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<document id="4AF6B6D1A9B9A3344CA128CD730CBBBB" ID-CLB-Dataset="20304" ID-DOI="10.5852/ejt.2022.798.1675" ID-GBIF-Dataset="eb4e5239-a946-4a99-a5a3-5e7197f39ca3" ID-ISSN="2118-9773" ID-Zenodo-Dep="6341178" ID-ZooBank="6C11886D-D217-4D37-AAD7-EA9B7EBE5361" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1646830985801" checkinUser="felipe" docAuthor="Molero-Baltanás, Rafael, Gaju-Ricart, Miquel, Fišer, Žiga, Bach de Roca, Carmen &amp; Mendes, Luís F." docDate="2022" docId="9B31C7625C4EFFAD76738778FF7BFA36" docLanguage="en" docName="ejt-798_127-161.pdf" docOrigin="European Journal of Taxonomy 798 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.798.1675" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Coletinia tinauti Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca 1997" docType="treatment" docVersion="5" lastPageNumber="153" masterDocId="6708BF1A5C57FFB777088325FF8BFFCB" masterDocTitle="Three new species of European Coletinia Wygodzinsky (Zygentoma, Nicoletiidae), with additional records and an updated identification key" masterLastPageNumber="161" masterPageNumber="127" pageNumber="152" updateTime="1699317574945" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1068EFDE89EF46A387B0B5CFDA239622">Three new species of European Coletinia Wygodzinsky (Zygentoma, Nicoletiidae), with additional records and an updated identification key</mods:title>
</mods:titleInfo>
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<mods:namePart id="C60384A2932E593EE29707683C9A75E8">Molero-Baltanás, Rafael</mods:namePart>
<mods:nameIdentifier id="F7543788748ED77BE135815EBB6028AF" type="LSID">826D3806-9744-4C48-8600-5EE8E034A13F</mods:nameIdentifier>
<mods:affiliation id="36A5312B1060434156572BDA0584F5F3">Department of Zoology, University of Córdoba, C- 1 Campus de Rabanales, 14071 Córdoba, Spain</mods:affiliation>
<mods:nameIdentifier id="809F462BEEF3097CE407981E7BFB5864" type="email">ba1mobar@uco.es</mods:nameIdentifier>
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<mods:namePart id="4B04930223E4BB5848A4D080F8C4D84B">Gaju-Ricart, Miquel</mods:namePart>
<mods:nameIdentifier id="330FB7C6AA887D00869D59B73567C6BF" type="LSID">89B6B7B3-20A7-4ADD-80C8-F801D40067F8</mods:nameIdentifier>
<mods:affiliation id="F440B26D24851A74A1EBFAB5608BD90F">Department of Zoology, University of Córdoba, C- 1 Campus de Rabanales, 14071 Córdoba, Spain</mods:affiliation>
<mods:nameIdentifier id="F40FA18AAEE601C84EEAB23DFC996E00" type="email">ba1garim@uco.es</mods:nameIdentifier>
</mods:name>
<mods:name id="AF5028338938C8A5F2E1860E0A905015" type="personal">
<mods:role id="7E562D24CFF4D0CC1ECCF878694E87D4">
<mods:roleTerm id="6819A5650A5BE18AEFAE90F3B9ED6A84">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FFFC4CA2C80D1114460FE597389FC509">Fišer, Žiga</mods:namePart>
<mods:nameIdentifier id="77DB5EF802F00BE2988798BDB5AEA988" type="LSID">6673CC2F-4C75-4309-A564-E9A1CCC30C67</mods:nameIdentifier>
<mods:affiliation id="C67880EE3A34A384FF772B48DEACB5F1">Department of Biology, Biotechnical Faculty, University of Ljubljana, Jamnikarjeva 101, SI- 1000 Ljubljana, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="008003CD438F919CFA57593853F2A0B7" type="email">ziga.fiser@bf.uni-lj.si</mods:nameIdentifier>
</mods:name>
<mods:name id="C8A79DA29FFC86CC5932F3A59AE07837" type="personal">
<mods:role id="099F7D89B991F0EDA743D038D99C48A1">
