treatments-xml/data/43/8B/EE/438BEE4825F3B89BB3AE5F5E15021196.xml
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.65.8454" ID-PMC="PMC4957019" ID-Pensoft-Pub="1314-2003-65-15" ID-Pensoft-UUID="FF92CB1F1C79FF802E2A8F4806001B0C" ID-PubMed="27489485" ID-Zenodo-Dep="122979" ModsDocID="1314-2003-65-15" checkinTime="1466061722947" checkinUser="pensoft" docAuthor="Ortiz-Rodriguez, Andres Ernesto, Burelo Ramos, Carlos Manuel &amp; Gomez-Dominguez, Hector" docDate="2016" docId="438BEE4825F3B89BB3AE5F5E15021196" docLanguage="en" docName="PhytoKeys 65: 15-23" docOrigin="PhytoKeys 65" docPubDate="2016-06-15" docSource="http://dx.doi.org/10.3897/phytokeys.65.8454" docTitle="Amphitecna loreae Ortiz-Rodr. &amp; Burelo 2016, sp. nov." docType="treatment" docVersion="7" id="FF92CB1F1C79FF802E2A8F4806001B0C" lastPageNumber="20" masterDocId="FF92CB1F1C79FF802E2A8F4806001B0C" masterDocTitle="A new species of Amphitecna (Bignoniaceae) endemic to Chiapas, Mexico" masterLastPageNumber="23" masterPageNumber="15" pageNumber="17" updateTime="1668141719996" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Amphitecna (Bignoniaceae) endemic to Chiapas, Mexico</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ortiz-Rodriguez, Andres Ernesto</mods:namePart>
<mods:affiliation>Instituto de Ecologia, A. C., Departamento de Biologia Evolutiva, Xalapa, Veracruz, Mexico</mods:affiliation>
<mods:nameIdentifier type="email">ortizrodriguez.ae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burelo Ramos, Carlos Manuel</mods:namePart>
<mods:affiliation>Herbario UJAT, Division Academica de Ciencias Biologicas, Universidad Juarez, Autonoma de Tabasco, Villahermosa Tabasco, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gomez-Dominguez, Hector</mods:namePart>
<mods:affiliation>Herbario Eizi Matuda (HEM) Facultad de Ciencias Biologicas, Universidad de Ciencias y Artes de Chiapas, Tuxtla Gutierrez, Chiapas, Mexico</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="pubDate">
<mods:number>2016-06-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>65</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>15</mods:start>
<mods:end>23</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.65.8454</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.65.8454</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-65-15</mods:identifier>
<mods:identifier type="Pensoft-UUID">FF92CB1F1C79FF802E2A8F4806001B0C</mods:identifier>
<mods:identifier type="Zenodo-Dep">122979</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127861537" LSID="urn:lsid:plazi:treatment:438BEE4825F3B89BB3AE5F5E15021196" httpUri="http://treatment.plazi.org/id/438BEE4825F3B89BB3AE5F5E15021196" lastPageId="5" lastPageNumber="20" pageId="2" pageNumber="17">
<subSubSection pageId="2" pageNumber="17" type="nomenclature">
<paragraph pageId="2" pageNumber="17">
<taxonomicName LSID="438BEE48-25F3-B89B-B3AE-5F5E15021196" authority="Ortiz-Rodr. &amp; Burelo" authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="loreae" status="sp. nov.">Amphitecna loreae Ortiz-Rodr. &amp; Burelo</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="17">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Amphitecna loreae sp. nov. A cauliflorous flowers with trilabiate calyx. B broadly elliptical to spherical fruits C corolla D buds rounded at apex. Photographs by Andres E. Ortiz-Rodriguez (A and C) and Marcos Escobar (B and D)." figureDoi="10.3897/phytokeys.65.8454.figure1" httpUri="https://binary.pensoft.net/fig/92590" pageId="2" pageNumber="17">Figures 1</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Leaf variation in Amphitecna loreae and related species. A Amphitecna tuxtlensis (H. Gomez 3710 HEM) B Amphitecna montana (N. Martinez 927 HEM) C Amphitecna loreae (M. Escobar 586 HEM) and D Amphitecna latifolia (E. Ucan E 251 XAL)." figureDoi="10.3897/phytokeys.65.8454.figure3" httpUri="https://binary.pensoft.net/fig/92592" pageId="2" pageNumber="17">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="17" type="materials_examined">
<paragraph pageId="2" pageNumber="17">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="2" pageNumber="17">
<materialsCitation collectingDate="2012-03-05" collectorName="de Berriozabal, Ortiz-Rodriguez A. E" country="Mexico" county="Municipio de Berriozabal" elevation="1068" latitude="16.864056" location="Cairo" longLatPrecision="1" longitude="-93.331024" municipality="Vista" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<collectingCountry name="Mexico">Mexico</collectingCountry>
.
