237 lines
28 KiB
XML
237 lines
28 KiB
XML
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<mods:title id="A829D9B7EDDFCF52411D905B5AF94B6F">Monograph of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species (Insecta: Coleoptera)</mods:title>
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<taxonomicName id="8BACF1CFFFDFC917FCF58B89851E711F" ID-CoL="8Y3YJ" authority="ASSING" authorityName="Assing & Brachat & Meybohm" authorityYear="2019" box="[892,1149,1514,1541]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="265" phylum="Arthropoda" rank="species" species="candicus" status="sp. nov.">
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<emphasis id="7ED8565EFFDFC917FCF58B89854F711F" box="[892,1068,1514,1541]" italics="true" pageId="26" pageNumber="265">Gabrius candicus</emphasis>
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ASSING
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</taxonomicName>
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<taxonomicNameLabel id="65EBEB25FFDFC917FB0D8B89858E711F" box="[1156,1261,1514,1541]" pageId="26" pageNumber="265" rank="species">spec. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="04B6D9C7FFDFC917FCF588518281715C" box="[892,994,1586,1606]" pageId="26" pageNumber="265" type="description">
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<paragraph id="4C138A4CFFDFC917FCF588518281715C" blockId="26.[892,1419,1514,1606]" box="[892,994,1586,1606]" pageId="26" pageNumber="265">
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(
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<figureCitation id="D49796C9FFDFC917FC09885182BD715C" box="[896,990,1586,1606]" captionStart-0="Figs 85–99" captionStart-1="Figs 100–113" captionStartId-0="11.[174,214,2005,2027]" captionStartId-1="12.[212,252,2002,2024]" captionTargetBox-0="[186,1413,263,1983]" captionTargetBox-1="[219,1459,263,1990]" captionTargetId-0="figure-49@11.[174,1421,255,1994]" captionTargetId-1="figure-41@12.[212,1459,255,1994]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Figs 85–99: Cretotyphlus idanus (85), C. chanianus (86–91), Pseudobium creticum (92–96), and Gabrius candicus (97–99): male sternite VIII (85, 88, 94); habitus (86, 92, 97); forebody in ventral view (87); aedeagus in lateral and in ventral view (89–90, 95–96, 98–99); female genital segments (91); forebody (93). Scale bars: 92–93, 97: 1.0 mm; 86: 0.5 mm; 98–99: 0.2 mm; 85, 87–91, 94–96: 0.1 mm." captionText-1="Figs 100–113: Gabrius candicus (100–101) and Cephennium spp. of the C. arcuatum group: C. arcuatum (102–103), C. thripticum (104–105), C. selinonum (106–108), C. meybohmi (109–110), and C. idanum (111–113): male sternite VIII (100); female abdominal segments IX–X (101); habitus (102, 106); aedeagus in lateral and in ventral view (103–105, 107–113). Scale bars: 102, 106: 0.5 mm; 100–101: 0.2 mm; 103–105, 107–113: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4755258" figureDoi-1="http://doi.org/10.5281/zenodo.4755262" httpUri-0="https://zenodo.org/record/4755258/files/figure.png" httpUri-1="https://zenodo.org/record/4755262/files/figure.png" pageId="26" pageNumber="265">Figs 97–101</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="04B6D9C7FFDFC917FCDF880382C37042" pageId="26" pageNumber="265" type="materials_examined">
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<paragraph id="4C138A4CFFDFC917FCDF880382C37042" blockId="26.[854,1459,1631,1880]" pageId="26" pageNumber="265">
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<emphasis id="7ED8565EFFDFC917FCDF88038293716D" bold="true" box="[854,1008,1631,1656]" pageId="26" pageNumber="265">Type material</emphasis>
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:
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: “GR –
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[4a],
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,
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,
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<location id="4973DC97FFDFC917FB8488E385337182" LSID="urn:lsid:plazi:treatment:C4053B5AFFDFC916FCF58B8982647F22:4973DC97FFDFC917FB8488E385337182" box="[1037,1104,1664,1688]" latitude="35.37111" longLatPrecision="19" longitude="23.631945" name="Limni" pageId="26" pageNumber="265" stateProvince="Crete">Limni</location>
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,
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,
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,
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, pond margin,
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<date id="3812AC8CFFDFC917FC7688C3851E71A2" box="[1023,1149,1696,1720]" pageId="26" pageNumber="265" value="2018-12-23">
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/ Holotypus ♂
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<taxonomicName id="8BACF1CFFFDFC917FCDF88A3856871C2" authority="Assing & Brachat & Meybohm, 2019" authorityName="Assing & Brachat & Meybohm" authorityYear="2019" box="[854,1035,1728,1752]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="265" phylum="Arthropoda" rank="species" species="candicus" status="sp. nov.">
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<emphasis id="7ED8565EFFDFC917FCDF88A3856871C2" box="[854,1035,1728,1752]" italics="true" pageId="26" pageNumber="265">Gabrius candicus</emphasis>
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<taxonomicNameLabel id="65EBEB25FFDFC917FB9888A3852A71CD" box="[1041,1097,1728,1751]" pageId="26" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
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det.
