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<document id="88D513819C7ED13D9B1DD993A96297AF" ID-ISSN="1175­5334" ID-ZooBank="C1FA90AF-1C31-45D6-9CB6-C7D3058E501C" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629820076988" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2012" docId="3601F35E9E579336A3F8F92BFE66D87A" docLanguage="en" docName="zt03531p095.pdf" docOrigin="Zootaxa 3531" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Epimetopus hintoni Balfour-Browne 1949" docType="treatment" docVersion="2" lastPageNumber="16" masterDocId="CA388B269E599339A36FFFE4FFC2DA6D" masterDocTitle="3531" masterLastPageNumber="95" masterPageNumber="1" pageNumber="15" updateTime="1699116377101" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:identifier id="362B8E64B77B306531C41152957E39A6" type="ISSN">1175­5334</mods:identifier>
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<treatment id="3601F35E9E579336A3F8F92BFE66D87A" LSID="urn:lsid:plazi:treatment:3601F35E9E579336A3F8F92BFE66D87A" httpUri="http://treatment.plazi.org/id/3601F35E9E579336A3F8F92BFE66D87A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="F6B211C39E579337A3F8F92BFD5DDC84" box="[151,671,1743,1769]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="BE1742489E579337A3F8F92BFD5DDC84" blockId="14.[151,671,1743,1802]" box="[151,671,1743,1769]" pageId="14" pageNumber="15">
<heading id="E55FF5249E579337A3F8F92BFD5DDC84" bold="true" box="[151,671,1743,1769]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="79A839CB9E579337A3F8F92BFD5DDC84" ID-CoL="6FTXN" authority="Balfour-Browne, 1949" authorityName="Balfour-Browne" authorityYear="1949" box="[151,671,1743,1769]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hintoni">
<emphasis id="8CDC9E5A9E579337A3F8F92BFD5DDC84" bold="true" box="[151,671,1743,1769]" pageId="14" pageNumber="15">
<emphasis id="8CDC9E5A9E579337A3F8F92BFE47DC84" bold="true" box="[151,389,1743,1769]" italics="true" pageId="14" pageNumber="15">Epimetopus hintoni</emphasis>
<bibRefCitation id="DA393FB99E579337A2E3F92BFD5DDC84" author="Balfour-Browne, J." box="[396,671,1743,1769]" pageId="14" pageNumber="15" pagination="12 - 18" refId="ref41025" refString="Balfour-Browne, J. (1949) Notes on the subfamily Epimetopinae (Col., Palpicornia) with the description of a new genus and of three new species. Proceedings of the Royal Entomological Society of London, Series B, 18, 12 - 18." type="journal article" year="1949">Balfour-Browne, 1949</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E579337A3F8F916FDB8DD67" box="[151,634,1778,1802]" pageId="14" pageNumber="15" type="description">
<paragraph id="BE1742489E579337A3F8F916FDB8DD67" blockId="14.[151,671,1743,1802]" box="[151,634,1778,1802]" pageId="14" pageNumber="15">
<figureCitation id="26935ECD9E579337A3F8F916FF21DD67" box="[151,227,1778,1802]" captionStart="FIGURES 8–11" captionStartId="12.[151,264,1728,1751]" captionTargetBox="[263,1347,203,1699]" captionTargetId="figure-88@12.[240,1348,195,1707]" captionTargetPageId="12" captionText="FIGURES 8–11. Epimetopus holotypes, dorsal and lateral habitus, and perpendicular to pronotum." pageId="14" pageNumber="15">Figs. 9</figureCitation>
(habitus), 13 (aedeagus), 127 (map)
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E579337A3F8F8DFFDC3DD02" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="BE1742489E579337A3F8F8DFFDB6DD3D" blockId="14.[151,628,1850,1903]" box="[151,628,1850,1872]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="9EB865669E579337A3F8F8DFFDB6DD3D" box="[151,628,1850,1872]" pageId="14" pageNumber="15">
<treatmentCitation id="3F0964599E579337A3F8F8DFFDB2DD3D" author="Balfour-Browne, J." box="[151,624,1850,1872]" page="17" pageId="14" pageNumber="15" year="1949">