<collectingRegion country="Mexico" name="Chiapas">Chiapas</collectingRegion>
,
<collectingCounty>
Municipio
<collectorName>
de
<normalizedToken originalValue="Berriozábal">Berriozabal</normalizedToken>
</collectorName>
</collectingCounty>
, zona sujeta a
<normalizedToken originalValue="protección">proteccion</normalizedToken>
<normalizedToken originalValue="ecológica">ecologica</normalizedToken>
&quot;La Pera&quot;, predio
<normalizedToken originalValue="“Peña">&quot;Pena</normalizedToken>
Flor&quot; camino
<normalizedToken originalValue="Berriozábal-">Berriozabal-</normalizedToken>
<collectingMunicipality>Vista</collectingMunicipality>
Hermosa-El
<location LSID="urn:lsid:plazi:treatment:438BEE4825F3B89BB3AE5F5E15021196:87FE8FE8375EB8AB3FD99C67B046A7C9" country="Mexico" county="Municipio de Berriozabal" latitude="16.864056" longLatPrecision="1" longitude="-93.331024" municipality="Vista" name="Cairo" stateProvince="Chiapas">Cairo</location>
, km. 12
<normalizedToken originalValue="desvío">desvio</normalizedToken>
al
<location LSID="urn:lsid:plazi:treatment:438BEE4825F3B89BB3AE5F5E15021196:883153E676D23EF05504020D060CF0C4" country="Mexico" county="Municipio de Berriozabal" latitude="16.864056" longLatPrecision="1" longitude="-93.331024" municipality="Vista" name="Pozo Turipache" stateProvince="Chiapas">Pozo Turipache</location>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.068" unit="m" value="1068.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.068" unit="m" value="1068.0">1068 m</elevation>
</quantity>
,
<geoCoordinate degrees="16" direction="north" minutes="51" orientation="latitude" precision="1" seconds="50.6" value="16.864056">16°51'50.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="93" direction="west" minutes="19" orientation="longitude" precision="1" seconds="51.7" value="-93.331024">93°19'51.7&quot;W</geoCoordinate>
,
<collectingDate value="2012-03-05">05 March 2012</collectingDate>
(fl, fr)
<emphasis italics="true" pageId="2" pageNumber="17">
<collectorName>
<normalizedToken originalValue="Ortiz-Rodríguez">Ortiz-Rodriguez</normalizedToken>
A. E
</collectorName>
0178
</emphasis>
(
<typeStatus>holotype</typeStatus>
HEM; isotypes: UJAT, XAL)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="17" type="diagnosis">
<paragraph pageId="2" pageNumber="17">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="17">
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna loreae</emphasis>
</taxonomicName>
is distinguishable from the other species of
<taxonomicName authorityName="Miers" authorityYear="1868" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna</emphasis>
</taxonomicName>
by a combination of its cauliflorous inflorescences, large flowers, buds rounded at apex, and broadly elliptical to spherical rather than acuminate fruits.