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<determinerName id="B9CF6688FFDFC917FBF788A3858371C2" box="[1150,1248,1728,1752]" pageId="26" pageNumber="265">
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V.
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<bibRefCitation id="283DF7BDFFDFC917FB1088A3858371C2" author="ASSING, V." box="[1177,1248,1728,1752]" pageId="26" pageNumber="265" pagination="197 - 238" refId="ref47469" refString="ASSING, V. 2019: Monograph of the Staphylinidae of Crete (Greece). Part I. Diversity and endemism (Insecta: Coleoptera). - Contributions to Entomology 69 (2): 197 - 238." type="journal article" year="2019">Assing</bibRefCitation>
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</determinerName>
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2019” (cAss)
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</materialsCitation>
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.
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<materialsCitation id="FCC48011FFDFC917FAF388A382FF7042" ID-GBIF-Occurrence="3458714580" collectingDate="2019-03-20" collectionCode="NHMW" collectorName="V. Assing" elevation="590" latitude="35.37111" location="Limni" longLatPrecision="19" longitude="35.371387" pageId="26" pageNumber="265" specimenCount="1" stateProvince="Kriti" typeStatus="paratype">
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<typeStatus id="931734EEFFDFC917FAF388A382F271E2" pageId="26" pageNumber="265" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="5AAA41C5FFDFC917FC148883828071E2" box="[925,995,1760,1784]" pageId="26" pageNumber="265" type="male">11 ♂♂</specimenCount>
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,
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<specimenCount id="5AAA41C5FFDFC917FC7A8883854F71E2" box="[1011,1068,1760,1784]" pageId="26" pageNumber="265" type="female">6 ♀♀</specimenCount>
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: same data as
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<typeStatus id="931734EEFFDFC917FB5B8883845371E2" box="[1234,1328,1760,1784]" pageId="26" pageNumber="265" type="holotype">holotype</typeStatus>
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(cAss, cFel,
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<collectionCode id="2ABD1289FFDFC917FCDF896382D77002" box="[854,948,1792,1816]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="26" pageNumber="265" type="Museum">NHMW</collectionCode>
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);
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<specimenCount id="5AAA41C5FFDFC917FC4E896385627002" box="[967,1025,1792,1816]" pageId="26" pageNumber="265" type="male">4 ♂♂</specimenCount>
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,
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||
<specimenCount id="5AAA41C5FFDFC917FB868963852B7002" box="[1039,1096,1792,1816]" pageId="26" pageNumber="265" type="female">4 ♀♀</specimenCount>
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: “
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<geoCoordinate id="2998EC8BFFDFC917FBD6896384237002" box="[1119,1344,1792,1816]" degrees="35" direction="north" minutes="22" orientation="latitude" pageId="26" pageNumber="265" precision="15" seconds="17" value="35.371387">N35°22'17 E23°37'57</geoCoordinate>
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||
, GR Westkreta, Limni
|
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<elevation id="C7816D7FFFDFC917FC5489438543702D" box="[989,1056,1824,1848]" metricMagnitude="2" metricUnit="m" metricValue="5.9" pageId="26" pageNumber="265" unit="m" value="590.0">590 m</elevation>
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</quantity>
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,
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<date id="3812AC8CFFDFC917FBA1894385F37022" box="[1064,1168,1824,1848]" pageId="26" pageNumber="265" value="2019-03-20">
