1156</td>
</tr>
<tr id="35753C2D426B002B0397BBA8436B0EC5" box="[151,1362,1244,1267]" gridrow="20" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BBA844FE0EC5" box="[151,711,1244,1267]" gridcol="0" gridrow="20" pageId="5" pageNumber="6">Body width</th>
<td id="76A45551426B002B008EBBA845DC0EC5" box="[910,997,1244,1267]" gridcol="1" gridrow="20" pageId="5" pageNumber="6">0.1993</td>
<td id="76A45551426B002B073EBBA842AC0EC5" box="[1086,1173,1244,1267]" gridcol="2" gridrow="20" pageId="5" pageNumber="6">0.1626</td>
<td id="76A45551426B002B07EEBBA8436B0EC5" box="[1262,1362,1244,1267]" gridcol="3" gridrow="20" pageId="5" pageNumber="6">-0.2184</td>
</tr>
<tr id="35753C2D426B002B0397BA72436B0F2B" box="[151,1362,1286,1309]" gridrow="21" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BA7244FE0F2B" box="[151,711,1286,1309]" gridcol="0" gridrow="21" pageId="5" pageNumber="6">Caudal peduncle length</th>
<td id="76A45551426B002B008EBA7245DC0F2B" box="[910,997,1286,1309]" gridcol="1" gridrow="21" pageId="5" pageNumber="6">0.1487</td>
<td id="76A45551426B002B073EBA7242AC0F2B" box="[1086,1173,1286,1309]" gridcol="2" gridrow="21" pageId="5" pageNumber="6">0.1404</td>
<td id="76A45551426B002B07EEBA72436B0F2B" box="[1262,1362,1286,1309]" gridcol="3" gridrow="21" pageId="5" pageNumber="6">0.1541</td>
</tr>
<tr id="35753C2D426B002B0397BA45436B0F7E" box="[151,1362,1329,1352]" gridrow="22" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BA4544FE0F7E" box="[151,711,1329,1352]" gridcol="0" gridrow="22" pageId="5" pageNumber="6">Caudal peduncle depth</th>
<td id="76A45551426B002B008EBA4545DC0F7E" box="[910,997,1329,1352]" gridcol="1" gridrow="22" pageId="5" pageNumber="6">0.2029</td>
<td id="76A45551426B002B073EBA4542AC0F7E" box="[1086,1173,1329,1352]" gridcol="2" gridrow="22" pageId="5" pageNumber="6">0.05258</td>
<td id="76A45551426B002B07EEBA45436B0F7E" box="[1262,1362,1329,1352]" gridcol="3" gridrow="22" pageId="5" pageNumber="6">-0.3216</td>
</tr>
<tr id="35753C2D426B002B0397BA28436B0F45" box="[151,1362,1372,1395]" gridrow="23" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BA2844FE0F45" box="[151,711,1372,1395]" gridcol="0" gridrow="23" pageId="5" pageNumber="6">Body depth at anal fin end</th>
<td id="76A45551426B002B008EBA2845DC0F45" box="[910,997,1372,1395]" gridcol="1" gridrow="23" pageId="5" pageNumber="6">0.1689</td>
<td id="76A45551426B002B073EBA2842AC0F45" box="[1086,1173,1372,1395]" gridcol="2" gridrow="23" pageId="5" pageNumber="6">0.1405</td>
<td id="76A45551426B002B07EEBA28436B0F45" box="[1262,1362,1372,1395]" gridcol="3" gridrow="23" pageId="5" pageNumber="6">0.004146</td>
</tr>
<tr id="35753C2D426B002B0397BAF2436B0FAB" box="[151,1362,1414,1437]" gridrow="24" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BAF244FE0FAB" box="[151,711,1414,1437]" gridcol="0" gridrow="24" pageId="5" pageNumber="6">Preanal distance</th>
<td id="76A45551426B002B008EBAF245DC0FAB" box="[910,997,1414,1437]" gridcol="1" gridrow="24" pageId="5" pageNumber="6">0.1858</td>
<td id="76A45551426B002B073EBAF242AC0FAB" box="[1086,1173,1414,1437]" gridcol="2" gridrow="24" pageId="5" pageNumber="6">0.04641</td>
<td id="76A45551426B002B07EEBAF2436B0FAB" box="[1262,1362,1414,1437]" gridcol="3" gridrow="24" pageId="5" pageNumber="6">0.09498</td>
</tr>
<tr id="35753C2D426B002B0397BAC5436B0FFE" box="[151,1362,1457,1480]" gridrow="25" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BAC544FE0FFE" box="[151,711,1457,1480]" gridcol="0" gridrow="25" pageId="5" pageNumber="6">Prepelvic distance</th>
<td id="76A45551426B002B008EBAC545DC0FFE" box="[910,997,1457,1480]" gridcol="1" gridrow="25" pageId="5" pageNumber="6">0.1732</td>
<td id="76A45551426B002B073EBAC542AC0FFE" box="[1086,1173,1457,1480]" gridcol="2" gridrow="25" pageId="5" pageNumber="6">0.02382</td>
<td id="76A45551426B002B07EEBAC5436B0FFE" box="[1262,1362,1457,1480]" gridcol="3" gridrow="25" pageId="5" pageNumber="6">0.3008</td>
</tr>
<tr id="35753C2D426B002B0397BAA8436B0FC5" box="[151,1362,1500,1523]" gridrow="26" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397BAA844FE0FC5" box="[151,711,1500,1523]" gridcol="0" gridrow="26" pageId="5" pageNumber="6">Distance of anus to base of anal fin</th>
<td id="76A45551426B002B008EBAA845DC0FC5" box="[910,997,1500,1523]" gridcol="1" gridrow="26" pageId="5" pageNumber="6">0.5703</td>
<td id="76A45551426B002B073EBAA842AC0FC5" box="[1086,1173,1500,1523]" gridcol="2" gridrow="26" pageId="5" pageNumber="6">0.7751</td>
<td id="76A45551426B002B07EEBAA8436B0FC5" box="[1262,1362,1500,1523]" gridcol="3" gridrow="26" pageId="5" pageNumber="6">0.03347</td>
</tr>
<tr id="35753C2D426B002B0397B972436B0C2B" box="[151,1362,1542,1565]" gridrow="27" pageId="5" pageNumber="6">
<th id="76A45551426B002B0397B97244FE0C2B" box="[151,711,1542,1565]" gridcol="0" gridrow="27" pageId="5" pageNumber="6">Dorsal-adipose-distance</th>
<td id="76A45551426B002B008EB97245DC0C2B" box="[910,997,1542,1565]" gridcol="1" gridrow="27" pageId="5" pageNumber="6">0.1888</td>
<td id="76A45551426B002B073EB97242AC0C2B" box="[1086,1173,1542,1565]" gridcol="2" gridrow="27" pageId="5" pageNumber="6">0.09721</td>
<td id="76A45551426B002B07EEB972436B0C2B" box="[1262,1362,1542,1565]" gridcol="3" gridrow="27" pageId="5" pageNumber="6">0.001643</td>