<taxonomicName id="79A839CB9E579337A3F8F8DFFDB2DD3D" ID-CoL="6FTXN" authority="Balfour-Browne, 1949: 17" authorityName="Balfour-Browne" authorityPageNumber="17" authorityYear="1949" box="[151,624,1850,1872]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hintoni">
<emphasis id="8CDC9E5A9E579337A3F8F8DFFE9EDD3D" box="[151,348,1850,1872]" italics="true" pageId="14" pageNumber="15">Epimetopus hintoni</emphasis>
<bibRefCitation id="DA393FB99E579337A20DF8DEFDB2DD3D" author="Balfour-Browne, J." box="[354,624,1850,1872]" pageId="14" pageNumber="15" pagination="12 - 18" refId="ref41025" refString="Balfour-Browne, J. (1949) Notes on the subfamily Epimetopinae (Col., Palpicornia) with the description of a new genus and of three new species. Proceedings of the Royal Entomological Society of London, Series B, 18, 12 - 18." type="journal article" year="1949">Balfour-Browne, 1949: 17</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="BE1742489E579337A3F8F8BEFDC3DD02" blockId="14.[151,628,1850,1903]" box="[151,513,1881,1903]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="9EB865669E579337A3F8F8BEFDC3DD02" box="[151,513,1881,1903]" pageId="14" pageNumber="15">
<treatmentCitation id="3F0964599E579337A3F8F8BEFE3FDD02" author="Oliva, A." box="[151,509,1881,1903]" page="8" pageId="14" pageNumber="15" year="1986">
<taxonomicName id="79A839CB9E579337A3F8F8BEFE3FDD02" ID-CoL="6FTXN" authority=": Oliva, 1986: 8" authorityName=": Oliva" authorityPageNumber="8" authorityYear="1986" box="[151,509,1881,1903]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hintoni">
<emphasis id="8CDC9E5A9E579337A3F8F8BEFE9EDD02" box="[151,348,1881,1903]" italics="true" pageId="14" pageNumber="15">Epimetopus hintoni</emphasis>
:
<bibRefCitation id="DA393FB99E579337A206F8BDFE3FDD02" author="Oliva, A." box="[361,509,1881,1903]" pageId="14" pageNumber="15" pagination="1 - 14" refId="ref41547" refString="Oliva, A. (1986) El genero Epimetopus Lacordaire en la Argentina (Coleoptera: Hydrophilidae). Revista de la Societa Entomologica de la Argentina, 44, 1 - 14." type="journal article" year="1986">Oliva, 1986: 8</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E579336A3F8F878FAD9DAB9" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="BE1742489E579337A3F8F878FB4DDDBA" blockId="14.[151,1435,1947,2007]" pageId="14" pageNumber="15">
<emphasis id="8CDC9E5A9E579337A3F8F878FE85DDD9" bold="true" box="[151,327,1947,1972]" pageId="14" pageNumber="15">Type material:</emphasis>
<materialsCitation id="0EC048159E579337A221F878FB4DDDBA" collectingDate="1937-06-12" collectingDateMax="1937-06-13" collectingDateMin="1937-06-12" collectionCode="BMNH" collectorName="H. E. Hinton Collector" country="Bolivia" determinerName="J. Balfour-Browne" location="Type" municipality="Yungas Valley" pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<emphasis id="8CDC9E5A9E579337A221F878FE79DDD9" bold="true" box="[334,443,1948,1972]" pageId="14" pageNumber="15">
<typeStatus id="6113FCEA9E579337A221F878FE79DDD9" box="[334,443,1948,1972]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
(male): &quot;
<location id="BB7714939E579337A14DF878FD9EDDD9" LSID="urn:lsid:plazi:treatment:3601F35E9E579336A3F8F92BFE66D87A:BB7714939E579337A14DF878FD9EDDD9" box="[546,604,1948,1972]" country="Bolivia" municipality="Yungas Valley" name="Type" pageId="14" pageNumber="15">Type</location>
[red disc];
<collectingMunicipality id="5E73D8329E579337A1B2F878FCBFDDD9" box="[733,893,1948,1972]" pageId="14" pageNumber="15">Yungas Valley</collectingMunicipality>
<collectingCountry id="C6BF02D89E579337A0EAF878FC19DDD9" box="[901,987,1948,1972]" name="Bolivia" pageId="14" pageNumber="15">Bolivia</collectingCountry>
.