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<collectingDate id="28565564FFDFC917FBA1894385F37022" box="[1064,1168,1824,1848]" pageId="26" pageNumber="265" value="2019-03-20">20.3.2019</collectingDate>
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</date>
|
||
, Brachat & Meybohm (12)” (cAss)
|
||
</materialsCitation>
|
||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B6D9C7FFDFC917FCDF89E3847E70A2" pageId="26" pageNumber="265" type="etymology">
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<paragraph id="4C138A4CFFDFC917FCDF89E3847E70A2" blockId="26.[854,1459,1920,1976]" pageId="26" pageNumber="265">
|
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<emphasis id="7ED8565EFFDFC917FCDF89E382AE7082" bold="true" box="[854,973,1920,1944]" pageId="26" pageNumber="265">Etymology</emphasis>
|
||
: The specific epithet is an adjective derived from
|
||
<collectingRegion id="8E6844AEFFDFC917FC1889C3828370A2" box="[913,992,1952,1976]" country="Greece" name="Kriti" pageId="26" pageNumber="265">Candia</collectingRegion>
|
||
, the Venetian name for
|
||
<collectingRegion id="8E6844AEFFDFC917FB5489C3847970A2" box="[1245,1306,1952,1976]" country="Greece" name="Kriti" pageId="26" pageNumber="265">Crete</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B6D9C7FFDFC916FCDF89BC832B71CD" lastPageId="27" lastPageNumber="266" pageId="26" pageNumber="265" type="description">
|
||
<paragraph id="4C138A4CFFDFC916FCDF89BC83267682" blockId="26.[854,1459,2015,2104]" lastBlockId="27.[174,779,256,1336]" lastPageId="27" lastPageNumber="266" pageId="26" pageNumber="265">
|
||
<emphasis id="7ED8565EFFDFC917FCDF89BC82BB70ED" bold="true" box="[854,984,2015,2039]" pageId="26" pageNumber="265">Description</emphasis>
|
||
: Body length 4.2–6.0 mm (measured to posterior margin of tergite VIII); length of forebody
|
||
<quantity id="8B5427A9FFDFC917FCDF864382B47F2D" box="[854,983,2080,2104]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="26" pageNumber="265" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4–2.9 mm</quantity>
|
||
. Habitus as in
|
||
<figureCitation id="D49796C9FFDFC917FB1E864385827F22" box="[1175,1249,2080,2104]" captionStart="Figs 85–99" captionStartId="11.[174,214,2005,2027]" captionTargetBox="[186,1413,263,1983]" captionTargetId="figure-49@11.[174,1421,255,1994]" captionTargetPageId="11" captionText="Figs 85–99: Cretotyphlus idanus (85), C. chanianus (86–91), Pseudobium creticum (92–96), and Gabrius candicus (97–99): male sternite VIII (85, 88, 94); habitus (86, 92, 97); forebody in ventral view (87); aedeagus in lateral and in ventral view (89–90, 95–96, 98–99); female genital segments (91); forebody (93). Scale bars: 92–93, 97: 1.0 mm; 86: 0.5 mm; 98–99: 0.2 mm; 85, 87–91, 94–96: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755258" httpUri="https://zenodo.org/record/4755258/files/figure.png" pageId="26" pageNumber="265">Fig. 97</figureCitation>
|
||
. Coloration: body black, except for the partly dark-yellowish abdominal segments VIII–X; legs dark-yellowish with the femora and tibiae partly infuscate; antennae blackish with (parts of) antennomere I, the base of antennomere II, and often also the base of antennomere III paler.
|
||
</paragraph>
|
||
<paragraph id="4C138A4CFFDEC916FF278FC3801275CD" blockId="27.[174,779,256,1336]" pageId="27" pageNumber="266">
|
||
Head without sexual dimorphism, approximately 1.15 times as long as broad, lateral margins behind eyes usually weakly converging in dorsal view; dorsal surface with fine microsculpture. Eyes more or less distinctly shorter than postocular region in dorsal view. punctation rather coarse and dense, sparse in median dorsal portion; interstices without microsculpture. Eyes large, approximately 0.7 times as long as postocular portion in dorsal view. Antenna
|
||
<quantity id="8B5427A9FFDEC916FEC58CC380AF75AD" box="[332,460,672,696]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="27" pageNumber="266" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
|
||
long; preapical antennomeres weakly transverse.