<mods:roleTerm id="02F85E23D03D5ED16DB973B56D39CB6C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1012D93F83860B4FA8229244D77A5A96">Bach de Roca, Carmen</mods:namePart>
<mods:nameIdentifier id="76BEC91FD78CE487CE0D5D7A359FA149" type="LSID">941C01DE-052D-465D-B5F4-976591FB514E</mods:nameIdentifier>
<mods:affiliation id="F93CF4F34BF15773E7C8DD526DFB6AA0">Calle Mestral, 13. 08230 Matadepera, Barcelona, Spain</mods:affiliation>
<mods:nameIdentifier id="DDA39F551B7343443E613E9F86317EA2" type="email">Carmen.Bach@uab.cat</mods:nameIdentifier>
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<mods:namePart id="EED742C892BE8B085459BFF5DE86254C">Mendes, Luís F.</mods:namePart>
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<mods:affiliation id="E3393B90D5CD6EB8F995CDBB28D10E3C">Universidade de Lisboa. Museu Nacional de História Natural (MUHNAC), Rua da Escola Politécnica, 58, PT- 1250 - 102, Lisbon, Portugal.</mods:affiliation>
<mods:nameIdentifier id="CFBE2ED9B0ECA403C9E93570D3DB3FD5" type="email">luisfmendes@edu.ulisboa.pt</mods:nameIdentifier>
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<mods:title id="49293C4B3F4258119E3F43427FC5392D">European Journal of Taxonomy</mods:title>
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<mods:date id="0323A5E5041CCE510C7C811E74DE0F73">2022</mods:date>
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<mods:number id="8E7DE7373B75D9755F171245F1D39AC6">2022-03-04</mods:number>
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<mods:number id="37B8E6070E8AE9948F787AE463F20345">798</mods:number>
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<mods:start id="D5DB9376193E49BE3575FE962CECA2E3">127</mods:start>
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<treatment id="9B31C7625C4EFFAD76738778FF7BFA36" ID-DOI="http://doi.org/10.5281/zenodo.6344265" ID-GBIF-Taxon="193895179" ID-Zenodo-Dep="6344265" LSID="urn:lsid:plazi:treatment:9B31C7625C4EFFAD76738778FF7BFA36" httpUri="http://treatment.plazi.org/id/9B31C7625C4EFFAD76738778FF7BFA36" lastPageId="26" lastPageNumber="153" pageId="25" pageNumber="152">
<subSubSection id="5B8225FF5C4EFFAE76738778FB32FBBC" box="[379,1209,1117,1144]" pageId="25" pageNumber="152" type="nomenclature">
<paragraph id="132776745C4EFFAE76738778FB32FBBC" blockId="25.[379,1209,1117,1144]" box="[379,1209,1117,1144]" pageId="25" pageNumber="152">
<heading id="486FC1185C4EFFAE76738778FB32FBBC" box="[379,1209,1117,1144]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="152" reason="2">
<taxonomicName id="D4980DF75C4EFFAE76738778FB32FBBC" authority="Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca, 1997" authorityName="Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca" authorityYear="1997" box="[379,1209,1117,1144]" class="Insecta" family="Nicoletiidae" genus="Coletinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="152" phylum="Arthropoda" rank="species" species="tinauti">
<emphasis id="21ECAA665C4EFFAE76738778FDCAFBBC" bold="true" box="[379,577,1117,1144]" italics="true" pageId="25" pageNumber="152">Coletinia tinauti</emphasis>
<bibRefCitation id="77090B855C4EFFAE75408778FB32FBBC" author="Molero-Baltanas R. &amp; Gaju-Ricart M. &amp; Bach de Roca C." box="[584,1209,1117,1144]" pageId="25" pageNumber="152" pagination="95 - 110" refId="ref18466" refString="Molero-Baltanas R., Gaju-Ricart M. &amp; Bach de Roca C. 1997. Coletinia tinauti n. sp., nueva especie de Nicoletiidae de Andalucia (Insecta, Apterygota, Zygentoma). Zoologica Baetica 8: 95 - 110." type="journal article" year="1997">Molero-Baltanás, Gaju-Ricart &amp; Bach de Roca, 1997</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="132776745C4EFFAE77B587BBFB5BFB72" blockId="25.[189,1232,1182,1209]" box="[189,1232,1182,1209]" pageId="25" pageNumber="152">
<treatmentCitationGroup id="3388515A5C4EFFAE77B587BBFB5BFB72" box="[189,1232,1182,1209]" pageId="25" pageNumber="152">
<treatmentCitation id="923950655C4EFFAE77B587BBFBD4FB72" author="Molero-Baltanas R. &amp; Gaju-Ricart M. &amp; Bach de Roca C." box="[189,1119,1182,1209]" page="104" pageId="25" pageNumber="152" year="1997">
<taxonomicName id="D4980DF75C4EFFAE77B587BBFBD4FB72" authority="Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca, 1997: 97 - 104" authorityName="Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca" authorityPageNumber="97 - 104" authorityYear="1997" box="[189,1119,1182,1209]" class="Insecta" family="Nicoletiidae" genus="Coletinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="152" phylum="Arthropoda" rank="species" species="tinauti">
<emphasis id="21ECAA665C4EFFAE77B587BBFEF4FB73" box="[189,383,1182,1208]" italics="true" pageId="25" pageNumber="152">Coletinia tinauti</emphasis>
<bibRefCitation id="77090B855C4EFFAE768D87BBFBD4FB72" author="Molero-Baltanas R. &amp; Gaju-Ricart M. &amp; Bach de Roca C." box="[389,1119,1182,1209]" pageId="25" pageNumber="152" pagination="95 - 110" refId="ref18466" refString="Molero-Baltanas R., Gaju-Ricart M. &amp; Bach de Roca C. 1997. Coletinia tinauti n. sp., nueva especie de Nicoletiidae de Andalucia (Insecta, Apterygota, Zygentoma). Zoologica Baetica 8: 95 - 110." type="journal article" year="1997">Molero-Baltanás, Gaju-Ricart &amp; Bach de Roca, 1997: 97–104</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1–4.
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</paragraph>
<paragraph id="132776745C4EFFAE77B587C5FB8AFB30" blockId="25.[189,1025,1248,1275]" box="[189,1025,1248,1275]" pageId="25" pageNumber="152">
<treatmentCitationGroup id="3388515A5C4EFFAE77B587C5FB8AFB30" box="[189,1025,1248,1275]" pageId="25" pageNumber="152">
<taxonomicName id="D4980DF75C4EFFAE77B587C5FEF4FB31" authorityName="Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca" authorityYear="1997" box="[189,383,1248,1274]" class="Insecta" family="Nicoletiidae" genus="Coletinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="152" phylum="Arthropoda" rank="species" species="tinauti">
<emphasis id="21ECAA665C4EFFAE77B587C5FEF4FB31" box="[189,383,1248,1274]" italics="true" pageId="25" pageNumber="152">Coletinia tinauti</emphasis>
</taxonomicName>
–
<treatmentCitation id="923950655C4EFFAE769387C5FD5AFB30" author="Molero R. &amp; Bach C. &amp; Sendra A. &amp; Montagud S. &amp; Barranco P. &amp; Gaju M." box="[411,721,1248,1275]" httpUri="http://treatment.plazi.org/id/03C0A74CFFFBFFD4FF4F6FE8E60C96DC" page="49" pageId="25" pageNumber="152" year="2013">
<bibRefCitation id="77090B855C4EFFAE769387C5FD5AFB30" author="Molero R. &amp; Bach C. &amp; Sendra A. &amp; Montagud S. &amp; Barranco P. &amp; Gaju M." box="[411,721,1248,1275]" pageId="25" pageNumber="152" pagination="1 - 60" refId="ref18336" refString="Molero R., Bach C., Sendra A., Montagud S., Barranco P. &amp; Gaju M. 2013. Revision of the genus Coletinia (Zygentoma: Nicoletiidae) in the Iberian Peninsula, with descriptions of nine new species. Zootaxa 3615 (1): 1 - 60. https: // doi. org / 10.11646 / zootaxa. 3615.1.1" type="journal article" year="2013">
Molero
<emphasis id="21ECAA665C4EFFAE76F187C4FDBFFB31" box="[505,564,1248,1274]" italics="true" pageId="25" pageNumber="152">et al.</emphasis>
2013: 49–52
</bibRefCitation>
</treatmentCitation>
, figs 1b, 6b, 9b, 153–163.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5B8225FF5C4EFFAD77B58604FF7BFA36" lastPageId="26" lastPageNumber="153" pageId="25" pageNumber="152" type="materials_examined">
<subSubSection id="5B8225FF5C4EFFAE77B58604FE9AFA45" pageId="25" pageNumber="152" type="materials_examined">
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<emphasis id="21ECAA665C4EFFAE77B58604FE21FAF0" bold="true" box="[189,426,1313,1339]" pageId="25" pageNumber="152">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="132776745C4EFFAE77B58675FE9AFA45" blockId="25.[189,1399,1360,1422]" pageId="25" pageNumber="152">
<materialsCitation id="A3F07C295C4EFFAE77B58675FE85FA45" ID-GBIF-Occurrence="3699120312" collectingDate="2013-12-24" collectionCode="UCO" collectorName="GEV" country="Spain" location="Jaen Province" municipality="Cueva de la Morciguilla" pageId="25" pageNumber="152" specimenCount="22" specimenCount-female="8" specimenCount-male="14">
<collectingCountry id="6B8F36E45C4EFFAE77B58675FE84FAA1" box="[189,271,1360,1386]" name="Spain" pageId="25" pageNumber="152">SPAIN</collectingCountry>
•
<specimenCount id="059EBDFD5C4EFFAE76238675FE0BFAA1" box="[299,384,1360,1386]" pageId="25" pageNumber="152" type="male">14 ♂♂</specimenCount>
,
<specimenCount id="059EBDFD5C4EFFAE76868675FE58FAA1" box="[398,467,1360,1386]" pageId="25" pageNumber="152" type="female">8 ♀♀</specimenCount>
;
<location id="164720AF5C4EFFAE76EB8675FD00FAA1" LSID="urn:lsid:plazi:treatment:9B31C7625C4EFFAD76738778FF7BFA36:164720AF5C4EFFAE76EB8675FD00FAA1" box="[483,651,1360,1386]" country="Spain" municipality="Cueva de la Morciguilla" name="Jaen Province" pageId="25" pageNumber="152">Jaén Province</location>
,
<location id="164720AF5C4EFFAE75918675FC36FAA1" LSID="urn:lsid:plazi:treatment:9B31C7625C4EFFAD76738778FF7BFA36:164720AF5C4EFFAE75918675FC36FAA1" box="[665,957,1360,1386]" country="Spain" municipality="Cueva de la Morciguilla" name="Cueva de la Morciguilla" pageId="25" pageNumber="152">Cueva de la Morciguilla</location>
;
<date id="672650B45C4EFFAE74C38675FBE4FAA1" box="[971,1135,1360,1386]" pageId="25" pageNumber="152" value="2013-12-24">
<collectingDate id="7762A95C5C4EFFAE74C38675FBE4FAA1" box="[971,1135,1360,1386]" pageId="25" pageNumber="152" value="2013-12-24">24 Dec. 2013</collectingDate>
</date>
;
<collectorName id="BE6D13A25C4EFFAE73758675FB32FAA1" box="[1149,1209,1360,1386]" pageId="25" pageNumber="152">GEV</collectorName>
leg.;
<collectionCode id="7589EEB15C4EFFAE73F68675FAB7FAA1" box="[1278,1340,1360,1386]" pageId="25" pageNumber="152">UCO</collectionCode>
Ref. Z2507
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5B8225FF5C4EFFAD77B58691FF7BFA36" lastPageId="26" lastPageNumber="153" pageId="25" pageNumber="152" type="description">
<paragraph id="132776745C4EFFAE77B58691FE3FFA04" blockId="25.[189,436,1460,1487]" box="[189,436,1460,1487]" pageId="25" pageNumber="152">
<heading id="486FC1185C4EFFAE77B58691FE3FFA04" bold="true" box="[189,436,1460,1487]" fontSize="11" level="3" pageId="25" pageNumber="152" reason="3">
<emphasis id="21ECAA665C4EFFAE77B58691FE3FFA04" bold="true" box="[189,436,1460,1487]" pageId="25" pageNumber="152">Variability remarks</emphasis>
</heading>
</paragraph>
<paragraph id="132776745C4EFFAE77B586C6FEC1F896" blockId="25.[188,1400,1507,1885]" pageId="25" pageNumber="152">
The abundant material belonging to this species collected at one locality (
<specimenCount id="059EBDFD5C4EFFAE731886C6FB3AFA35" box="[1040,1201,1507,1534]" pageId="25" pageNumber="152" type="generic">22 specimens</specimenCount>
) allows to assess the variability of
<taxonomicName id="D4980DF75C4EFFAE768E8522FE7EF9EB" authorityName="Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca" authorityYear="1997" box="[390,501,1543,1569]" class="Insecta" family="Nicoletiidae" genus="Coletinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="152" phylum="Arthropoda" rank="species" species="tinauti">
<emphasis id="21ECAA665C4EFFAE768E8522FE7EF9EB" box="[390,501,1543,1569]" italics="true" pageId="25" pageNumber="152">C. tinauti</emphasis>
</taxonomicName>
. This sample comes from a cave about
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distance from the
<typeStatus id="CC23C8D65C4EFFAE73ED8522FA93F9EA" box="[1253,1304,1543,1569]" pageId="25" pageNumber="152">type</typeStatus>
locality of the species, and the insects collected completely fit the original description. Some specimens of this sample have been dissected and examined, concluding that most of their characters are proven to be constant inside the population, such as the shape of the urosternite VIII of males or the number of divisions of the ovipositor. Nevertheless, the variability of some characters proves to be wider than previously known for this species (see Table 1). For example, the shape of the hind margin of the urotergite X in males proves to be variable, since in some specimens it is folded in its median part or is more convex dorsally but more straight ventrally; this agrees with the variability detected for this character in
<taxonomicName id="D4980DF75C4EFFAE76458405FE3CF8F1" baseAuthorityName="Grassi" baseAuthorityYear="1887" box="[333,439,1824,1850]" class="Insecta" family="Nicoletiidae" genus="Coletinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="152" phylum="Arthropoda" rank="species" species="maggi">
<emphasis id="21ECAA665C4EFFAE76458405FE3CF8F1" box="[333,439,1824,1850]" italics="true" pageId="25" pageNumber="152">C. maggi</emphasis>
</taxonomicName>
by
<bibRefCitation id="77090B855C4EFFAE76EB8405FC99F8F0" author="Gilgado J. D. &amp; Ortuno V. M." box="[483,786,1824,1851]" pageId="25" pageNumber="152" pagination="85 - 100" refId="ref17981" refString="Gilgado J. D. &amp; Ortuno V. M. 2015. Intra- and inter-population polymorphism in Coletinia maggii (Grassi, 1887) (Zygentoma: Nicoletiidae), an inhabitant of soil, mesovoid shallow substratum (MSS) and caves - A challenge for the strict classification of subterranean fauna? Zootaxa 3920 (1): 85 - 100. https: // doi. org / 10.11646 / zootaxa. 3920.1.5" type="journal article" year="2015">Gilgado &amp; Ortuño (2015)</bibRefCitation>
, although this variability is not detected in
<taxonomicName id="D4980DF75C4EFFAE72008405FAFCF8F1" authorityName="Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca" authorityYear="1997" box="[1288,1399,1824,1850]" class="Insecta" family="Nicoletiidae" genus="Coletinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="25" pageNumber="152" phylum="Arthropoda" rank="species" species="tinauti">
<emphasis id="21ECAA665C4EFFAE72008405FAFCF8F1" box="[1288,1399,1824,1850]" italics="true" pageId="25" pageNumber="152">C. tinauti</emphasis>
</taxonomicName>
for females.
</paragraph>
<paragraph id="132776745C4EFFAE77B584A1FAFDF82E" blockId="25.[189,1399,1924,2021]" pageId="25" pageNumber="152">
The terminal filaments of this species present few pegs with blunt apex (at most 5 but usually 4 or less in each cercus). The paracercus has only, as usual, short, pigmented spines. The formula of the paracercus of the
<typeStatus id="CC23C8D65C4EFFAE768684EEFE7EF82E" box="[398,501,1995,2021]" pageId="25" pageNumber="152" type="holotype">holotype</typeStatus>
is P1 (1[1
<emphasis id="21ECAA665C4EFFAE756784EEFDF5F82E" box="[623,638,1995,2021]" italics="true" pageId="25" pageNumber="152">d</emphasis>
]) +P2 (1[0]); in this specimen the paracercus is broken and only
</paragraph>
<paragraph id="132776745C4DFFAD77B5822CFD39FE8C" blockId="26.[188,1398,265,327]" pageId="26" pageNumber="153">
the first division is preserved. The illustration presented by
<bibRefCitation id="77090B855C4DFFAD7479822CFB46FEE8" author="Molero-Baltanas R. &amp; Gaju-Ricart M. &amp; Bach de Roca C." box="[881,1229,265,291]" pageId="26" pageNumber="153" pagination="95 - 110" refId="ref18466" refString="Molero-Baltanas R., Gaju-Ricart M. &amp; Bach de Roca C. 1997. Coletinia tinauti n. sp., nueva especie de Nicoletiidae de Andalucia (Insecta, Apterygota, Zygentoma). Zoologica Baetica 8: 95 - 110." type="journal article" year="1997">
Molero-Baltanás
<emphasis id="21ECAA665C4DFFAD7335822FFBFCFEE8" box="[1085,1143,265,291]" italics="true" pageId="26" pageNumber="153">et al.</emphasis>
(1997)
</bibRefCitation>
in the original description corresponds in fact to a cercus.
</paragraph>
<paragraph id="132776745C4DFFAD77B5824BFD6BFD6A" blockId="26.[185,1400,366,674]" pageId="26" pageNumber="153">
Variability of the paracercus in other male specimens examined: the first division of the cercus bears in some specimens (as the
<typeStatus id="CC23C8D65C4DFFAD76D582B4FDCDFE67" box="[477,582,401,428]" pageId="26" pageNumber="153" type="holotype">holotype</typeStatus>
) one thin acute and small dorsal spine; in other specimens this spine is absent. The second division shows 3 more robust and short dorsal spines (the third longer than the preceding ones) inserted on alternate rings of setae. If the dorsal spine on P1 is absent, these spines are present on the first, third and fifth rings of setae; if the dorsal spine on P1 is present, the modified spines are inserted on the second, fourth and sixth rings of setae. So, the formula can be P1 (1[0]) +P2 (1[1d] +2[0]+ 3[1d] +4[0]+ 5[1d]) or P1 (1[1d])+P2 (1[0]+ 2[1d] +3[0] +4[1d] +5[0] +6[1d]). All these spines are acute but clearly different (shorter, more robust, and pigmented) to the unmodified setae of the dorsal side of the paracercus of females.
</paragraph>
<paragraph id="132776745C4DFFAD77B581ECFB9CFC11" blockId="26.[185,1400,713,986]" pageId="26" pageNumber="153">
In cerci, the inner-dorsal spines are more robust than the inner-ventral ones and usually show blunt apex, except those in the C3, and can be considered as pegs, but there is some variability (for example, in the
<typeStatus id="CC23C8D65C4DFFAD77B5802AFEAFFCE2" box="[189,292,783,809]" pageId="26" pageNumber="153" type="holotype">holotype</typeStatus>
there is only one peg in the first ring of C2, see
<figureCitation id="8BA36AF15C4DFFAD745D802AFC24FCE2" box="[853,943,783,809]" captionStart="Fig" captionStartId="3.[189,232,1770,1796]" captionTargetBox="[197,1388,546,1721]" captionTargetId="figure-105@3.[189,1398,538,1732]" captionTargetPageId="3" captionText="Fig. 1. Characters of taxonomic interest in Coletinia Wygodzinsky, 1980.A–C. Coletinia tinauti Molero, Gaju &amp; Bach, 1997, holotype. A. Micrographs of a pedicellar apophysis. B. Urotergite X. C. Basal part of the left cercus. Abbreviations: C1, C2 and C3 = first, second and third divisions of the cercus (the limit between the first and the second is better in focus); D = disc of the urotergite; id = spines or pegs inserted inner dorsal position; the first id spine in the C2 has blunt apex and can be considered as a peg; iL = inner length of the apophysis; iv = spines inserted in inner ventral position; Ms = marginal setae; oL = outer length of the apophysis; P = pedicel; Ss = submarginal setae." figureDoi="http://doi.org/10.5281/zenodo.6341184" httpUri="https://zenodo.org/record/6341184/files/figure.png" pageId="26" pageNumber="153">Fig. 1C</figureCitation>
). The remaining spines are thinner and frequently acute but always shortened compared with setae in similar position in females. The formula of the left cercus of the
<typeStatus id="CC23C8D65C4DFFAD76D08070FDB4FCA4" box="[472,575,853,879]" pageId="26" pageNumber="153" type="holotype">holotype</typeStatus>
(shown in
<figureCitation id="8BA36AF15C4DFFAD75CC8070FC81FCBB" box="[708,778,853,880]" captionStart="Fig" captionStartId="8.[189,232,1886,1912]" captionTargetBox="[201,1359,288,1837]" captionTargetId="figure-16@8.[197,1390,280,1845]" captionTargetPageId="8" captionText="Fig. 3. Coletinia dalmatica Molero-Baltanás, Fišer, Bach de Roca &amp; Gaju-Ricart sp. nov. Holotype, ♂ (MNCN_Ent 283558). A. Mandible. B. Maxilla and maxillary palp. C. Labial palps and anterior part of the labium. D. Thoracic nota. E. First leg, excluding apical tarsomeres. F. Femur, tibia, tarsal articles and praetarsus of the third leg. Scale bars: A, C, F = 0.1 mm; B, D–E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6341188" httpUri="https://zenodo.org/record/6341188/files/figure.png" pageId="26" pageNumber="153">Fig. 3</figureCitation>
) is: C1 (1[1id +1iv]) +C2 (1[1
<emphasis id="21ECAA665C4DFFAD737D8070FB07FCA4" box="[1141,1164,853,879]" italics="true" pageId="26" pageNumber="153">id</emphasis>
+1iv] +2[1id])+C3 (1[1id + 1iv]+ 2[1id]); the right cercus is drawn by Molero
<emphasis id="21ECAA665C4DFFAD747E805CFC39FC58" box="[886,946,889,915]" italics="true" pageId="26" pageNumber="153">et al.</emphasis>
(1997: fig. 3.6), except for the second ring of setae of C3, which is damaged (in the caption it is incorrectly indicated as paracercus) and its formula is C1 (1[1
<emphasis id="21ECAA665C4DFFAD769E809AFE26FC12" box="[406,429,959,985]" italics="true" pageId="26" pageNumber="153">id</emphasis>
+1
<emphasis id="21ECAA665C4DFFAD76DC80E5FE62FC12" box="[468,489,960,985]" italics="true" pageId="26" pageNumber="153">iv</emphasis>
])+C2 (1[1
<emphasis id="21ECAA665C4DFFAD7567809AFD0DFC12" box="[623,646,959,985]" italics="true" pageId="26" pageNumber="153">id</emphasis>
+ 1iv]+ 2[1id]) +C3 (1[1id +1iv]).
</paragraph>
<paragraph id="132776745C4DFFAD77B58725FECFFB6C" blockId="26.[188,1401,1024,1191]" pageId="26" pageNumber="153">
Variability of the cercus in other male specimens examined: the limit between the second and the third division of the cercus has a more distal position in most of the remaining specimens observed, so the spines of the first and second ring of setae of C
<quantity id="D460DB915C4DFFAD74498762FCE5FBAA" box="[833,878,1095,1121]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="26" pageNumber="153" unit="in" value="3.0">3 in</quantity>
the
<typeStatus id="CC23C8D65C4DFFAD74AA8762FB81FBAA" box="[930,1034,1095,1121]" pageId="26" pageNumber="153" type="holotype">holotype</typeStatus>
correspond in these additional specimens to the third and fourth ring of setae of C2. But the arrangement of pegs is similar, with some exceptions:
</paragraph>
<paragraph id="132776745C4DFFAD77DD87EAFD86FAC7" blockId="26.[213,1398,1231,1363]" pageId="26" pageNumber="153">– The pegs of C1 are absent in some specimens; in this case, the apex of the spine of the third ring of setae of the C2 is blunt.</paragraph>
<paragraph id="132776745C4DFFAD77DD8633FEFFFA98" blockId="26.[213,1398,1231,1363]" pageId="26" pageNumber="153">– Some spines inserted on inner-ventral position can be absent or reduced to thin usual setae (not modified).</paragraph>
<paragraph id="132776745C4DFFAD77B5865FFF7BFA36" blockId="26.[189,1399,1402,1533]" pageId="26" pageNumber="153">
Considering this variability, the formula of cerci of most specimens of
<taxonomicName id="D4980DF75C4DFFAD7315865FFB04FA5F" authorityName="Molero-Baltanas, Gaju-Ricart &amp; Bach de Roca" authorityYear="1997" box="[1053,1167,1402,1428]" class="Insecta" family="Nicoletiidae" genus="Coletinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Zygentoma" pageId="26" pageNumber="153" phylum="Arthropoda" rank="species" species="tinauti">
<emphasis id="21ECAA665C4DFFAD7315865FFB04FA5F" box="[1053,1167,1402,1428]" italics="true" pageId="26" pageNumber="153">C. tinauti</emphasis>
</taxonomicName>
where the division between C2 and C3 is placed beyond the fourth ring of setae is: C1(1[0–1
<emphasis id="21ECAA665C4DFFAD734A86B8FBD2FA7C" box="[1090,1113,1437,1463]" italics="true" pageId="26" pageNumber="153">id</emphasis>
+0–1iv]) + C2(1[1
<emphasis id="21ECAA665C4DFFAD723986B8FAC3FA7C" box="[1329,1352,1437,1463]" italics="true" pageId="26" pageNumber="153">id</emphasis>
+0– 1iv] + 2[1
<emphasis id="21ECAA665C4DFFAD762686E5FECEFA11" box="[302,325,1472,1498]" italics="true" pageId="26" pageNumber="153">id</emphasis>
+0–1iv] +3[1
<emphasis id="21ECAA665C4DFFAD76E686E5FD8EFA11" box="[494,517,1472,1498]" italics="true" pageId="26" pageNumber="153">id</emphasis>
+ 0–1iv] + 4 [0–1
<emphasis id="21ECAA665C4DFFAD75DD86E5FD67FA11" box="[725,748,1472,1498]" italics="true" pageId="26" pageNumber="153">id</emphasis>
+ 0–1iv]). The maximum number of blunt pegs in C2 is 4.
</paragraph>
</subSubSection>
</subSubSection>
</treatment>
</document>
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