</tr>
</table>
</paragraph>
<subSubSection id="C35F6DE4426BFFD103C5B91347FE0D5C" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BFA3E6F426BFFD103C5B91347FE0D5C" blockId="5.[151,1436,1639,2014]" pageId="5" pageNumber="6">
<emphasis id="B931E27D426BFFD103C5B913477C0CB7" bold="true" box="[197,325,1639,1665]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
<taxonomicName id="4C4545EC426BFFD1024DB91C44010CB7" box="[333,568,1640,1665]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D426BFFD1024DB91C44010CB7" box="[333,568,1640,1665]" italics="true" pageId="5" pageNumber="6">Nannocharax zebra</emphasis>
</taxonomicName>
is distinguished from all other described
<taxonomicName id="4C4545EC426BFFD10726B91C42FF0CB7" box="[1062,1222,1640,1665]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
<emphasis id="B931E27D426BFFD10726B91C42FF0CB7" box="[1062,1222,1640,1665]" italics="true" pageId="5" pageNumber="6">Nannocharax</emphasis>
</taxonomicName>
species except
<emphasis id="B931E27D426BFFD10681B91C44400C9E" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C4545EC426BFFD10681B91C471F0C9E" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="schoutedeni">N. schoutedeni</taxonomicName>
,
<taxonomicName id="4C4545EC426BFFD10233B9FB47D00C9E" box="[307,489,1679,1704]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="latifasciatus">N. latifasciatus</taxonomicName>
,
<taxonomicName id="4C4545EC426BFFD102F5B9FB44400C9E" box="[501,633,1679,1704]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ansorgii">N. ansorgii</taxonomicName>
</emphasis>
and
<emphasis id="B931E27D426BFFD101B3B9FB45590C9E" box="[691,864,1679,1704]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C4545EC426BFFD101B3B9FB45650C9E" box="[691,860,1679,1704]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="procatopus">N. procatopus</taxonomicName>
,
</emphasis>
by its shorter distance between posterior border of anus and articulation of first anal-fin ray (2.2–3.8 vs. 4.2–14.2% of SL). It differs from
<taxonomicName id="4C4545EC426BFFD107C7B9C1434F0CF8" box="[1223,1398,1717,1742]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="schoutedeni">
<emphasis id="B931E27D426BFFD107C7B9C1434F0CF8" box="[1223,1398,1717,1742]" italics="true" pageId="5" pageNumber="6">N. schoutedeni</emphasis>
</taxonomicName>
by having 39–43 vs. 36 or 37 lateral line scales, from
<taxonomicName id="4C4545EC426BFFD1000AB9A845FA0CC3" box="[778,963,1756,1781]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="latifasciatus">
<emphasis id="B931E27D426BFFD1000AB9A845FA0CC3" box="[778,963,1756,1781]" italics="true" pageId="5" pageNumber="6">N. latifasciatus</emphasis>
</taxonomicName>
by its major depth of caudal peduncle (55.9–63.8% vs. 48.1–50.8% of SL), from
<taxonomicName id="4C4545EC426BFFD10194B87745210D2A" box="[660,792,1795,1820]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ansorgii">
<emphasis id="B931E27D426BFFD10194B87745210D2A" box="[660,792,1795,1820]" italics="true" pageId="5" pageNumber="6">N. ansorgii</emphasis>
</taxonomicName>
by having 12–13 rather than 9 anal-fin rays, and from
<taxonomicName id="4C4545EC426BFFD10397B85D47050D74" box="[151,316,1833,1858]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="procatopus">
<emphasis id="B931E27D426BFFD10397B85D47050D74" box="[151,316,1833,1858]" italics="true" pageId="5" pageNumber="6">N. procatopus</emphasis>
</taxonomicName>
by having a lower caudal peduncle length (11.3–15.2% versus 18.0%), by having 39–43 rather than 36 lateral line scales.
</paragraph>
</subSubSection>
<subSubSection id="C35F6DE4426BFFDD03C5B80244EB0AEE" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6" type="description">
<paragraph id="8BFA3E6F426BFFD303C5B80242100B10" blockId="5.[151,1436,1639,2014]" lastBlockId="7.[151,1436,152,294]" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<emphasis id="B931E27D426BFFD103C5B80247670DA6" bold="true" box="[197,350,1910,1936]" pageId="5" pageNumber="6">Description.</emphasis>
Morphometric and meristic data of the
<typeStatus id="54FE80CD426BFFD10038B80245A60DA6" box="[824,927,1910,1936]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
and 29
<typeStatus id="54FE80CD426BFFD100FFB80242490DA6" box="[1023,1136,1910,1936]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
are presented in
<tableCitation id="C6C70BD4426BFFD1063DB80243AF0DA6" box="[1341,1430,1910,1936]" captionStart="TABLE 3" captionStartId="8.[151,239,155,179]" captionText="TABLE 3. Measurements and counts for types of N. zebra sp. nov and N. zebra. MRAC 87 - 24 - P- 87." pageId="5" pageNumber="6">Table 3</tableCitation>
. See
<figureCitation id="137E22EA426BFFD103CAB8E947030D81" box="[202,314,1949,1975]" captionStart="FIGURE 3" captionStartId="7.[151,260,1242,1266]" captionTargetBox="[151,1435,341,1241]" captionTargetId="figure@7.[151,1436,341,1286]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Nannocharax zebra (ZSM 35730), 29.1 mm SL, Cameroon, Cross River at Mamfé, a. Photograph of holotype showing preserved coloration in alcohol, b. Illustration of holotype, drawing by R. Kühbandner." httpUri="https://zenodo.org/record/186236/files/figure.png" pageId="5" pageNumber="6">Figures 3</figureCitation>
a &amp; b for general appearance.