|
||
</paragraph>
|
||
<paragraph id="4C138A4CFFDEC916FF278C8380F07442" blockId="27.[174,779,256,1336]" pageId="27" pageNumber="266">Pronotum approximately 1.25 times as long as broad and 1.15 times as broad as head; dorsal series composed of 1+5 punctures; disc with nearly obsolete to distinct transverse microsculpture.</paragraph>
|
||
<paragraph id="4C138A4CFFDEC916FF278D03804274C2" blockId="27.[174,779,256,1336]" pageId="27" pageNumber="266">Elytra approximately 0.7 times as long as pronotum; punctation moderately dense and moderately coarse; interstices without microsculpture. Hind wings fully developed.</paragraph>
|
||
<paragraph id="4C138A4CFFDEC916FF278D8383FB7362" blockId="27.[174,779,256,1336]" pageId="27" pageNumber="266">Abdomen with dense and fine punctation, the density gradually decreasing from anterior towards posterior tergites; interstices with very fine and shallow transverse microsculpture visible only at high magnification; posterior margin of tergite VII with palisade fringe.</paragraph>
|
||
<paragraph id="4C138A4CFFDEC916FF278AE281927202" blockId="27.[174,779,256,1336]" pageId="27" pageNumber="266">
|
||
♂: sternite VIII (
|
||
<figureCitation id="D49796C9FFDEC916FEE88AE380DA7382" box="[353,441,1152,1176]" captionStart="Figs 100–113" captionStartId="12.[212,252,2002,2024]" captionTargetBox="[219,1459,263,1990]" captionTargetId="figure-41@12.[212,1459,255,1994]" captionTargetPageId="12" captionText="Figs 100–113: Gabrius candicus (100–101) and Cephennium spp. of the C. arcuatum group: C. arcuatum (102–103), C. thripticum (104–105), C. selinonum (106–108), C. meybohmi (109–110), and C. idanum (111–113): male sternite VIII (100); female abdominal segments IX–X (101); habitus (102, 106); aedeagus in lateral and in ventral view (103–105, 107–113). Scale bars: 102, 106: 0.5 mm; 100–101: 0.2 mm; 103–105, 107–113: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755262" httpUri="https://zenodo.org/record/4755262/files/figure.png" pageId="27" pageNumber="266">Fig. 100</figureCitation>
|
||
) posteriorly membranous, with shallowly concave posterior excision, and with long setae and longitudinal sculpture in posterior third; aedeagus (
|
||
<figureCitation id="D49796C9FFDEC916FF3C8A83804F73E2" box="[181,300,1248,1272]" captionStart="Figs 85–99" captionStartId="11.[174,214,2005,2027]" captionTargetBox="[186,1413,263,1983]" captionTargetId="figure-49@11.[174,1421,255,1994]" captionTargetPageId="11" captionText="Figs 85–99: Cretotyphlus idanus (85), C. chanianus (86–91), Pseudobium creticum (92–96), and Gabrius candicus (97–99): male sternite VIII (85, 88, 94); habitus (86, 92, 97); forebody in ventral view (87); aedeagus in lateral and in ventral view (89–90, 95–96, 98–99); female genital segments (91); forebody (93). Scale bars: 92–93, 97: 1.0 mm; 86: 0.5 mm; 98–99: 0.2 mm; 85, 87–91, 94–96: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755258" httpUri="https://zenodo.org/record/4755258/files/figure.png" pageId="27" pageNumber="266">Figs 98–99</figureCitation>
|
||
) 1.00–
|
||
<quantity id="8B5427A9FFDEC916FEFA8A8380B273ED" box="[371,465,1248,1272]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="27" pageNumber="266" unit="mm" value="1.05">1.05 mm</quantity>
|
||
long and of highly distinctive shape.
|
||
</paragraph>
|
||
<paragraph id="4C138A4CFFDEC916FF278B4280B67222" blockId="27.[174,779,256,1336]" box="[174,469,1312,1336]" pageId="27" pageNumber="266">
|
||
♀: terminalia as in
|
||
<figureCitation id="D49796C9FFDEC916FEF38B4380B27222" box="[378,465,1312,1336]" captionStart="Figs 100–113" captionStartId="12.[212,252,2002,2024]" captionTargetBox="[219,1459,263,1990]" captionTargetId="figure-41@12.[212,1459,255,1994]" captionTargetPageId="12" captionText="Figs 100–113: Gabrius candicus (100–101) and Cephennium spp. of the C. arcuatum group: C. arcuatum (102–103), C. thripticum (104–105), C. selinonum (106–108), C. meybohmi (109–110), and C. idanum (111–113): male sternite VIII (100); female abdominal segments IX–X (101); habitus (102, 106); aedeagus in lateral and in ventral view (103–105, 107–113). Scale bars: 102, 106: 0.5 mm; 100–101: 0.2 mm; 103–105, 107–113: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4755262" httpUri="https://zenodo.org/record/4755262/files/figure.png" pageId="27" pageNumber="266">Fig. 101</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="4C138A4CFFDEC916FF278B3C832B71CD" blockId="27.[174,779,1375,1752]" pageId="27" pageNumber="266">