<taxonomicName authorityName="A. H. Gentry" authorityYear="1980" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna tuxtlensis" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="tuxtlensis">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna tuxtlensis</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="L. O. Williams" authorityYear="1973" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna montana" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="montana">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna montana</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna latifolia" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna latifolia</emphasis>
</taxonomicName>
, also distributed in Mexico, have
<normalizedToken originalValue="affinities">affinities</normalizedToken>
with
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna loreae</emphasis>
</taxonomicName>
and share the cauliflorous inflorescences and leaves less than 50 cm long. However,
<taxonomicName authorityName="A. H. Gentry" authorityYear="1980" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna tuxtlensis" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="tuxtlensis">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna tuxtlensis</emphasis>
</taxonomicName>
differs in having the flower buds pointed at the apex and fruits elliptic, acute to acuminate at apex, and
<taxonomicName authorityName="L. O. Williams" authorityYear="1973" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna montana" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="montana">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna montana</emphasis>
</taxonomicName>
differs in having larger leaves, long pedicellate flowers and elliptical fruits shortly pointed at the tip, while
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna latifolia" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna latifolia</emphasis>
</taxonomicName>
differs in having obovate to widely elliptic leaves, rounded to mucronate at apex with poorly defined petioles (Figure
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Amphitecna loreae sp. nov. A cauliflorous flowers with trilabiate calyx. B broadly elliptical to spherical fruits C corolla D buds rounded at apex. Photographs by Andres E. Ortiz-Rodriguez (A and C) and Marcos Escobar (B and D)." figureDoi="10.3897/phytokeys.65.8454.figure1" httpUri="https://binary.pensoft.net/fig/92590" pageId="2" pageNumber="17">1</figureCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="980339" doi="10.3897/phytokeys.65.8454.figure1" httpUri="https://binary.pensoft.net/fig/92590" pageId="2" pageNumber="17" start="Figure 1" startId="F1">
<paragraph pageId="2" pageNumber="17">
<emphasis bold="true" pageId="2" pageNumber="17">Figure 1.</emphasis>
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="2" pageNumber="17" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="2" pageNumber="17">Amphitecna loreae</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="2" pageNumber="17">A</emphasis>
cauliflorous flowers with trilabiate calyx.
<emphasis bold="true" pageId="2" pageNumber="17">B</emphasis>
broadly elliptical to spherical fruits
<emphasis bold="true" pageId="2" pageNumber="17">C</emphasis>
corolla
<emphasis bold="true" pageId="2" pageNumber="17">D</emphasis>
buds rounded at apex. Photographs by Andres E. Ortiz-Rodriguez (
<emphasis bold="true" pageId="2" pageNumber="17">A</emphasis>
and
<emphasis bold="true" pageId="2" pageNumber="17">C</emphasis>
) and Marcos Escobar (
<emphasis bold="true" pageId="2" pageNumber="17">B</emphasis>
and
<emphasis bold="true" pageId="2" pageNumber="17">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="18" pageId="2" pageNumber="17" type="description">
<paragraph pageId="2" pageNumber="17">Description.</paragraph>
<paragraph lastPageId="3" lastPageNumber="18" pageId="2" pageNumber="17">
Trees, 15-25 m and 15-50 cm DBH, the secondary branches terete.
<emphasis italics="true" pageId="2" pageNumber="17">Leaves</emphasis>
, alternate-verticillate, clustered near the apex of the branches, olive-green when dry, glabrous, 10-20 cm long, 2-5 cm wide, oblanceolate to narrowly elliptic, acuminate, subcoriaceous, acute to attenuate basis, midrib slightly raised on the upper surface, prominent on the lower surface; secondary veins 11-14 on a side, slightly raised above, prominent below; petiole short, to 2 cm long, merging with attenuate leaf base.
<emphasis italics="true" pageId="2" pageNumber="17">Inflorescences</emphasis>
, groups of two or three flowers, with an unpleasant odor, which are borne on leafless portions of old branches and along the main trunk (cauliflory). Flower buds, rounded at apex.
<emphasis italics="true" pageId="2" pageNumber="17">Flowers</emphasis>
pendant, pedicel 35-60 mm long; calyx campanulate, 28-38 mm long, more or less coriaceous, evenly 2 to 3-labiate to below the middle, circumscissile; corolla radially symmetric, pale green, tubular-infundibuliform, 48-60 mm long, 30-40 mm wide at the mouth of the tube, the basal part of the corolla a straight tube, 15-25 mm long, the lobes fused in to frilly-margined rim; stamens included, inserted 18-28 mm from base of the tube, the anther thecae divergent, 4-7
<pageBreakToken pageId="3" pageNumber="18" start="start">mm</pageBreakToken>
long, the filaments 18-30 mm long; the staminode, when present, less than 20 mm long, inserted 10-20 mm from base of the tube, sometimes well developed (with one or two small thecae) to give the impression of being a fifth stamen; ovary, up to 3 mm long and 2.5 mm wide, broadly elliptical, glandular-papillose; pistil 40-60 mm long with the stigma bifurcate; disc annular-pulvinate, about 6 mm in diameter; flowers are often found with signs of herbivory, in which the ovule and disc are not present.
<emphasis italics="true" pageId="3" pageNumber="18">Fruits</emphasis>
broadly elliptical to spherical, 70-100 mm long, 60-80 mm wide.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="18" type="habitat">
<paragraph pageId="3" pageNumber="18">Habitat and ecology.</paragraph>
<paragraph pageId="3" pageNumber="18">
The species is only known from Chiapas, Mexico. It is a rare species at the type locality in the ecological reserve La Pera. The species inhabits the karst areas, mainly in the tropical rainforest. It is a canopy tree and coexists with species of
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus lancifolia" order="Fagales" pageId="3" pageNumber="18" phylum="Angiospermae" rank="species" species="lancifolia">
<emphasis italics="true" pageId="3" pageNumber="18">Quercus lancifolia</emphasis>
</taxonomicName>
Schltdl. &amp; Cham.,
<taxonomicName class="Dicotyledoneae" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quercus corrugata" order="Fagales" pageId="3" pageNumber="18" phylum="Angiospermae" rank="species" species="corrugata">
<emphasis italics="true" pageId="3" pageNumber="18">Quercus corrugata</emphasis>
</taxonomicName>
Hook.,
<taxonomicName class="Magnoliopsida" family="Icacinaceae" genus="Calatola" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Calatola costaricensis" order="Celastrales" pageId="3" pageNumber="18" phylum="Magnoliophyta" rank="species" species="costaricensis">
<emphasis italics="true" pageId="3" pageNumber="18">Calatola costaricensis</emphasis>
</taxonomicName>
Standl.,
<taxonomicName class="Magnoliopsida" family="Acanthaceae" genus="Spathacanthus" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" lsidName="Spathacanthus hahnianus" order="Lamiales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="species" species="hahnianus">
<emphasis italics="true" pageId="3" pageNumber="18">Spathacanthus hahnianus</emphasis>
</taxonomicName>
Baill, and
<taxonomicName class="Magnoliopsida" family="Bombacaceae" genus="Quararibea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Quararibea funebris" order="Malvales" pageId="3" pageNumber="18" phylum="Magnoliophyta" rank="species" species="funebris">
<emphasis italics="true" pageId="3" pageNumber="18">Quararibea funebris</emphasis>
</taxonomicName>