<taxonomicName id="4C4545EC426BFFD101AAB8E945A10D80" box="[682,920,1949,1974]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D426BFFD101AAB8E945A10D80" box="[682,920,1949,1974]" italics="true" pageId="5" pageNumber="6">Nannocharax zebra</emphasis>
</taxonomicName>
is a medium-sized species (maximum size observed
<quantity id="4CBD938A426BFFD1020AB8B0474D0DE8" box="[266,372,1988,2014]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" pageId="5" pageNumber="6" unit="mm" value="39.3">39.3 mm</quantity>
SL). Body elongate, sides slightly compressed, greatest body depth at anterior dorsal-fin region. Predorsal body profile slightly convex. Body profile at base of dorsal fin slightly posteroventrally sloped. Dorsal body profile posterior to dorsal fin straight to caudal peduncle. Ventral body profile straight or very slightly posterodorsally sloped from tip of lower jaw to base of pectoral fin and from pectoral-fin insertion nearly straight to insertion of anal fin. Body profile at base of anal-fin base slightly posterodorsally inclined. Ventral body profile posterior to anal fin straight to caudale peduncle. Caudal peduncle length more than 10% of SL. Body depth at anal-fin end is located at the vertical distance between posterior base of anal fin and adipose fin. Head pointed in dorsal view. Eyes large and situated closer to the tip of the snout than to the posterior end of the operculum. Eye diameter always greater than interorbital width. Snout length equals interorbital width. Nostrils closer to anterior border of eye than to tip of snout. Mouth small and subterminal. Body covered with ctenoid scales. Lateral line complete and originating midlaterally behind operculum. Two
<emphasis id="B931E27D4268FFD203C5B814473B0D4C" bold="true" box="[197,258,1888,1914]" pageId="6" pageNumber="7">Fins.</emphasis>
Origin of dorsal fin on level with origin of pelvic fin. Length of dorsal-fin base short and first ray usually unbranched and very short. Dorsal rays
<quantity id="4CBD938A4268FFD201CAB8F3453F0D97" box="[714,774,1927,1953]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="6" pageNumber="7" unit="in" value="12.0">12 in</quantity>
<typeStatus id="54FE80CD4268FFD2000DB8F3454D0D97" box="[781,884,1927,1953]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
and
<quantity id="4CBD938A4268FFD200AEB8F3422F0D97" box="[942,1046,1927,1953]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="6" pageNumber="7" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">12–14 in</quantity>
<typeStatus id="54FE80CD4268FFD2071EB8F342AB0D97" box="[1054,1170,1927,1953]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
. Length of adipose-fin base very short and located approximately on the vertical through posterior end of anal-fin base. Caudal fin deeply forked with equal-sized lobes. Length of anal-fin base nearly equals length of dorsal-fin base. Anal rays
<quantity id="4CBD938A4269FFD303CEBFEC47300A84" box="[206,265,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="7" pageNumber="8" unit="in" value="12.0">12 in</quantity>
<typeStatus id="54FE80CD4269FFD30210BFEC474F0A84" box="[272,374,152,178]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
and 12 or
<quantity id="4CBD938A4269FFD302F1BFEC44150A84" box="[497,556,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="7" pageNumber="8" unit="in" value="13.0">13 in</quantity>
<typeStatus id="54FE80CD4269FFD30133BFEC449F0A84" box="[563,678,152,178]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
, sometimes first ray very small. Pelvic-fin rays
<quantity id="4CBD938A4269FFD307D2BFEC42C70A84" box="[1234,1278,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="7" pageNumber="8" unit="in" value="8.0">8 in</quantity>
<typeStatus id="54FE80CD4269FFD30604BFEC43520A84" box="[1284,1387,152,178]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
and
<quantity id="4CBD938A4269FFD30397BFCA46DA0AEE" box="[151,227,190,216]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" pageId="7" pageNumber="8" unit="in" value="8.0" valueMax="9.0" valueMin="7.0">7–9 in</quantity>
<typeStatus id="54FE80CD4269FFD303ECBFCA475B0AEE" box="[236,354,190,216]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
, third pelvic ray always longest. First, second and third ray slightly thicker than all other pelvic rays. Tip of longest pelvic-fin ray not reaching anus. Pectoral-fin rays
<quantity id="4CBD938A4269FFD30744BF9142BB0AC9" box="[1092,1154,229,255]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="7" pageNumber="8" unit="in" value="13.0">13 in</quantity>
<typeStatus id="54FE80CD4269FFD3078BBF9142CC0AC9" box="[1163,1269,229,255]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
and
<quantity id="4CBD938A4269FFD30631BF9143A50AC9" box="[1329,1436,229,255]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="3.556" metricValueMin="3.048" pageId="7" pageNumber="8" unit="in" value="13.0" valueMax="14.0" valueMin="12.0">12–14 in</quantity>
<typeStatus id="54FE80CD4269FFD30397BE7847330B10" box="[151,266,268,294]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
, tip of longest pectoral-fin ray does not reach insertion of pelvic fin.
</paragraph>
<caption id="DF3A6EE74268FFD20397B95042460D18" httpUri="https://zenodo.org/record/186235/files/figure.png" pageId="6" pageNumber="7" targetBox="[151,1379,528,1510]" targetPageId="6">
<paragraph id="8BFA3E6F4268FFD20397B95042460D18" blockId="6.[151,1436,1572,1838]" pageId="6" pageNumber="7">
<emphasis id="B931E27D4268FFD20397B95047190C0A" bold="true" box="[151,288,1572,1596]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
Plot of sheared PC II morphometric data vs. PC I meristic data shown as 95% confidence interval as ellipses. Red (cross) =
<emphasis id="B931E27D4268FFD20294B93347C10C68" box="[404,504,1607,1630]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C4545EC4268FFD20294B93347CD0C68" box="[404,500,1607,1630]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="zebra">N. zebra</taxonomicName>
,
</emphasis>
pink (filled square) =
<emphasis id="B931E27D4268FFD201F3B933455B0C68" box="[755,866,1607,1630]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="4C4545EC4268FFD201F3B93345670C68" box="[755,862,1607,1630]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="usongo">N usongo</taxonomicName>
,
</emphasis>
blue (open square) = remainig
<taxonomicName id="4C4545EC4268FFD207BEB933436B0C68" box="[1214,1362,1607,1630]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B931E27D4268FFD207BEB933436B0C68" box="[1214,1362,1607,1630]" italics="true" pageId="6" pageNumber="7">Nannocharax</emphasis>
</taxonomicName>
, black (dots) = five specimens, which clustering close with
<taxonomicName id="4C4545EC4268FFD201E0B91E45070CB7" box="[736,830,1642,1665]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4268FFD201E0B91E45070CB7" box="[736,830,1642,1665]" italics="true" pageId="6" pageNumber="7">N. zebra</emphasis>
</taxonomicName>
: 1=
<taxonomicName id="4C4545EC4268FFD20072B91E45F60CB7" box="[882,975,1642,1665]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4268FFD20072B91E45F60CB7" box="[882,975,1642,1665]" italics="true" pageId="6" pageNumber="7">N. zebra</emphasis>
</taxonomicName>
(MRAC 87-24-P-87), 2 =
<taxonomicName id="4C4545EC4268FFD207F8B91E43A50CB7" box="[1272,1436,1642,1665]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="schoutedeni">
<emphasis id="B931E27D4268FFD207F8B91E43A50CB7" box="[1272,1436,1642,1665]" italics="true" pageId="6" pageNumber="7">N. schoutedeni</emphasis>
</taxonomicName>
(ZSM 18926), 3 =
<taxonomicName id="4C4545EC4268FFD20262B9F9443B0C92" box="[354,514,1677,1700]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="schoutedeni">
<emphasis id="B931E27D4268FFD20262B9F9443B0C92" box="[354,514,1677,1700]" italics="true" pageId="6" pageNumber="7">N. schoutedeni</emphasis>
</taxonomicName>
(MRAC P-39705), 4 =
<taxonomicName id="4C4545EC4268FFD20003B9F945FD0C92" box="[771,964,1676,1700]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="schoutedeni">
<emphasis id="B931E27D4268FFD20003B9F945250C92" box="[771,796,1677,1700]" italics="true" pageId="6" pageNumber="7">N.</emphasis>
cf.
<emphasis id="B931E27D4268FFD20045B9F945FD0C92" box="[837,964,1677,1700]" italics="true" pageId="6" pageNumber="7">schoutedeni</emphasis>
</taxonomicName>
(ZSM 35332) and 5 =
<taxonomicName id="4C4545EC4268FFD207BEB9F9436C0C92" box="[1214,1365,1677,1700]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="procatopus">
<emphasis id="B931E27D4268FFD207BEB9F9436C0C92" box="[1214,1365,1677,1700]" italics="true" pageId="6" pageNumber="7">N. procatopus</emphasis>
</taxonomicName>
(ZSM 30030); 1 red (cross) =
<taxonomicName id="4C4545EC4268FFD20294B9DB47C90CF0" box="[404,496,1711,1734]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4268FFD20294B9DB47C90CF0" box="[404,496,1711,1734]" italics="true" pageId="6" pageNumber="7">N. zebra</emphasis>
</taxonomicName>
(MRAC 88-53-P-129-153) (River Badi), 2 red (cross) =
<taxonomicName id="4C4545EC4268FFD20755B9DB42960CF0" box="[1109,1199,1711,1734]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4268FFD20755B9DB42960CF0" box="[1109,1199,1711,1734]" italics="true" pageId="6" pageNumber="7">N. zebra</emphasis>
</taxonomicName>
(ZSM 35731); 1 pink (filled square) =
<taxonomicName id="4C4545EC4268FFD20248B9A6478D0CDF" box="[328,436,1746,1769]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="usongo">
<emphasis id="B931E27D4268FFD20248B9A6478D0CDF" box="[328,436,1746,1769]" italics="true" pageId="6" pageNumber="7">N. usongo</emphasis>
</taxonomicName>
(MRAC 88-53-P-116-128) (River Badi); 1 blue (open square) =
<taxonomicName id="4C4545EC4268FFD20766B9A642FE0CDF" box="[1126,1223,1746,1769]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="taenia">
<emphasis id="B931E27D4268FFD20766B9A642FE0CDF" box="[1126,1223,1746,1769]" italics="true" pageId="6" pageNumber="7">N. taenia</emphasis>
</taxonomicName>
(MRAC P-1283), 2 blue (open square) =
<taxonomicName id="4C4545EC4268FFD20284B98144360D3A" box="[388,527,1781,1804]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gobioides">
<emphasis id="B931E27D4268FFD20284B98144360D3A" box="[388,527,1781,1804]" italics="true" pageId="6" pageNumber="7">N. gobioides</emphasis>
</taxonomicName>
(MRAC 141442), 3 blue (open square) =
<taxonomicName id="4C4545EC4268FFD200E4B981425D0D3A" box="[996,1124,1781,1804]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="ogoensis">
<emphasis id="B931E27D4268FFD200E4B981425D0D3A" box="[996,1124,1781,1804]" italics="true" pageId="6" pageNumber="7">N. ogoensis</emphasis>
</taxonomicName>
(ZSM 27101), 4 blue (open square) =
<taxonomicName id="4C4545EC4268FFD20202B86347B90D18" box="[258,384,1815,1838]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="B931E27D4268FFD20202B86347B90D18" box="[258,384,1815,1838]" italics="true" pageId="6" pageNumber="7">N. fasciatus</emphasis>
</taxonomicName>
(ZSM 27426) and 5 blue (open square) =
<taxonomicName id="4C4545EC4268FFD20043B86345D90D18" box="[835,992,1815,1838]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B931E27D4268FFD20043B86345D90D18" box="[835,992,1815,1838]" italics="true" pageId="6" pageNumber="7">N. intermedius</emphasis>
</taxonomicName>
(ZSM 30033).
</paragraph>
</caption>
<caption id="DF3A6EE74269FFD30397BBAE43A30F22" httpUri="https://zenodo.org/record/186236/files/figure.png" pageId="7" pageNumber="8" targetBox="[151,1435,341,1241]" targetPageId="7">
<paragraph id="8BFA3E6F4269FFD30397BBAE43A30F22" blockId="7.[151,1437,1242,1301]" pageId="7" pageNumber="8">
<emphasis id="B931E27D4269FFD30397BBAE471B0EC4" bold="true" box="[151,290,1242,1266]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
Holotype of
<taxonomicName id="4C4545EC4269FFD302BCBBAF44A60EC4" box="[444,671,1243,1266]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4269FFD302BCBBAF44A60EC4" box="[444,671,1243,1266]" italics="true" pageId="7" pageNumber="8">Nannocharax zebra</emphasis>
</taxonomicName>
(ZSM 35730), 29.1 mm SL, Cameroon, Cross River at Mamfé,
<emphasis id="B931E27D4269FFD30687BBAE43AD0EC4" bold="true" box="[1415,1428,1242,1266]" pageId="7" pageNumber="8">a</emphasis>
. Photograph of holotype showing preserved coloration in alcohol,
<emphasis id="B931E27D4269FFD30052BB8945500F23" bold="true" box="[850,873,1277,1301]" pageId="7" pageNumber="8">b.</emphasis>
Illustration of holotype, drawing by R. Kühbandner.
</paragraph>
</caption>
<paragraph id="8BFA3E6F4269FFD303C5BA3644BB0FB5" blockId="7.[151,1437,1346,2030]" pageId="7" pageNumber="8">
<emphasis id="B931E27D4269FFD303C5BA36477A0F6A" bold="true" box="[197,323,1346,1372]" pageId="7" pageNumber="8">Dentition.</emphasis>
Upper jaw with a single row of 12 bicuspid, acute teeth in
<typeStatus id="54FE80CD4269FFD3070DBA37424C0F6B" box="[1037,1141,1347,1373]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
and
<quantity id="4CBD938A4269FFD307B0BA3743230F6B" box="[1200,1306,1347,1373]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="4.064" metricValueMin="2.54" pageId="7" pageNumber="8" unit="in" value="13.0" valueMax="16.0" valueMin="10.0">10–16 in</quantity>
<typeStatus id="54FE80CD4269FFD30622BA3743A10F6B" box="[1314,1432,1347,1373]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
. Upper jaw teeth limited to the premaxilla.
</paragraph>
<paragraph id="8BFA3E6F4269FFD303C5BAE447ED0DD8" blockId="7.[151,1437,1346,2030]" pageId="7" pageNumber="8">
<emphasis id="B931E27D4269FFD303C5BAE447ED0F9C" bold="true" box="[197,468,1424,1450]" pageId="7" pageNumber="8">Coloration in alcohol.</emphasis>
Ground coloration pale yellow, with few scattered dark melanophores in
<typeStatus id="54FE80CD4269FFD30635BAE443A50F9C" box="[1333,1436,1424,1450]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
and
<typeStatus id="54FE80CD4269FFD303C8BAC347010FE7" box="[200,312,1463,1489]" pageId="7" pageNumber="8" type="paratype">paratypes</typeStatus>
(
<figureCitation id="137E22EA4269FFD30247BAC347B50FE7" box="[327,396,1463,1489]" captionStart="FIGURE 3" captionStartId="7.[151,260,1242,1266]" captionTargetBox="[151,1435,341,1241]" captionTargetId="figure@7.[151,1436,341,1286]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Nannocharax zebra (ZSM 35730), 29.1 mm SL, Cameroon, Cross River at Mamfé, a. Photograph of holotype showing preserved coloration in alcohol, b. Illustration of holotype, drawing by R. Kühbandner." httpUri="https://zenodo.org/record/186236/files/figure.png" pageId="7" pageNumber="8">Fig. 3</figureCitation>
a). Melanophore concentration higher on the dorsal region. Melanophores located mainly on borders of scales, numerous on upper lip, fewer on lower lip. Region around nostrils unpigmented. A thin line of melanophores extending middorsally from tip of snout to interorbital area. Another thin horizontal line of melanophores between tip of snout and eye. Operculum ventrally unpigmented and dorsally slightly pigmented, similar to infraorbitals. Dorsal surface of head strongly pigmented at nape. Ventral surface of head ranging from unpigmented to having scattered, small, dark melanophores. Body covered with 9–12 vertical, dark bars largely under lateral line and 12–15 small, dark blotches along dorsum. Usually wide bars in the predorsal region not ventrally connected. Lateral flank bars in caudal region ventrally connected with each other. Small saddle blotches on dorsum may be faintly connected with flank bars. Caudal peduncle with a large blotch, extending onto caudal fin. Dorsal fin coloration ranging from unpigmented to 1–4 small spots on the first rays. Some young specimens with completely hyaline fins. Adipose fin without coloration. Caudal fin with narrow dark band in each lobe a basally situated, both bands are at the same level, but are not connected to each other. First two rays of anal fin coloured in center with a small spot. Pelvic fin with a dark band in the center, well visible on first three rays. Pectoral fin almost without coloration. Anterior region of the pectoralfin base with a small mark.
</paragraph>
<caption id="DF3A6EE74266FFDC0397BFEF42F40A84" box="[151,1229,155,179]" pageId="8" pageNumber="9">
<paragraph id="8BFA3E6F4266FFDC0397BFEF42F40A84" blockId="8.[151,1429,155,1547]" box="[151,1229,155,179]" pageId="8" pageNumber="9">
<emphasis id="B931E27D4266FFDC0397BFEF47310A85" bold="true" box="[151,264,155,179]" pageId="8" pageNumber="9">TABLE 3.</emphasis>
Measurements and counts for types of
<taxonomicName id="4C4545EC4266FFDC01ADBFEF45310A84" box="[685,776,155,178]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="zebra" status="sp. nov.">
<emphasis id="B931E27D4266FFDC01ADBFEF45310A84" box="[685,776,155,178]" italics="true" pageId="8" pageNumber="9">N. zebra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2025F064266FFDC000EBFEF45620A84" box="[782,859,155,178]" pageId="8" pageNumber="9" rank="species">sp. nov</taxonomicNameLabel>
and
<emphasis id="B931E27D4266FFDC008FBFEF45C80A84" box="[911,1009,155,178]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C4545EC4266FFDC008FBFEF45D50A84" box="[911,1004,155,178]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="zebra">N. zebra</taxonomicName>
.
</emphasis>
MRAC 87-24-P-87.
</paragraph>
</caption>
<paragraph id="8BFA3E6F4266FFDC0150BFB947EA0BB4" blockId="8.[151,1429,155,1547]" pageId="8" pageNumber="9">
<emphasis id="B931E27D4266FFDC0150BFB943BA0B3C" bold="true" pageId="8" pageNumber="9">
<typeStatus id="54FE80CD4266FFDC0150BFB945870AD4" box="[592,958,205,226]" pageId="8" pageNumber="9" type="holotype">holotype holotype</typeStatus>
+
<typeStatus id="54FE80CD4266FFDC00D8BFB942040AD4" box="[984,1085,205,226]" pageId="8" pageNumber="9" type="paratype">paratypes</typeStatus>
MRAC 87-24-P-87 Measurements min max mean SD n
</emphasis>
Total length (mm) 43.5 31.5 48.3 48.3 31 Standard length (SL) (mm) 29.1 25.0 40.7 40.7 31
<emphasis id="B931E27D4266FFDC039FBE1947EA0BB4" bold="true" box="[159,467,365,386]" pageId="8" pageNumber="9">In percents of standard length</emphasis>
</paragraph>
<paragraph id="8BFA3E6F4266FFDC039FBEE2479009EC" blockId="8.[151,1429,155,1547]" pageId="8" pageNumber="9">
Head length 27.2 25.4 28.3 27.1 0.7 26.7 31 Dorsal-fin length 23.9 18.9 25.7 24.3 1.3 22.6 31 Length of dorsal-fin base 13.5 12.2 14.7 13.2 0.6 12.7 31 Length of anal-fin base 13.3 12.2 13.9 13.1 0.4 11.6 31 Length of adipose-fin base 2.1 1.5 3.1 2.2 0.4 1.8 31 Pelvic-fin length 20.6 19.0 22.0 20.6 0.7 18.8 31 Pectoral-fin length 18.3 15.2 20.5 19.0 1.2 17.9 31 Body depth 18.6 17.0 19.4 17.9 0.6 16.9 31 Body width 10.0 9.6 11.1 10.4 0.3 10.3 31 Caudal peduncle length 13.6 11.3 15.2 13.1 0.8 13.7 31 Preanal distance 72.9 68.7 72.9 70.7 1.1 72.5 31 Prepelvic distance 46.6 42.5 46.6 44.5 0.9 46.1 31 Distance of anus to base of anal fin 3.3 2.2 3.8 2.8 0.4 3.1 31 Dorsal-adipose-distance 25.9 23.0 26.2 24.2 0.8 26.2 31
<emphasis id="B931E27D4266FFDC039FBCB1479009EC" bold="true" box="[159,425,965,986]" pageId="8" pageNumber="9">In percents of head length</emphasis>
</paragraph>
<paragraph id="8BFA3E6F4266FFDC039FBC9A479D0EC4" blockId="8.[151,1429,155,1547]" pageId="8" pageNumber="9">
Interorbital width 28.9 25.5 29.5 27.9 0.9 23.3 31 Eye diameter 33.1 29.9 34.6 32.6 1.1 28.6 31 Internostril distance 20.3 16.5 20.3 18.2 1.0 16.2 31 Minimal distance between nostril - eye 4.3 4.3 6.9 5.8 0.6 4.5 31 Snout length 26.4 25.1 29.7 27.0 0.9 29.7 31 Upper jaw length 20.6 17.7 23.3 21.4 1.2 16.1 31
<emphasis id="B931E27D4266FFDC039FBBA9479D0EC4" bold="true" box="[159,420,1245,1266]" pageId="8" pageNumber="9">In percents of head width</emphasis>
</paragraph>
<paragraph id="8BFA3E6F4266FFDC039FBA72479C0F74" blockId="8.[151,1429,155,1547]" pageId="8" pageNumber="9">
Lower jaw width 36.6 31.7 40.9 36.4 1.9 36.1 31
<emphasis id="B931E27D4266FFDC039FBA59479C0F74" bold="true" box="[159,421,1325,1346]" pageId="8" pageNumber="9">In percents of body width</emphasis>
</paragraph>
<paragraph id="8BFA3E6F4266FFDC039FBA22479C0F8D" blockId="8.[151,1429,155,1547]" pageId="8" pageNumber="9">
Head width 124.7 111.5 128.7 121.9 4.1 121.9 31 Width of pelvic-fin base 63.6 40.9 63.6 52.6 5.9 52.6 31
<emphasis id="B931E27D4266FFDC039FBAD2479C0F8D" bold="true" box="[159,421,1446,1467]" pageId="8" pageNumber="9">In percents of body depth</emphasis>
</paragraph>
<paragraph id="8BFA3E6F4266FFDD039FBABA44EB0AEE" blockId="8.[151,1429,155,1547]" lastBlockId="9.[151,1437,151,796]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Caudal peduncle depth 63.7 55.9 64.5 63.3 1.9 63.3 31 Body depth at anal-fin end 68.8 57.2 71.1 72.4 3.3 72.4 31
<emphasis id="B931E27D4267FFDD03C5BFE347950A87" bold="true" box="[197,428,151,177]" pageId="9" pageNumber="10">Coloration in life.</emphasis>
Field observations show that the species is an overall transparent fish, with a thin reddish band above the clearly visible backbone.
</paragraph>
</subSubSection>
<paragraph id="8BFA3E6F4266FFDC039FB94F43AF0D8E" pageId="8" pageNumber="9">
<table id="F945CCCF4266002B039FB94F43AF0D8E" box="[159,1430,1595,1976]" gridcols="3" gridrows="10" pageId="8" pageNumber="9">
<tr id="35753C2D4266002B039FB94F43AF0C66" box="[159,1430,1595,1616]" gridrow="0" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB94F44190C66" box="[159,544,1595,1616]" gridcol="0" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B931E27D4266FFDC039FB94F46D10C66" bold="true" box="[159,232,1595,1616]" pageId="8" pageNumber="9">Counts</emphasis>
</th>
<th id="76A455514266002B004CB94F459D0C66" box="[844,932,1595,1616]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B931E27D4266FFDC004CB94F459D0C66" bold="true" box="[844,932,1595,1616]" pageId="8" pageNumber="9">holotype</emphasis>
</th>
<th id="76A455514266002B00F4B94F43AF0C66" box="[1012,1430,1595,1616]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">
<emphasis id="B931E27D4266FFDC0714B94F43AF0C66" bold="true" box="[1044,1430,1595,1616]" pageId="8" pageNumber="9">paratypes + MRAC 87-24-P-87 (bold)</emphasis>
</th>
</tr>
<tr id="35753C2D4266002B039FB91743AF0C4E" box="[159,1430,1635,1656]" gridrow="1" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB91744190C4E" box="[159,544,1635,1656]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">Dorsal-fin rays</th>
<td id="76A455514266002B004CB917459D0C4E" box="[844,932,1635,1656]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">12</td>
<td id="76A455514266002B00F4B91743AF0C4E" box="[1012,1430,1635,1656]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">
12 (3), 13 (
<emphasis id="B931E27D4266FFDC0630B91743710C4E" bold="true" box="[1328,1352,1635,1656]" pageId="8" pageNumber="9">24</emphasis>
), 14 (4)
</td>
</tr>
<tr id="35753C2D4266002B039FB9FF43AF0C96" box="[159,1430,1675,1696]" gridrow="2" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB9FF44190C96" box="[159,544,1675,1696]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">Anal-fin rays</th>
<td id="76A455514266002B004CB9FF459D0C96" box="[844,932,1675,1696]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">12</td>
<td id="76A455514266002B00F4B9FF43AF0C96" box="[1012,1430,1675,1696]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">
12 (
<emphasis id="B931E27D4266FFDC0630B9FF43710C96" bold="true" box="[1328,1352,1675,1696]" pageId="8" pageNumber="9">24</emphasis>
), 13 (7)
</td>
</tr>
<tr id="35753C2D4266002B039FB9C743AF0CFE" box="[159,1430,1715,1736]" gridrow="3" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB9C744190CFE" box="[159,544,1715,1736]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">Pelvic-fin rays</th>
<td id="76A455514266002B004CB9C7459D0CFE" box="[844,932,1715,1736]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">8</td>
<td id="76A455514266002B00F4B9C743AF0CFE" box="[1012,1430,1715,1736]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">
7 (1), 8 (
<emphasis id="B931E27D4266FFDC063CB9C7436D0CFE" bold="true" box="[1340,1364,1715,1736]" pageId="8" pageNumber="9">24</emphasis>
), 9 (2)
</td>
</tr>
<tr id="35753C2D4266002B039FB9AF43AF0CC6" box="[159,1430,1755,1776]" gridrow="4" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB9AF44190CC6" box="[159,544,1755,1776]" gridcol="0" gridrow="4" pageId="8" pageNumber="9">Pectoral-fin rays</th>
<td id="76A455514266002B004CB9AF459D0CC6" box="[844,932,1755,1776]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">13</td>
<td id="76A455514266002B00F4B9AF43AF0CC6" box="[1012,1430,1755,1776]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">
12 (2), 13 (
<emphasis id="B931E27D4266FFDC0630B9AF43710CC6" bold="true" box="[1328,1352,1755,1776]" pageId="8" pageNumber="9">24</emphasis>
), 14 (5)
</td>
</tr>
<tr id="35753C2D4266002B039FB87743AF0D2E" box="[159,1430,1795,1816]" gridrow="5" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB87744190D2E" box="[159,544,1795,1816]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">Scales on lateral line</th>
<td id="76A455514266002B004CB877459D0D2E" box="[844,932,1795,1816]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">40</td>
<td id="76A455514266002B00F4B87743AF0D2E" box="[1012,1430,1795,1816]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">
39 (2), 40 (5), 41 (15), 42 (6), 43 (
<emphasis id="B931E27D4266FFDC0682B87743B70D2E" bold="true" box="[1410,1422,1795,1816]" pageId="8" pageNumber="9">3</emphasis>
)
</td>
</tr>
<tr id="35753C2D4266002B039FB85F43AF0D76" box="[159,1430,1835,1856]" gridrow="6" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB85F44190D76" box="[159,544,1835,1856]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">Scales between dorsal and adipose fin</th>
<td id="76A455514266002B004CB85F459D0D76" box="[844,932,1835,1856]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">12</td>
<td id="76A455514266002B00F4B85F43AF0D76" box="[1012,1430,1835,1856]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">11 (1), 12 (22), 13 (7)</td>
</tr>
<tr id="35753C2D4266002B039FB82743AF0D5E" box="[159,1430,1875,1896]" gridrow="7" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB82744190D5E" box="[159,544,1875,1896]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">Scales between LL and dorsal-fin origin</th>
<td id="76A455514266002B004CB827459D0D5E" box="[844,932,1875,1896]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">5.5</td>
<td id="76A455514266002B00F4B82743AF0D5E" box="[1012,1430,1875,1896]" gridcol="2" gridrow="7" pageId="8" pageNumber="9">
4.5 (
<emphasis id="B931E27D4266FFDC062BB827430E0D5E" bold="true" box="[1323,1335,1875,1896]" pageId="8" pageNumber="9">2</emphasis>
), 5.5 (29)
</td>
</tr>
<tr id="35753C2D4266002B039FB80F43AF0DA6" box="[159,1430,1915,1936]" gridrow="8" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB80F44190DA6" box="[159,544,1915,1936]" gridcol="0" gridrow="8" pageId="8" pageNumber="9">Scale between LL and pelvic-fin origin</th>
<td id="76A455514266002B004CB80F459D0DA6" box="[844,932,1915,1936]" gridcol="1" gridrow="8" pageId="8" pageNumber="9">5</td>
<td id="76A455514266002B00F4B80F43AF0DA6" box="[1012,1430,1915,1936]" gridcol="2" gridrow="8" pageId="8" pageNumber="9">
4 (
<emphasis id="B931E27D4266FFDC063CB80F436D0DA6" bold="true" box="[1340,1364,1915,1936]" pageId="8" pageNumber="9">29</emphasis>
