treatments-xml/data/81/6E/AF/816EAF991457F6E11EC5505CE004DC46.xml
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<document ID-CLB-Dataset="297124" ID-DOI="10.5281/zenodo.7316519" ID-GBIF-Dataset="3f0f641c-cd4b-4694-8e47-c47be23ac2a9" ID-ISBN="0-8018-8221-4" checkinTime="1670905116760" checkinUser="guido" docAuthor="Wilson, Don E. &amp; Reeder, DeeAnn" docDate="2005" docId="816EAF991457F6E11EC5505CE004DC46" docLanguage="en" docName="MammalSpeciesofTheWorld.2005.Carnivora.532-628.xml" docOrigin="Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1, Baltimore: The Johns Hopkins University Press" docTitle="Helarctos Horsfield 1825" docType="treatment" docVersion="4" id="E69BF2968277AF6992A8FF32E9D4E7ED" lastPageNumber="587" masterDocId="E69BF2968277AF6992A8FF32E9D4E7ED" masterDocTitle="Order Carnivora" masterLastPageNumber="628" masterPageNumber="532" pageNumber="587" updateTime="1716792096583" updateUser="ExternalLinkService">
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<mods:title id="9C9D9E9F599B2656CC97FF346730F57E">Order Carnivora</mods:title>
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<mods:namePart id="966A5E4DAB281500FEC8D0B2A018DA15">Wilson, Don E.</mods:namePart>
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<mods:namePart id="224DF02EBF2072AED1293E37C8C4B479">Reeder, DeeAnn</mods:namePart>
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<mods:title id="CB82B3159A07F021F615FFDC9664DB3B">Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 1</mods:title>
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<treatment id="816EAF991457F6E11EC5505CE004DC46" ID-DOI="http://doi.org/10.5281/zenodo.11344926" ID-GBIF-Taxon="231548378" ID-Zenodo-Dep="11344926" LSID="urn:lsid:plazi:treatment:816EAF991457F6E11EC5505CE004DC46" httpUri="http://treatment.plazi.org/id/816EAF991457F6E11EC5505CE004DC46" lastPageNumber="587" pageNumber="587" sourceUrl="http://www.departments.bucknell.edu/biology/resources/msw3/browse.asp?id=14000942">
<subSubSection id="DC3C5E3583837F025D73A95AE0960495" pageNumber="587" type="nomenclature">
<paragraph id="56DD00EA7D848DBA445311F7EA815EB3" pageNumber="587">
<taxonomicName id="FBF229CB027AC53ABD5A5D0980BB6C7B" ID-CoL="62TH6" authority="Horsfield 1825" authorityName="Horsfield" authorityYear="1825" class="Mammalia" family="Ursidae" genus="Helarctos" kingdom="Animalia" order="Carnivora" pageNumber="587" phylum="Chordata" rank="genus">
<emphasis id="630DE407AB606E3277AC1DB80B4465BA" italics="true" pageNumber="587">Helarctos</emphasis>
Horsfield 1825
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="DAFC680138205945BF7012EC25847F8C" pageNumber="587" type="reference_group">
<paragraph id="72420405EE7B2B54F6BCACBE096E1A29" pageNumber="587">
<treatmentCitation id="A7A854AD99838069CA044156F66180B7" author="Horsfield" authority="Horsfield 1825" authorityName="Horsfield" authorityYear="1825" class="Mammalia" family="Ursidae" genus="Helarctos" issue="6" journal="Zool. J." kingdom="Animalia" order="Carnivora" page="221" pageNumber="587" phylum="Chordata" rank="genus" title="Helarctos" volume="2" year="1825">
<taxonomicName id="DBA14678123D613FD7BE1C22F8358B79" ID-CoL="62TH6" authority="Horsfield 1825" authorityName="Horsfield" authorityYear="1825" class="Mammalia" family="Ursidae" genus="Helarctos" kingdom="Animalia" order="Carnivora" pageNumber="587" phylum="Chordata" rank="genus">
<emphasis id="51BF900BF45A5DAA29DFFF041713FF9B" italics="true" pageNumber="587">Helarctos</emphasis>
<bibRefCitation id="1BCC92BD64BA66CE66F22086C4F531FC" author="Horsfield, T." refId="ref159312" refString="Horsfield, T. 1825. Description of the RIMAU-DAHAN of the inhabitants of Sumatra, a new species of Felis, discovered in the forests of Bencoolen, by Sir T. Stamford Raffles, late Lieutenant Governor of Fort Marlborough, &amp; c., &amp; c., &amp; c. Zoological Journal, 1 (4): 542 - 554." year="1825">Horsfield 1825</bibRefCitation>