(La Llave) Vischer.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="18" type="phenology">
<paragraph pageId="3" pageNumber="18">Phenology.</paragraph>
<paragraph pageId="3" pageNumber="18">Mature flowers and fruits were collected in March and April; buds, ripe and immature fruits were observed in the same months.</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="18" type="etymology">
<paragraph pageId="3" pageNumber="18">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="18">
The
<normalizedToken originalValue="specific">specific</normalizedToken>
epithet honors Francisco Lorea
<normalizedToken originalValue="Hernández">Hernandez</normalizedToken>
, in recognition of his many important contributions to our knowledge of the Mexican flora.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18" type="conservation status">
<paragraph pageId="3" pageNumber="18">Conservation status.</paragraph>
<paragraph lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18">
Currently we lack the necessary information to objectively define the conservation status of
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="3" pageNumber="18" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="3" pageNumber="18">Amphitecna loreae</emphasis>
</taxonomicName>
. However, according to the criteria
<pageBreakToken pageId="4" pageNumber="19" start="start">established</pageBreakToken>
by the IUCN, it is possible to tentatively determine that the species is Critically Endangered [CR B1ab (iii)]. Although the only known population of the species is located within a protected natural area,
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna loreae</emphasis>
</taxonomicName>
appears to be rare ecologically and only eight individuals in one hectare of sampling were recorded (
<bibRefCitation author="Escobar-Castellanos, M" journalOrPublisher="Universidad de Ciencias y Artes de Chiapas, Mexico" pageId="7" pageNumber="22" refId="B4" refString="Escobar-Castellanos, M, 2016. Estructura, diversidad y composicion floristica del bosque tropical perennifolio de la zona sujeta a conservacion ecologica La Pera, Berriozabal, Chiapas. Universidad de Ciencias y Artes de Chiapas, Mexico" title="Estructura, diversidad y composicion floristica del bosque tropical perennifolio de la zona sujeta a conservacion ecologica La Pera, Berriozabal, Chiapas" year="2016">Escobar-Castellanos 2016</bibRefCitation>
). The Area of occupancy (AOO) is 12,000 km² and the Extent of occurrence (EOO) is 0.763 km², suggesting a very restricted overall distribution. Furthermore, the tropical rain forest in this region of Chiapas is seriously fragmented and only small remnants persist.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="19" type="materials_examined">
<paragraph pageId="4" pageNumber="19">Additional specimens examined.</paragraph>
<paragraph pageId="4" pageNumber="19">
<materialsCitation collectingDate="2014-05-04" collectingDateMax="2015-05-16" collectingDateMin="2014-05-04" collectorName="M. A. Escobar Castellanos, Y. Licona-Vera" country="Mexico" county="Berriozabal" elevation="1114" latitude="16.864056" location="Joaquin Miguel Gutierrez" longLatPrecision="1" longitude="-93.331024" municipality="Rancho" specimenCount="1" stateProvince="Chiapas">
<collectingCountry name="Mexico">Mexico</collectingCountry>
.
<collectingRegion country="Mexico" name="Chiapas">Chiapas</collectingRegion>
,
<collectingCounty>Berriozabal</collectingCounty>
:
<collectingMunicipality>Rancho</collectingMunicipality>
&quot;El Retiro&quot;,
<normalizedToken originalValue="atrás">atras</normalizedToken>
de el rancho &quot;El Zapote&quot;.
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.3" unit="km" value="13.0">13 km</quantity>
al
<locationDeviation location="Berriozabal">
N de
<normalizedToken originalValue="Berriozábal">Berriozabal</normalizedToken>
</locationDeviation>
camino a
<location LSID="urn:lsid:plazi:treatment:438BEE4825F3B89BB3AE5F5E15021196:5092AD61D5D90121F9ACFF3E38923A32" country="Mexico" county="Berriozabal" latitude="16.864056" longLatPrecision="1" longitude="-93.331024" municipality="Rancho" name="Joaquin Miguel Gutierrez" stateProvince="Chiapas">
<normalizedToken originalValue="Joaquín">Joaquin</normalizedToken>
Miguel
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
</location>
,
<geoCoordinate degrees="16" direction="north" minutes="52" orientation="latitude" precision="1" seconds="09.2" value="16.869223">16°52'09.2&quot;N</geoCoordinate>
,
<geoCoordinate degrees="93" direction="west" minutes="19" orientation="longitude" precision="1" seconds="26.4" value="-93.324">93°19'26.4&quot;W</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.114" unit="m" value="1114.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.114" unit="m" value="1114.0">1114 m</elevation>
</quantity>
.,
<collectingDate value="2014-05-04">04 May 2014</collectingDate>
,
<emphasis italics="true" pageId="4" pageNumber="19">
<collectorName>M. A. Escobar Castellanos</collectorName>
586
</emphasis>
(HEM); same locality,
<emphasis italics="true" pageId="4" pageNumber="19">
<collectorName>M. A. Escobar Castellanos</collectorName>
675
</emphasis>
(HEM); zona sujeta a
<normalizedToken originalValue="protección">proteccion</normalizedToken>
<normalizedToken originalValue="ecológica">ecologica</normalizedToken>
&quot;La Pera&quot;, predio
<normalizedToken originalValue="“Peña">&quot;Pena</normalizedToken>
Flor&quot; camino
<normalizedToken originalValue="Berriozábal-">Berriozabal-</normalizedToken>
Vista Hermosa-El Cairo, km. 12
<normalizedToken originalValue="desvío">desvio</normalizedToken>
al Pozo Turipache,
<geoCoordinate degrees="16" direction="north" minutes="51" orientation="latitude" precision="1" seconds="50.6" value="16.864056">16°51'50.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="93" direction="west" minutes="19" orientation="longitude" precision="1" seconds="51.7" value="-93.331024">93°19'51.7&quot;W</geoCoordinate>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<collectingDate value="2015-05-16">16 May 2015</collectingDate>
,
<emphasis italics="true" pageId="4" pageNumber="19">
<collectorName>Y. Licona-Vera</collectorName>
190
</emphasis>
(XAL)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19" type="discussion">
<paragraph pageId="4" pageNumber="19">Discussion.</paragraph>
<paragraph pageId="4" pageNumber="19">
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna loreae</emphasis>
</taxonomicName>
sp. nov. has a combination of characters that clearly separate it from other species of
<taxonomicName authorityName="Miers" authorityYear="1868" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna</emphasis>
</taxonomicName>
: its strictly cauliflorous inflorescences distinguish it from those species with terminal inflorescences (
<taxonomicName authorityName="A. H. Gentry" authorityYear="1977" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna apiculata" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="apiculata">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna apiculata</emphasis>
</taxonomicName>
A.H. Gentry,
<taxonomicName authorityName="A. H. Gentry" authorityYear="1977" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna breedlovei" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="breedlovei">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna breedlovei</emphasis>
</taxonomicName>
A.H. Gentry,
<taxonomicName authorityName="L. O. Williams" authorityYear="1973" baseAuthorityName="Sprague" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna donnell-smithii" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="donnell-smithii">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna donnell-smithii</emphasis>
</taxonomicName>
(Sprague) L.O. Williams,
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna isthmica" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="isthmica">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna isthmica</emphasis>
</taxonomicName>
(A.H. Gentry) A.H. Gentry,
<taxonomicName authorityName="L. O. Williams" authorityYear="1973" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna molinae" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="molinae">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna molinae</emphasis>
</taxonomicName>
L.O. Williams and
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna steyermarkii" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="steyermarkii">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna steyermarkii</emphasis>
</taxonomicName>
(A.H. Gentry) A.H. Gentry).
</paragraph>
<paragraph lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
The four cauliferous species discussed in the diagnoses have different distribution ranges with different climatic preferences (Figure
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution range and climatic preferences of Amphitecna loreae and related species. Amphitecna latifolia (purple circles) Amphitecna montana (green cross), Amphitecna loreae (black dots) and Amphitecna tuxtlensis (blue squares). In colours similar to those of the species the 95 % confidence ellipses produced by PCA analysis." figureDoi="10.3897/phytokeys.65.8454.figure2" httpUri="https://binary.pensoft.net/fig/92591" pageId="4" pageNumber="19">2</figureCitation>
).
<taxonomicName authorityName="A. H. Gentry" authorityYear="1980" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna tuxtlensis" order="Lamiales" pageId="4" pageNumber="19" phylum="Tracheophyta" rank="species" species="tuxtlensis">
<emphasis italics="true" pageId="4" pageNumber="19">Amphitecna tuxtlensis</emphasis>
</taxonomicName>
has two
<pageBreakToken pageId="5" pageNumber="20" start="start">disjunct</pageBreakToken>
populations in Veracruz, one in the area of the Los Tuxtlas and another in the Uxpanapa-Chimalpas region, where it inhabits the tropical rainforest.
<taxonomicName authorityName="L. O. Williams" authorityYear="1973" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna montana" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="montana">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna montana</emphasis>
</taxonomicName>
is distributed along the Sierra Madre de Chiapas and inhabits the cloud forest above 1200 m.
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna latifolia" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna latifolia</emphasis>
</taxonomicName>
is distributed intermittently in areas near to the Atlantic coast of Mexico, where it inhabits mainly in riparian vegetation and mangrove associations. In contrast,
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna loreae</emphasis>
</taxonomicName>
is endemic to Chiapas and it is known only from a single locality at the municipality of Berriozabal, Chiapas. The species grows on a karstic zone at approximately 900-1,150 m and it inhabits the tropical rainforest (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of diagnostic morphological characters of Amphitecna loreae with its close relatives." httpUri="http://table.plazi.org/id/D623E2964ACA36D18DE86E6D96841B0F" pageId="5" pageNumber="20" tableUuid="D623E2964ACA36D18DE86E6D96841B0F">1</tableCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="980341" doi="10.3897/phytokeys.65.8454.figure2" httpUri="https://binary.pensoft.net/fig/92591" pageId="5" pageNumber="20" start="Figure 2" startId="F2">
<paragraph pageId="5" pageNumber="20">
<emphasis bold="true" pageId="5" pageNumber="20">Figure 2.</emphasis>
Distribution range and climatic preferences of
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna loreae</emphasis>
</taxonomicName>
and related species.
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna latifolia" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna latifolia</emphasis>
</taxonomicName>
(purple circles)
<taxonomicName authorityName="L. O. Williams" authorityYear="1973" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna montana" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="montana">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna montana</emphasis>
</taxonomicName>
(green cross),
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna loreae</emphasis>
</taxonomicName>
(black dots) and
<taxonomicName authorityName="A. H. Gentry" authorityYear="1980" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna tuxtlensis" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="tuxtlensis">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna tuxtlensis</emphasis>
</taxonomicName>
(blue squares). In colours similar to those of the species the 95% confidence ellipses produced by PCA analysis.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="980343" doi="10.3897/phytokeys.65.8454.figure3" httpUri="https://binary.pensoft.net/fig/92592" pageId="5" pageNumber="20" start="Figure 3" startId="F3">
<paragraph pageId="5" pageNumber="20">
<emphasis bold="true" pageId="5" pageNumber="20">Figure 3.</emphasis>
Leaf variation in
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna loreae</emphasis>
</taxonomicName>
and related species.
<emphasis bold="true" pageId="5" pageNumber="20">A</emphasis>
<taxonomicName authorityName="A. H. Gentry" authorityYear="1980" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna tuxtlensis" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="tuxtlensis">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna tuxtlensis</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="5" pageNumber="20">H. Gomez 3710</emphasis>
HEM)
<emphasis bold="true" pageId="5" pageNumber="20">B</emphasis>
<taxonomicName authorityName="L. O. Williams" authorityYear="1973" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna montana" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="montana">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna montana</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="5" pageNumber="20">N. Martinez 927</emphasis>
HEM)
<emphasis bold="true" pageId="5" pageNumber="20">C</emphasis>
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna loreae</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="5" pageNumber="20">M. Escobar 586</emphasis>
HEM) and
<emphasis bold="true" pageId="5" pageNumber="20">D</emphasis>
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna latifolia" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna latifolia</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="5" pageNumber="20">E. Ucan E 251</emphasis>
XAL).
</paragraph>
</caption>
<caption ID-Table-UUID="D623E2964ACA36D18DE86E6D96841B0F" httpUri="http://table.plazi.org/id/D623E2964ACA36D18DE86E6D96841B0F" pageId="5" pageNumber="20" start="Table 1" startId="T1">
<paragraph pageId="5" pageNumber="20">
<emphasis bold="true" pageId="5" pageNumber="20">Table 1.</emphasis>
Comparison of diagnostic morphological characters of
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna loreae</emphasis>
</taxonomicName>
with its close relatives.
</paragraph>
</caption>
<paragraph pageId="5" pageNumber="20">
<table pageId="5" pageNumber="20">
<tr pageId="5" pageNumber="20">
<th colspan="1" pageId="5" pageNumber="20" rowspan="1">Characters</th>
<th colspan="1" pageId="5" pageNumber="20" rowspan="1">
<taxonomicName class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna latifolia" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna latifolia</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="5" pageNumber="20" rowspan="1">
<taxonomicName authorityName="L. O. Williams" authorityYear="1973" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna montana" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="montana">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna montana</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="5" pageNumber="20" rowspan="1">
<taxonomicName authorityName="A. H. Gentry" authorityYear="1980" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna tuxtlensis" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="tuxtlensis">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna tuxtlensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="5" pageNumber="20" rowspan="1">
<taxonomicName authorityName="Ortiz-Rodr. &amp; Burelo" authorityYear="2016" class="Magnoliopsida" family="Bignoniaceae" genus="Amphitecna" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Amphitecna loreae" order="Lamiales" pageId="5" pageNumber="20" phylum="Tracheophyta" rank="species" species="loreae">
<emphasis italics="true" pageId="5" pageNumber="20">Amphitecna loreae</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Habit</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Tree to 10 m tall</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Large tree, 10-20 m tall</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Tree, 5-15 m tall</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Large tree, 10-25 m tall</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Leaf length</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 19 cm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 34 cm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 18 cm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 20 cm</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Leaf width</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 11 cm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 11 cm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 5 cm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 5 cm</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Petiole</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">poorly defined</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">clearly differentiated</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">defined</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">defined</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Leaf shape</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Broadly obovate</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Oblanceolate to narrowly obovate</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Oblanceolate</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Oblanceolate</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Leaf apex</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">rounded to acute, usually apiculate</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">acute to short-acuminate</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">acuminate</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">acuminate</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Length of the flower pedicel</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 36 mm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 100 mm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 26 mm</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">to 60 mm</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Tip of flower buds</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">rounded</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">rounded</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Pointed</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">rounded</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Fruit shape</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">broadly elliptical to spherical</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Oblong-ovoid or ellipsoid</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">ellipsoid</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">broadly elliptical to spherical</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Fruit apex</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">rounded (rare shortly pointed)</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">shortly pointed to acute</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">acute to acuminate</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">rounded</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Habitat</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">always near sea level, mostly in mangrove associations and flooded vegetation</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Mountain cloud forest</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Tropical rain forest</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Tropical rain forest</td>
</tr>
<tr pageId="5" pageNumber="20">
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">
<emphasis bold="true" pageId="5" pageNumber="20">Distribution</emphasis>
</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Mexico (Campeche, Tabasco, Veracruz and Yucatan); Central America, West Indies to Venezuela and Ecuador</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Mexico (Chiapas); Guatemala</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Mexico (Veracruz and Oaxaca)</td>
<td colspan="1" pageId="5" pageNumber="20" rowspan="1">Mexico (Chiapas)</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>