), 5 (2)
</td>
</tr>
<tr id="35753C2D4266002B039FB8D743AF0D8E" box="[159,1430,1955,1976]" gridrow="9" pageId="8" pageNumber="9">
<th id="76A455514266002B039FB8D744190D8E" box="[159,544,1955,1976]" gridcol="0" gridrow="9" pageId="8" pageNumber="9">Teeth in upper jaw</th>
<td id="76A455514266002B004CB8D7459D0D8E" box="[844,932,1955,1976]" gridcol="1" gridrow="9" pageId="8" pageNumber="9">12</td>
<td id="76A455514266002B00F4B8D743AF0D8E" box="[1012,1430,1955,1976]" gridcol="2" gridrow="9" pageId="8" pageNumber="9">
10 (1), 11 (1), 12 (9), 13 (1), 14 (
<emphasis id="B931E27D4266FFDC0630B8D743710D8E" bold="true" box="[1328,1352,1955,1976]" pageId="8" pageNumber="9">14</emphasis>
), 16 (1)
</td>
</tr>
</table>
</paragraph>
<subSubSection id="C35F6DE44267FFDD03C5BF9142A40B45" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BFA3E6F4267FFDD03C5BF9142A40B45" blockId="9.[151,1437,151,796]" pageId="9" pageNumber="10">
<emphasis id="B931E27D4267FFDD03C5BF91475C0AC9" bold="true" box="[197,357,229,255]" pageId="9" pageNumber="10">Distribution.</emphasis>
Almost all samples of
<taxonomicName id="4C4545EC4267FFDD0172BF9145600AC8" box="[626,857,229,254]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4267FFDD0172BF9145600AC8" box="[626,857,229,254]" italics="true" pageId="9" pageNumber="10">Nannocharax zebra</emphasis>
</taxonomicName>
were collected in the Cameroonian portion of the Cross basin, i.e., riffles in the outlet of the Badi River into the Cross River and in the Cross River directly at Mamfé at a distance of less than one mile north of outlet of the Badi River. One additional albeit slighty aberrant specimen (see below) was collected in the Sombreiro River at Odieke,
<collectingCountry id="F3527EFF4267FFDD073EBE2D42A00B45" box="[1086,1177,345,371]" name="Nigeria" pageId="9" pageNumber="10">Nigeria</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F6DE44267FFDD03C5BE0B4343089E" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BFA3E6F4267FFDD03C5BE0B470E0BF6" blockId="9.[151,1437,151,796]" pageId="9" pageNumber="10">
<emphasis id="B931E27D4267FFDD03C5BE0B476B0BAF" bold="true" box="[197,338,383,409]" pageId="9" pageNumber="10">Etymology.</emphasis>
The species name
<taxonomicName id="4C4545EC4267FFDD0134BEF4444C0BAF" box="[564,629,384,409]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4267FFDD0134BEF4444C0BAF" box="[564,629,384,409]" italics="true" pageId="9" pageNumber="10">zebra</emphasis>
</taxonomicName>
is an allusion to the striped pattern of preserved specimens. A noun in apposition.
</paragraph>
<paragraph id="8BFA3E6F4267FFDD03C5BEB94343089E" blockId="9.[151,1437,151,796]" pageId="9" pageNumber="10">
<emphasis id="B931E27D4267FFDD03C5BEB947A20BD1" bold="true" box="[197,411,461,487]" pageId="9" pageNumber="10">Ecological notes.</emphasis>
<typeStatus id="54FE80CD4267FFDD02A4BEB947D90BD1" box="[420,480,461,487]" pageId="9" pageNumber="10">Type</typeStatus>
specimens were collected inshore in areas of relatively calm water among fine twigs of a fallen tree. These twigs were not reaching soil, but remained attached to the crown of the fallen tree and reaching into the free water column. At the peak of the dry season (February), water level is very low in the Cross River at Mamfé and visibility is high resulting in clear water. Specimens sometimes rested on the fine twigs, but mostly were seen moving or hovering in the free water column in a water depth of approx.
<quantity id="4CBD938A4267FFDD0397BDFA4723089E" box="[151,282,654,680]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="9" pageNumber="10" unit="cm" value="75.0" valueMax="100.0" valueMin="50.0">50–100 cm</quantity>
(between the twigs), well above the bottom which consisted of leaflitter, sand and fine detritus.
</paragraph>
</subSubSection>
<subSubSection id="C35F6DE44267FFDD03C5BDC1433B092A" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BFA3E6F4267FFDD03C5BDC1433B092A" blockId="9.[151,1437,151,796]" pageId="9" pageNumber="10">
<emphasis id="B931E27D4267FFDD03C5BDC1446C08F9" bold="true" box="[197,597,693,719]" pageId="9" pageNumber="10">Remarks on special specimen.</emphasis>
The single specimen MRAC 87-24-P-87 from Sombreiro River in
<collectingCountry id="F3527EFF4267FFDD0397BDA846D608C0" box="[151,239,732,758]" name="Nigeria" pageId="9" pageNumber="10">Nigeria</collectingCountry>
(
<figureCitation id="137E22EA4267FFDD03FFBDA8477F08C0" box="[255,326,732,758]" captionStart="FIGURE 4" captionStartId="9.[151,259,1248,1272]" captionTargetBox="[151,1435,844,1236]" captionTargetId="figure@9.[151,1436,838,1237]" captionTargetPageId="9" captionText="FIGURE 4. Nannocharax zebra, MRAC 87 - 24 - P- 87, 40.7 mm SL, Nigeria, Sombreiro River at Odieke, showing preserved coloration in alcohol." httpUri="https://zenodo.org/record/186237/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
,
<tableCitation id="C6C70BD44267FFDD0253BDA8479308C0" box="[339,426,732,758]" captionStart="TABLE 3" captionStartId="8.[151,239,155,179]" captionText="TABLE 3. Measurements and counts for types of N. zebra sp. nov and N. zebra. MRAC 87 - 24 - P- 87." pageId="9" pageNumber="10">Table 3</tableCitation>
) is similar but slightly different from the
<typeStatus id="54FE80CD4267FFDD0090BDA845FA08C0" box="[912,963,732,758]" pageId="9" pageNumber="10">type</typeStatus>
series of
<taxonomicName id="4C4545EC4267FFDD0734BDA842AE08C3" box="[1076,1175,732,757]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4267FFDD0734BDA842AE08C3" box="[1076,1175,732,757]" italics="true" pageId="9" pageNumber="10">N. zebra</emphasis>
</taxonomicName>
. Unfortunately, this is the only specimen. It is considered conspecific with
<taxonomicName id="4C4545EC4267FFDD01FDBC764559092D" box="[765,864,770,795]" class="Actinopterygii" family="Distichodontidae" genus="Nannocharax" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="zebra">
<emphasis id="B931E27D4267FFDD01FDBC764559092D" box="[765,864,770,795]" italics="true" pageId="9" pageNumber="10">N. zebra</emphasis>
</taxonomicName>
, but not included in the
<typeStatus id="54FE80CD4267FFDD077EBC764288092A" box="[1150,1201,770,796]" pageId="9" pageNumber="10">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>
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