</taxonomicName>
,
<bibCitation id="8746ECC1BABD65A56215179E758AE9EA" author="Horsfield" issue="6" journal="Zool. J." pageNumber="587" pagination="221" title="Helarctos" volume="2" year="1825">Zool. J., 2 (6): 221</bibCitation>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E31804439CE96A07F0EC85E3EAEBA8C5" pageNumber="587" type="type_taxon">
<paragraph id="52A1968408BDB015B1C9882204AA7479" pageNumber="587">
<emphasis id="AB8F88CDDE23EF93F589933F75C571E1" inLineHeading="true" pageNumber="587">Type Species:</emphasis>
<taxonomicName id="CF129B1B5D0C43E925A252A3E094B0EA" authority="Horsfield 1825" authorityName="Horsfield" authorityYear="1825" class="Mammalia" family="Ursidae" genus="Helarctos" kingdom="Animalia" order="Carnivora" pageNumber="587" phylum="Chordata" rank="species" species="euryspilus">
<emphasis id="06CCD718035A97EBE0678505F20345D5" italics="true" pageNumber="587">Helarctos euryspilus</emphasis>
<bibRefCitation id="AAFD23B10EC00138D64602157822FDCB" author="Horsfield, T." refId="ref159312" refString="Horsfield, T. 1825. Description of the RIMAU-DAHAN of the inhabitants of Sumatra, a new species of Felis, discovered in the forests of Bencoolen, by Sir T. Stamford Raffles, late Lieutenant Governor of Fort Marlborough, &amp; c., &amp; c., &amp; c. Zoological Journal, 1 (4): 542 - 554." year="1825">Horsfield 1825</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="08C1943FD12938DEA04A1AC4E79604FC" pageNumber="587" type="synonym_list">
<paragraph id="69D72597795CC845BA7890BF102643C2" pageNumber="587">
<emphasis id="5243E7AB680C6817CD90750B1B2FCE70" inLineHeading="true" pageNumber="587">Synonyms:</emphasis>
<taxonomicName id="A7980E5B9CB3A03DB0DB2EF293FBC7CE" authority="Gloger 1841" authorityName="Gloger" authorityYear="1841" class="Mammalia" family="Ursidae" genus="Helarctus" kingdom="Animalia" order="Carnivora" pageNumber="587" phylum="Chordata" rank="genus">
<emphasis id="D1EC6DF75F72C906A524519F60979B7C" italics="true" pageNumber="587">Helarctus</emphasis>
Gloger 1841
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E9F1A5B864B26F259CADE7F62BD1E4BF" pageNumber="587" type="child_taxa">
<paragraph id="3186F550F2301439D3F73676D989564C" pageNumber="587">
<emphasis id="0A9A67652063988F09418F1EF0560D7F" inLineHeading="true" pageNumber="587">Species and subspecies:</emphasis>
1 species with 2 subspecies:
</paragraph>
<paragraph id="8D44BA306A91661DD36D4A9FC553B41C" pageNumber="587">
Species
<taxonomicName id="7F11D43E82AA259A7571C38E5ED2D064" authority="(Raffles 1821)" baseAuthorityName="Raffles" baseAuthorityYear="1821" class="Mammalia" family="Ursidae" genus="Helarctos" kingdom="Animalia" order="Carnivora" pageNumber="587" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="F9A0E958D1107505E2BFB8C64637B3D2" italics="true" pageNumber="587">Helarctos malayanus</emphasis>
(
<bibRefCitation id="7031B402E440500E145B17A9D302D8A5" author="Raffles, T. S." refId="ref295881" refString="Raffles, T. S. 1821. Descriptive catalogue of a zoological collection, made on account of the honorable East Indian Company, in the island of Sumatra and its vicinity, under the direction of Sir Thomas Stamford Raffles, Lieutenant-Governor of Fort Marlborough, with additional notices illurstrative of the natural history. Transactions of the Linnean Society of London, 13: 239 - 274." year="1821">Raffles 1821</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8253E31ACF70C7553F9D8C9E71A1CA04" pageNumber="587">
Subspecies
<taxonomicName id="62DF44BD687E956DA66F0F1A5494F3DF" authority="Raffles 1821" authorityName="Raffles" authorityYear="1821" class="Mammalia" family="Ursidae" genus="Helarctos" kingdom="Animalia" order="Carnivora" pageNumber="587" phylum="Chordata" rank="subSpecies" species="malayanus" subSpecies="malayanus">
<emphasis id="6926B73FC8C31FE717E30397E5B09ACB" italics="true" pageNumber="587">Helarctos malayanus</emphasis>
subsp.