<date id="CA1664889E579337A08AF878FB48DDD9" box="[997,1162,1948,1972]" pageId="14" pageNumber="15" value="1937-06-12" valueMax="1937-06-13" valueMin="1937-06-12">
<collectingDate id="DA529D609E579337A08AF878FB48DDD9" box="[997,1162,1948,1972]" pageId="14" pageNumber="15" value="1937-06-12" valueMax="1937-06-13" valueMin="1937-06-12">12–13.vi.1937</collectingDate>
</date>
;
<collectorName id="135D279E9E579337A7F9F878FA5ADDD9" box="[1174,1432,1948,1972]" pageId="14" pageNumber="15">H. E. Hinton Collector</collectorName>
;
<taxonomicName id="79A839CB9E579337A3F8F824FEB3DDBA" authorityName=": Oliva" authorityYear="1986" box="[151,369,1983,2007]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hintoni">
<emphasis id="8CDC9E5A9E579337A3F8F824FED9DDBA" box="[151,283,1984,2007]" italics="true" pageId="14" pageNumber="15">Epimetopus</emphasis>
hintoni
</taxonomicName>
Type!
<determinerName id="4BCBAE8C9E579337A2AFF824FD52DDBA" box="[448,656,1983,2007]" pageId="14" pageNumber="15">J. Balfour-Browne</determinerName>
det.; Hinton Coll. B.M. 1939-583&quot; (
<collectionCode id="D8B9DA8D9E579337A747F85BFB40DDBA" box="[1064,1154,1983,2007]" pageId="14" pageNumber="15">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="BE1742489E569336A3A8FF7CFAD9DAB9" blockId="15.[151,1436,151,536]" pageId="15" pageNumber="16">
<emphasis id="8CDC9E5A9E569336A3A8FF7CFE10DADD" bold="true" box="[199,466,151,176]" pageId="15" pageNumber="16">New collection record:</emphasis>
<materialsCitation id="0EC048159E569336A2B6FF73FAD5DAB9" collectingDate="2004-11-20" collectingDateMax="2004-12-12" collectingDateMin="2004-11-20" collectionCode="BMNH" collectorName="Los Volcanes, MV &amp; M. V. L. Barclay &amp; H. Mendel" country="Bolivia" elevation="1000" latitude="-18.1" location="Amboro National Park" longLatPrecision="1276" longitude="-63.6" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Santa Cruz">
<collectingCountry id="C6BF02D89E569336A2B6FF73FDF2DAC2" box="[473,560,151,175]" name="Bolivia" pageId="15" pageNumber="16">Bolivia</collectingCountry>
:
<collectingRegion id="7C6C8CAA9E569336A155FF73FD7ADAC2" box="[570,696,151,175]" country="Bolivia" name="Santa Cruz" pageId="15" pageNumber="16">Santa Cruz</collectingRegion>
,
<location id="BB7714939E569336A1ACFF73FC07DAC2" LSID="urn:lsid:plazi:treatment:3601F35E9E579336A3F8F92BFE66D87A:BB7714939E569336A1ACFF73FC07DAC2" box="[707,965,151,175]" country="Bolivia" latitude="-18.1" longLatPrecision="1276" longitude="-63.6" name="Amboro National Park" pageId="15" pageNumber="16" stateProvince="Santa Cruz">Amboro National Park</location>
,
<collectorName id="135D279E9E569336A0A0FF7CFB62DAC2" box="[975,1184,151,175]" pageId="15" pageNumber="16">Los Volcanes, MV</collectorName>
Light Sheet on stream beach, elev.
<quantity id="7950EFAD9E569336A24EFF58FEBADABE" box="[289,376,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="1000.0">
<elevation id="3585A57B9E569336A24EFF58FEBADABE" box="[289,376,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<geoCoordinate id="DB9C248F9E569336A2EBFF58FE1FDAB9" box="[388,477,188,212]" degrees="18" direction="south" minutes="6" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="-18.1">18° 6' S</geoCoordinate>
,
<geoCoordinate id="DB9C248F9E569336A285FF58FD99DABE" box="[490,603,188,212]" degrees="63" direction="west" minutes="36" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="-63.6">63° 36' W</geoCoordinate>
,
<date id="CA1664889E569336A10AFF58FCEDDAB9" box="[613,815,188,212]" pageId="15" pageNumber="16" value="2004-11-20" valueMax="2004-12-12" valueMin="2004-11-20">
<collectingDate id="DA529D609E569336A10AFF58FCEDDAB9" box="[613,815,188,212]" pageId="15" pageNumber="16" value="2004-11-20" valueMax="2004-12-12" valueMin="2004-11-20">20 xi–12 xii 2004</collectingDate>
</date>
,
<collectorName id="135D279E9E569336A055FF58FC25DAB9" box="[826,999,188,212]" pageId="15" pageNumber="16">M.V.L. Barclay</collectorName>
&amp;
<collectorName id="135D279E9E569336A764FF58FB40DAB9" box="[1035,1154,188,212]" pageId="15" pageNumber="16">H. Mendel</collectorName>
(34
<collectionCode id="D8B9DA8D9E569336A7D9FF58FACDDAB9" box="[1206,1295,188,212]" pageId="15" pageNumber="16">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E569336A3A8FF3BFD3EDBC1" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="BE1742489E569336A3A8FF3BFD3EDBC1" blockId="15.[151,1436,151,536]" pageId="15" pageNumber="16">
<emphasis id="8CDC9E5A9E569336A3A8FF3BFE0CDA95" bold="true" box="[199,462,223,248]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Differential diagnosis.</emphasis>
The habitus of this species (
<figureCitation id="26935ECD9E569336A07DFF3BFC97DA9A" box="[786,853,223,247]" captionStart="FIGURES 8–11" captionStartId="12.[151,264,1728,1751]" captionTargetBox="[263,1347,203,1699]" captionTargetId="figure-88@12.[240,1348,195,1707]" captionTargetPageId="12" captionText="FIGURES 8–11. Epimetopus holotypes, dorsal and lateral habitus, and perpendicular to pronotum." pageId="15" pageNumber="16">Fig. 9</figureCitation>
) is very similar to that of several other members of the
<emphasis id="8CDC9E5A9E569336A3ADFEE0FEE6DB71" box="[194,292,260,284]" italics="true" pageId="15" pageNumber="16">Costatus</emphasis>
group; reliable determinations will be based on examination of the aedeagus (
<figureCitation id="26935ECD9E569336A7C6FEE0FB3CDB71" box="[1193,1278,260,284]" captionStart="FIGURES 12–15" captionStartId="14.[151,264,1641,1664]" captionTargetBox="[175,1414,226,1578]" captionTargetId="figure-73@14.[151,1436,195,1619]" captionTargetPageId="14" captionText="FIGURES 12–15. Epimetopus, aedeagi of holotypes." pageId="15" pageNumber="16">Fig. 13</figureCitation>
). The median lobe is nearly parallel-sided in dorsal view and quite blunt apically; in lateral aspect it is strongly curved. The aedeagus is similar in general shape to that of
<taxonomicName id="79A839CB9E569336A1C6FEA9FC9EDB09" authorityName="Perkins" authorityYear="2012" box="[681,860,332,356]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="8CDC9E5A9E569336A1C6FEA9FC9EDB09" box="[681,860,332,356]" italics="true" pageId="15" pageNumber="16">E. ecuadorensis</emphasis>
</taxonomicName>
(
<figureCitation id="26935ECD9E569336A002FEA8FC03DB09" box="[877,961,332,356]" captionStart="FIGURES 20–23" captionStartId="18.[151,264,1672,1695]" captionTargetBox="[256,1332,191,1643]" captionTargetId="figure-126@18.[227,1360,167,1679]" captionTargetPageId="18" captionText="FIGURES 20–23. Epimetopus, aedeagi of holotypes." pageId="15" pageNumber="16">Fig. 21</figureCitation>
), but differs in shapes of the median lobe and parameres, and curvature of the medial lobe. Externally,
<taxonomicName id="79A839CB9E569336A026FE95FC7ADBE5" authorityName=": Oliva" authorityYear="1986" box="[841,952,368,392]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hintoni">
<emphasis id="8CDC9E5A9E569336A026FE95FC7ADBE5" box="[841,952,368,392]" italics="true" pageId="15" pageNumber="16">E. hintoni</emphasis>
</taxonomicName>
is slightly larger than
<taxonomicName id="79A839CB9E569336A7D7FE95FAA9DBE5" authorityName="Perkins" authorityYear="2012" box="[1208,1387,368,392]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="8CDC9E5A9E569336A7D7FE95FAA9DBE5" box="[1208,1387,368,392]" italics="true" pageId="15" pageNumber="16">E. ecuadorensis</emphasis>
</taxonomicName>
(ca. 1.71 vs.
<quantity id="7950EFAD9E569336A399FE70FE9CDBC1" box="[246,350,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="15" pageNumber="16" unit="mm" value="1.56">1.56 mm</quantity>
), but otherwise they are very similar.
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E569336A3A8FE53FE66D87A" pageId="15" pageNumber="16" type="distribution">
<paragraph id="BE1742489E569336A3A8FE53FE66D87A" blockId="15.[151,1436,151,536]" pageId="15" pageNumber="16">
<emphasis id="8CDC9E5A9E569336A3A8FE53FEA2DBBD" bold="true" box="[199,352,439,464]" inLineHeading="true" pageId="15" pageNumber="16" reason="1">Distribution.</emphasis>
Currently known from two localities in
<collectingCountry id="C6BF02D89E569336A051FE53FC52DBA2" box="[830,912,439,463]" name="Bolivia" pageId="15" pageNumber="16">Bolivia</collectingCountry>
(
<figureCitation id="26935ECD9E569336A0CCFE53FBC8DBA2" box="[931,1034,439,463]" captionStart="FIGURE 127" captionStartId="87.[151,250,1935,1958]" captionTargetBox="[283,1304,402,1914]" captionTargetId="figure-58@87.[283,1304,402,1914]" captionTargetPageId="87" captionText="FIGURE 127. Geographical distributions of Epimetopus species." pageId="15" pageNumber="16">Fig. 127</figureCitation>
).
<bibRefCitation id="DA393FB99E569336A74EFE53FB76DBA2" author="Oliva, A." box="[1057,1204,439,463]" pageId="15" pageNumber="16" pagination="1 - 14" refId="ref41547" refString="Oliva, A. (1986) El genero Epimetopus Lacordaire en la Argentina (Coleoptera: Hydrophilidae). Revista de la Societa Entomologica de la Argentina, 44, 1 - 14." type="journal article" year="1986">Oliva (1986)</bibRefCitation>
reported specimens from
<collectingRegion id="7C6C8CAA9E569336A3BBFE38FE68DB99" box="[212,426,476,500]" country="Argentina" name="Tucuman" pageId="15" pageNumber="16">Tucuman Province</collectingRegion>
,
<collectingCountry id="C6BF02D89E569336A2DAFE38FDEADB99" box="[437,552,476,500]" name="Argentina" pageId="15" pageNumber="16">Argentina</collectingCountry>
; these specimens were not seen during this revision, and therefore could not be confirmed as
<taxonomicName id="79A839CB9E569336A240FDE5FE5FD875" authorityName=": Oliva" authorityYear="1986" box="[303,413,512,536]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hintoni">
<emphasis id="8CDC9E5A9E569336A240FDE5FE5FD875" box="[303,413,512,536]" italics="true" pageId="15" pageNumber="16">E. hintoni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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