|
||
<emphasis id="7ED8565EFFDEC916FF278B3C80E67262" bold="true" box="[174,389,1375,1400]" pageId="27" pageNumber="266">Comparative notes</emphasis>
|
||
: Based on external and the male sexual characters,
|
||
<taxonomicName id="8BACF1CFFFDEC916FEE68BE3808B7282" authorityName="Assing & Brachat & Meybohm" authorityYear="2019" box="[367,488,1408,1432]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="266" phylum="Arthropoda" rank="species" species="candicus">
|
||
<emphasis id="7ED8565EFFDEC916FEE68BE3808B7282" box="[367,488,1408,1432]" italics="true" pageId="27" pageNumber="266">G. candicus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="8BACF1CFFFDEC916FD0E8BE382687282" baseAuthorityName="GRAVENHORST" baseAuthorityYear="1802" box="[647,779,1408,1432]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="266" phylum="Arthropoda" rank="species" species="nigritulus">
|
||
<emphasis id="7ED8565EFFDEC916FD0E8BE382687282" box="[647,779,1408,1432]" italics="true" pageId="27" pageNumber="266">G. nigritulus</emphasis>
|
||
</taxonomicName>
|
||
group. It is easily distinguished from other representatives of this group only by the male sexual characters, particularly the conspicuous shape of the aedeagus. It is additionally distinguished from the similar
|
||
<taxonomicName id="8BACF1CFFFDEC916FD0E886380C67122" authority="(GRAVENHORST, 1802)" baseAuthorityName="GRAVENHORST" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="266" phylum="Arthropoda" rank="species" species="nigritulus">
|
||
<emphasis id="7ED8565EFFDEC916FD0E886382687102" box="[647,779,1536,1560]" italics="true" pageId="27" pageNumber="266">G. nigritulus</emphasis>
|
||
(GRAVENHORST, 1802)
|
||
</taxonomicName>
|
||
, the only other
|
||
<taxonomicName id="8BACF1CFFFDEC916FDE9884382687122" box="[608,779,1568,1592]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="266" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="7ED8565EFFDEC916FDE9884383D77122" box="[608,692,1568,1592]" italics="true" pageId="27" pageNumber="266">Gabrius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
known from
|
||
<collectingRegion id="8E6844AEFFDEC916FEB7882380197142" box="[318,378,1600,1624]" country="Greece" name="Kriti" pageId="27" pageNumber="266">Crete</collectingRegion>
|
||
, by more distinctly infuscate femora and tibiae, a posteriorly usually tapering head, on average larger eyes, less pronounced microsculpture on the abdomen, and by a posteriorly more acute female tergite IX with more numerous long apical setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="04B6D9C7FFDEC916FF27889C82647F22" pageId="27" pageNumber="266" type="distribution">
|
||
<paragraph id="4C138A4CFFDEC916FF27889C82647F22" blockId="27.[174,780,1791,2104]" pageId="27" pageNumber="266">
|
||
<emphasis id="7ED8565EFFDEC916FF27889C8343700D" bold="true" box="[174,544,1791,1816]" pageId="27" pageNumber="266">Distribution and natural history</emphasis>
|
||
: The
|
||
<typeStatus id="931734EEFFDEC916FDEE896383F67002" box="[615,661,1792,1816]" pageId="27" pageNumber="266">type</typeStatus>
|
||
locality is situated to the south of Kissamos in West
|
||
<collectingRegion id="8E6844AEFFDEC916FD1C894383B17022" box="[661,722,1824,1848]" country="Greece" name="Kriti" pageId="27" pageNumber="266">Crete</collectingRegion>
|
||
. The specimens were sifted from debris and grass roots at the margin of an artificial pond at an altitude of
|
||
<specimenCount id="5AAA41C5FFDEC916FD18890383B8706D" box="[657,731,1888,1912]" pageId="27" pageNumber="266" type="generic">
|
||
<quantity id="8B5427A9FFDEC916FD18890383B4706D" box="[657,727,1888,1912]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="27" pageNumber="266" unit="m" value="580.0">580 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
The species is most likely endemic in
|
||
<collectingRegion id="8E6844AEFFDEC916FD9689E3833A7082" box="[543,601,1920,1944]" country="Greece" name="Kriti" pageId="27" pageNumber="266">Crete</collectingRegion>
|
||
and was probably widespread in the island. The fact that it had never been recorded before suggests that it is very rare today, presumably because suitable habitats (natural lakes, ponds, swamps) have almost completely disappeared as a result of extensive agricultural and other human activity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |