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<document id="5DD571BF2ED3BAE1A4D8F1F685562795" ID-ISSN="1175­5334" ID-ZooBank="C1FA90AF-1C31-45D6-9CB6-C7D3058E501C" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629820076988" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2012" docId="3601F35E9E489328A3F8FADEFCBCDDB4" docLanguage="en" docName="zt03531p095.pdf" docOrigin="Zootaxa 3531" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Epimetopus steineri Perkins 2012, new species" docType="treatment" docVersion="2" lastPageNumber="18" masterDocId="CA388B269E599339A36FFFE4FFC2DA6D" masterDocTitle="3531" masterLastPageNumber="95" masterPageNumber="1" pageNumber="18" updateTime="1699116377101" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="00B6359EEBE6F491DFC5346273EE9F01" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:namePart id="AC01624E7376EAA6149120CAE9841089">Perkins, Philip D.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B013A3E20FDC23E65362880727E8E709">text</mods:typeOfResource>
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<mods:part id="71B7329AF28F38D730AD127F91069846">
<mods:date id="3B040B5FCCA51BF6A5A56F8176B325B1">2012</mods:date>
<mods:detail id="227513C759B6DC9871FF42C12DE09E95" type="pubDate">
<mods:number id="9BF781018B4752B9CD70333FCA8C0BFC">2012-10-30</mods:number>
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<mods:number id="AEAC9BE1B307895917D70E5466EDFE8A">3531</mods:number>
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<mods:classification id="ED49BCFF59840C6833C8082281A8636E">journal article</mods:classification>
<mods:identifier id="EA6B0C02C514D6371DE7DC5177FB801E" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="4C4AC7963BEAAF92CBB00AD4B39FB997" type="ZooBank">C1FA90AF-1C31-45D6-9CB6-C7D3058E501C</mods:identifier>
</mods:mods>
<treatment id="3601F35E9E489328A3F8FADEFCBCDDB4" LSID="urn:lsid:plazi:treatment:3601F35E9E489328A3F8FADEFCBCDDB4" httpUri="http://treatment.plazi.org/id/3601F35E9E489328A3F8FADEFCBCDDB4" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="F6B211C39E489328A3F8FADEFDE7DF39" box="[151,549,1338,1364]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="BE1742489E489328A3F8FADEFDE7DF39" blockId="17.[151,648,1338,1397]" box="[151,549,1338,1364]" pageId="17" pageNumber="18">
<heading id="E55FF5249E489328A3F8FADEFDE7DF39" bold="true" box="[151,549,1338,1364]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="8CDC9E5A9E489328A3F8FADEFDE7DF39" bold="true" box="[151,549,1338,1364]" pageId="17" pageNumber="18">
<taxonomicName id="79A839CB9E489328A3F8FADEFE44DF39" ID-CoL="6G5Y2" authority="Perkins, 2012" authorityName="Perkins" authorityYear="2012" box="[151,390,1338,1364]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="steineri" status="sp. nov.">
<emphasis id="8CDC9E5A9E489328A3F8FADEFE44DF39" bold="true" box="[151,390,1338,1364]" italics="true" pageId="17" pageNumber="18">Epimetopus steineri</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="97EF23219E489328A2FAFADEFDE7DF39" box="[405,549,1338,1364]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E489328A3F8FAB9FD4ADF18" box="[151,648,1373,1397]" pageId="17" pageNumber="18" type="description">
<paragraph id="BE1742489E489328A3F8FAB9FD4ADF18" blockId="17.[151,648,1338,1397]" box="[151,648,1373,1397]" pageId="17" pageNumber="18">
<figureCitation id="26935ECD9E489328A3F8FAB9FF33DF18" box="[151,241,1373,1397]" captionStart="FIGURES 16–19" captionStartId="16.[151,264,1728,1751]" captionTargetBox="[252,1331,206,1685]" captionTargetId="figure-102@16.[231,1356,195,1707]" captionTargetPageId="16" captionText="FIGURES 16–19. Epimetopus holotypes, dorsal and lateral habitus, and perpendicular to pronotum." pageId="17" pageNumber="18">Figs. 18</figureCitation>
(habitus), 22 (aedeagus), 128 (map)
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E489328A3F8FA41FA55DC68" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="BE1742489E489328A3F8FA41FA55DC68" blockId="17.[151,1436,1445,2010]" pageId="17" pageNumber="18">
<emphasis id="8CDC9E5A9E489328A3F8FA41FE8BDFD3" bold="true" box="[151,329,1445,1470]" pageId="17" pageNumber="18">Type Material.</emphasis>
<materialsCitation id="0EC048159E489328A23EFA41FA5ADFD0" box="[337,1432,1445,1469]" collectingDate="1977-05-27" collectionCode="USNM" collectorName="W. E. Steiner" country="Ecuador" latitude="0.98333335" location="Tena" longLatPrecision="1308" longitude="-77.816666" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Napo" typeStatus="holotype">
<emphasis id="8CDC9E5A9E489328A23EFA41FE7CDFD0" bold="true" box="[337,446,1445,1469]" pageId="17" pageNumber="18">
<typeStatus id="6113FCEA9E489328A23EFA41FE7CDFD0" box="[337,446,1445,1469]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<emphasis id="8CDC9E5A9E489328A170FA41FD47DFD0" bold="true" box="[543,645,1445,1469]" pageId="17" pageNumber="18">
<collectingCountry id="C6BF02D89E489328A170FA41FD47DFD0" box="[543,645,1445,1469]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
</emphasis>
,
<collectingRegion id="7C6C8CAA9E489328A1FBFA42FD11DFD0" box="[660,723,1446,1469]" country="Ecuador" name="Napo" pageId="17" pageNumber="18">Napo</collectingRegion>
,
<location id="BB7714939E489328A18FFA42FCDBDFD0" LSID="urn:lsid:plazi:treatment:3601F35E9E489328A3F8FADEFCBCDDB4:BB7714939E489328A18FFA42FCDBDFD0" box="[736,793,1446,1469]" country="Ecuador" latitude="0.98333335" longLatPrecision="1308" longitude="-77.816666" name="Tena" pageId="17" pageNumber="18" stateProvince="Napo">Tena</location>
,
<geoCoordinate id="DB9C248F9E489328A04AFA41FC44DFD0" box="[805,902,1445,1469]" degrees="0" direction="north" minutes="59" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="0.98333335">0° 59' N</geoCoordinate>
,
<geoCoordinate id="DB9C248F9E489328A0FDFA41FBC4DFD0" box="[914,1030,1445,1469]" degrees="77" direction="west" minutes="49" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-77.816666">77° 49' W</geoCoordinate>
,
<date id="CA1664889E489328A77DFA41FB44DFD0" box="[1042,1158,1445,1469]" pageId="17" pageNumber="18" value="1977-05-27">
<collectingDate id="DA529D609E489328A77DFA41FB44DFD0" box="[1042,1158,1445,1469]" pageId="17" pageNumber="18" value="1977-05-27">27 v 1977</collectingDate>
</date>
,
<collectorName id="135D279E9E489328A7FCFA42FAE8DFD0" box="[1171,1322,1445,1469]" pageId="17" pageNumber="18">W. E. Steiner</collectorName>
(
<collectionCode id="D8B9DA8D9E489328A656FA41FA52DFD0" box="[1337,1424,1445,1469]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0EC048159E489328A3F8FA2DFC90DF8C" box="[151,850,1481,1506]" collectingDate="1977-05-27" collectionCode="USNM" collectorName="W. E. Steiner" country="Ecuador" latitude="0.98333335" location="Same" longLatPrecision="1308" longitude="-77.816666" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<emphasis id="8CDC9E5A9E489328A3F8FA2DFE04DF8C" bold="true" box="[151,454,1481,1506]" pageId="17" pageNumber="18">
<typeStatus id="6113FCEA9E489328A3F8FA2DFECDDF8C" box="[151,271,1481,1505]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
(18):
<collectingCountry id="C6BF02D89E489328A238FA2DFE00DF8C" box="[343,450,1481,1505]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
:
</emphasis>
<location id="BB7714939E489328A2A1FA2DFDCEDF8C" LSID="urn:lsid:plazi:treatment:3601F35E9E489328A3F8FADEFCBCDDB4:BB7714939E489328A2A1FA2DFDCEDF8C" box="[462,524,1481,1505]" country="Ecuador" name="Same" pageId="17" pageNumber="18">Same</location>
data as holotype (5
<collectionCode id="D8B9DA8D9E489328A19BFA2DFC88DF8C" box="[756,842,1481,1505]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0EC048159E489328A031FA2EFEB4DC68" collectingDate="1977-05-26" collectionCode="USNM" collectorName="W. E. Steiner" country="Ecuador" latitude="0.98333335" location="Tena" longLatPrecision="1308" longitude="-77.816666" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Napo" typeStatus="paratype">
<collectingRegion id="7C6C8CAA9E489328A031FA2EFC5FDF8C" box="[862,925,1482,1505]" country="Ecuador" name="Napo" pageId="17" pageNumber="18">Napo</collectingRegion>
,
<location id="BB7714939E489328A0C5FA2EFC21DF8C" LSID="urn:lsid:plazi:treatment:3601F35E9E489328A3F8FADEFCBCDDB4:BB7714939E489328A0C5FA2EFC21DF8C" box="[938,995,1482,1505]" country="Ecuador" latitude="0.98333335" longLatPrecision="1308" longitude="-77.816666" name="Tena" pageId="17" pageNumber="18" stateProvince="Napo">Tena</location>
,
<geoCoordinate id="DB9C248F9E489328A080FA2DFB92DF8C" box="[1007,1104,1481,1505]" degrees="0" direction="north" minutes="59" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="0.98333335">0° 59' N</geoCoordinate>
,
<geoCoordinate id="DB9C248F9E489328A733FA2DFB12DF8C" box="[1116,1232,1481,1505]" degrees="77" direction="west" minutes="49" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-77.816666">77° 49' W</geoCoordinate>
,
<date id="CA1664889E489328A7B3FA2DFA92DF8C" box="[1244,1360,1481,1505]" pageId="17" pageNumber="18" value="1977-05-26">
<collectingDate id="DA529D609E489328A7B3FA2DFA92DF8C" box="[1244,1360,1481,1505]" pageId="17" pageNumber="18" value="1977-05-26">26 v 1977</collectingDate>
</date>
,
<collectorName id="135D279E9E489328A632FA2EFF24DC68" pageId="17" pageNumber="18">W. E. Steiner</collectorName>
(11
<collectionCode id="D8B9DA8D9E489328A277FA09FEACDC68" box="[280,366,1517,1541]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="0EC048159E489328A2EEFA0AFA51DC68" box="[385,1427,1517,1541]" collectingDate="1977-05-23" collectionCode="USNM" collectorName="P. J. Spangler &amp; D. R. Givens" country="Ecuador" latitude="-1.05" location="Tena" longLatPrecision="1308" longitude="-77.8" pageId="17" pageNumber="18" specimenCount="1" typeStatus="paratype">
<location id="BB7714939E489328A2EEFA0AFE75DC68" LSID="urn:lsid:plazi:treatment:3601F35E9E489328A3F8FADEFCBCDDB4:BB7714939E489328A2EEFA0AFE75DC68" box="[385,439,1518,1541]" country="Ecuador" latitude="-1.05" longLatPrecision="1308" longitude="-77.8" name="Tena" pageId="17" pageNumber="18">Tena</location>
(
<quantity id="7950EFAD9E489328A2A9FA09FDC2DC68" box="[454,512,1517,1541]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="18" unit="km" value="4.0">4 km</quantity>
S),
<geoCoordinate id="DB9C248F9E489328A141FA09FDBBDC68" box="[558,633,1517,1541]" degrees="1" direction="south" minutes="3" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="-1.05">1° 3' S</geoCoordinate>
,
<geoCoordinate id="DB9C248F9E489328A1E9FA09FD35DC68" box="[646,759,1517,1541]" degrees="77" direction="west" minutes="48" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-77.8">77° 48' W</geoCoordinate>
,
<date id="CA1664889E489328A06DFA09FCB1DC68" box="[770,883,1517,1541]" pageId="17" pageNumber="18" value="1977-05-23">
<collectingDate id="DA529D609E489328A06DFA09FCB1DC68" box="[770,883,1517,1541]" pageId="17" pageNumber="18" value="1977-05-23">23 v 1977</collectingDate>
</date>
,
<collectorName id="135D279E9E489328A010FA0AFBD4DC68" box="[895,1046,1517,1541]" pageId="17" pageNumber="18">P. J. Spangler</collectorName>
&amp;
<collectorName id="135D279E9E489328A755FA0AFB0EDC68" box="[1082,1228,1517,1541]" pageId="17" pageNumber="18">D. R. Givens</collectorName>
(#70) (2
<collectionCode id="D8B9DA8D9E489328A65AFA09FA49DC68" box="[1333,1419,1517,1541]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E489328A3A8F9F5FB48DCD4" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="BE1742489E489328A3A8F9F5FB48DCD4" blockId="17.[151,1436,1445,2010]" pageId="17" pageNumber="18">
<emphasis id="8CDC9E5A9E489328A3A8F9F5FE10DC47" bold="true" box="[199,466,1553,1578]" pageId="17" pageNumber="18">Differential Diagnosis.</emphasis>
The habitus of this species (
<figureCitation id="26935ECD9E489328A060F9F5FCA2DC44" box="[783,864,1553,1577]" captionStart="FIGURES 16–19" captionStartId="16.[151,264,1728,1751]" captionTargetBox="[252,1331,206,1685]" captionTargetId="figure-102@16.[231,1356,195,1707]" captionTargetPageId="16" captionText="FIGURES 16–19. Epimetopus holotypes, dorsal and lateral habitus, and perpendicular to pronotum." pageId="17" pageNumber="18">Fig. 18</figureCitation>