<emphasis id="7CA3718B1CF4F14C4E98A840DA2D8183" italics="true" pageNumber="587">malayanus</emphasis>
<bibRefCitation id="A84A631953984229B9E3EDC9B0C14689" author="Raffles, T. S." refId="ref295881" refString="Raffles, T. S. 1821. Descriptive catalogue of a zoological collection, made on account of the honorable East Indian Company, in the island of Sumatra and its vicinity, under the direction of Sir Thomas Stamford Raffles, Lieutenant-Governor of Fort Marlborough, with additional notices illurstrative of the natural history. Transactions of the Linnean Society of London, 13: 239 - 274." year="1821">Raffles 1821</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="80B0A8E2D03516DBF5E5A558AF9D9716" pageNumber="587">
Subspecies
<taxonomicName id="BC1E47F8D72D728670687714FA8B90DE" authority="Horsfield 1825" authorityName="Horsfield" authorityYear="1825" class="Mammalia" family="Ursidae" genus="Helarctos" kingdom="Animalia" order="Carnivora" pageNumber="587" phylum="Chordata" rank="subSpecies" species="malayanus" subSpecies="euryspilus">
<emphasis id="0ECB9DBF06D27F2EF4EE3D8D51689021" italics="true" pageNumber="587">Helarctos malayanus</emphasis>
subsp.
<emphasis id="B4180D421A750B3E69523E9298D7E90D" italics="true" pageNumber="587">euryspilus</emphasis>
<bibRefCitation id="7033D6B1D6C55149332A4CF43FB8F5A3" author="Horsfield, T." refId="ref159312" refString="Horsfield, T. 1825. Description of the RIMAU-DAHAN of the inhabitants of Sumatra, a new species of Felis, discovered in the forests of Bencoolen, by Sir T. Stamford Raffles, late Lieutenant Governor of Fort Marlborough, &amp; c., &amp; c., &amp; c. Zoological Journal, 1 (4): 542 - 554." year="1825">Horsfield 1825</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="0725E7A83A95A7E45B5EABDC66700265" pageNumber="587" type="discussion">
<paragraph id="1771FE686AF2FDAE592590438AF6C900" pageNumber="587">
<emphasis id="25793322C86C4ADC57BA1EE4CA734BD4" inLineHeading="true" pageNumber="587">Discussion:</emphasis>
Revised by
<bibRefCitation id="EAE252305DA96AE37789FE5341836B0C" author="Pocock, R. I." refId="ref290006" refString="Pocock, R. I. 1932 b. The black and brown bears of Europe and Asia. Part II. The sloth bear (Melursus), the Himalayan black bear (Selenarctos) and the Malayan bear (Helarctos). Journal of the Bombay Natural History Society, 36 (1): 101 - 138." year="1932">
Pocock (1932
<emphasis id="2AC8C7D0D5A5F9E876CEF9DAFB601A61" italics="true" pageNumber="587">b</emphasis>
)
</bibRefCitation>
.
<bibRefCitation id="D0A29FEB837E3D906137816DFCCA5290" author="Van Gelder, R. G." refId="ref366220" refString="Van Gelder, R. G. 1977 b. Mammalian hybrids and generic limits. American Museum Novitates, 2635: 1 - 25." year="1977">
<collectingRegion id="17DEECB41F31E4BEEDB61BF096E5F8D4" country="Turkey" name="Van">Van</collectingRegion>
Gelder (1977
<emphasis id="42859374409A183660C04FF4C53333F6" italics="true" pageNumber="587">b</emphasis>
)
</bibRefCitation>
placed
<taxonomicName id="584D6939FA98F753B8E540AF9724B02D" authorityName="Horsfield" authorityYear="1825" class="Mammalia" family="Ursidae" genus="Helarctos" kingdom="Animalia" order="Carnivora" phylum="Chordata" rank="genus">
<emphasis id="897099FBCBA744B1FD545096ACD4A145" italics="true" pageNumber="587">Helarctos</emphasis>
</taxonomicName>
in
<taxonomicName id="D03AC16F0B50DEA490A044F7ECB8C49A" authorityName="Meyer" authorityYear="1793" class="Mammalia" family="Ursidae" genus="Melursus" kingdom="Animalia" order="Carnivora" phylum="Chordata" rank="genus">
<emphasis id="1CCCF12D2E842B4F017D5470ABC5E2C6" italics="true" pageNumber="587">Melursus</emphasis>
</taxonomicName>
.
<bibRefCitation id="B86155BCD74B6CBC9D309F35964C4A27" author="Pocock, R. I." refId="ref290688" refString="Pocock, R. I. 1941 a. The fauna of British India, including Ceylon and Burma. Mammalia. Vol. II. Carnivora (suborders Aeluroidae (part) and Arctoidae). Taylor and Francis, Ltd., London, 503 pp." year="1941">
Pocock (1941
<emphasis id="B7986FEDF62D87AF02EFA0B4A4F257D0" italics="true" pageNumber="587">a</emphasis>
)
</bibRefCitation>
and
<bibRefCitation id="04DFF89CF56B56D52CCBF722154717DB" author="Bininda-Emonds, O. R. P. &amp; J. L. Gittleman &amp; A. Purvis" refId="ref32387" refString="Bininda-Emonds, O. R. P., J. L. Gittleman, and A. Purvis. 1999. Building large trees by combining phylogenetic information: A complete phylogeny of the extant Carnivora (Mammalia). Biological Reviews, 74: 143 - 175." year="1999">Bininda-Emonds et al. (1999)</bibRefCitation>
suggested a close relationship between
<taxonomicName id="AF19CDBA31E083B49C1A15785AEF5FC5" baseAuthorityName="Shaw" baseAuthorityYear="1791" class="Mammalia" family="Ursidae" genus="Melursus" kingdom="Animalia" order="Carnivora" phylum="Chordata" rank="species" species="ursinus">
<emphasis id="0E166017A4D8F6521A6BD9F1A878A37C" italics="true" pageNumber="587">M. ursinus</emphasis>
</taxonomicName>
and
<taxonomicName id="6B12FB76A38D1F8A82F0B20EF41B2711" baseAuthorityName="Raffles" baseAuthorityYear="1821" class="Mammalia" family="Ursidae" genus="Helarctos" kingdom="Animalia" order="Carnivora" phylum="Chordata" rank="species" species="malayanus">
<emphasis id="6F904EBE95449125CBFE12C71DDADDB8" italics="true" pageNumber="587">H. malayanus</emphasis>
</taxonomicName>
; however, this was not supported by
<bibRefCitation id="C96C8F73453139E551565B9E0213228B" author="Goldman, D. P. &amp; R. Giri &amp; S. J. O'Brien" refId="ref122531" refString="Goldman, D. P., R. Giri, and S. J. O'Brien. 1989. Molecular genetic-distance estimates among the Ursidae as indicated by one- and two-dimensional protein electrophoresis. Evolution, 43 (2): 282 - 295." year="1989">Goldman et al. (1989)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>