) is very similar to that of several other members of the
<emphasis id="8CDC9E5A9E489328A3AEF9D1FEE1DC20" box="[193,291,1589,1613]" italics="true" pageId="17" pageNumber="18">Costatus</emphasis>
group; reliable determinations will be based on examination of the aedeagus. The aedeagus (
<figureCitation id="26935ECD9E489328A62EF9D1FA56DC20" box="[1345,1428,1589,1613]" captionStart="FIGURES 20–23" captionStartId="18.[151,264,1672,1695]" captionTargetBox="[256,1332,191,1643]" captionTargetId="figure-126@18.[227,1360,167,1679]" captionTargetPageId="18" captionText="FIGURES 20–23. Epimetopus, aedeagi of holotypes." pageId="17" pageNumber="18">Fig. 22</figureCitation>
) has a large and distinctively shaped median lobe, widest at about midlength, tapering toward the apex, then widening slightly apically. The internal sac has many fine ridges and spicules. It might be compared with
<taxonomicName id="79A839CB9E489328A6EBF99AFEF8DCD4" authorityName="Perkins" authorityYear="2012" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="transversoides">
<emphasis id="8CDC9E5A9E489328A6EBF99AFEF8DCD4" italics="true" pageId="17" pageNumber="18">E. transversoides</emphasis>
</taxonomicName>
(
<figureCitation id="26935ECD9E489328A226F945FE59DCD4" box="[329,411,1697,1721]" captionStart="FIGURES 12–15" captionStartId="14.[151,264,1641,1664]" captionTargetBox="[175,1414,226,1578]" captionTargetId="figure-73@14.[151,1436,195,1619]" captionTargetPageId="14" captionText="FIGURES 12–15. Epimetopus, aedeagi of holotypes." pageId="17" pageNumber="18">Fig. 15</figureCitation>
), but the median lobe is shaped differently and significantly larger.
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E489328A3A8F921FEB9DDFC" pageId="17" pageNumber="18" type="description">
<paragraph id="BE1742489E489328A3A8F921FEB9DDFC" blockId="17.[151,1436,1445,2010]" pageId="17" pageNumber="18">
<emphasis id="8CDC9E5A9E489328A3A8F921FE9ADCB0" box="[199,344,1733,1758]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">
<emphasis id="8CDC9E5A9E489328A3A8F921FE93DCB3" bold="true" box="[199,337,1733,1758]" pageId="17" pageNumber="18">Description</emphasis>
.
</emphasis>
Size:
<typeStatus id="6113FCEA9E489328A2CAF921FDC5DCB0" box="[421,519,1733,1757]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
(length/width, mm): body (length from anterior margin of pronotum to elytral apices) 2.22/1.69; head (width) 0.44; pronotum 0.61/0.68; elytra 1.11/0.85. Habitus and sculpture as illustrated (
<figureCitation id="26935ECD9E489328A3F0F8E9FF34DD48" box="[159,246,1805,1829]" captionStart="FIGURES 16–19" captionStartId="16.[151,264,1728,1751]" captionTargetBox="[252,1331,206,1685]" captionTargetId="figure-102@16.[231,1356,195,1707]" captionTargetPageId="16" captionText="FIGURES 16–19. Epimetopus holotypes, dorsal and lateral habitus, and perpendicular to pronotum." pageId="17" pageNumber="18">Fig. 18</figureCitation>
). Head black, dorsum red, venter and coxae dark brown, maxillary palpi testaceous. Eye with ca. 3–4 facets between canthus and posterior margin. Elytral punctures round, each with distinct granule at anterior margin, most granules not linking punctures. Protibiae slightly arcuate. Metaventral depression moderately deep, ca. six granules along base.
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E489328A3A8F879FA82DDD8" box="[199,1344,1949,1973]" pageId="17" pageNumber="18" type="etymology">
<paragraph id="BE1742489E489328A3A8F879FA82DDD8" blockId="17.[151,1436,1445,2010]" box="[199,1344,1949,1973]" pageId="17" pageNumber="18">
<emphasis id="8CDC9E5A9E489328A3A8F879FE8CDDD8" bold="true" box="[199,334,1949,1973]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">Etymology.</emphasis>
Named in honor of the expert collector, long-time friend and colleague Warren E. Steiner.
</paragraph>
</subSubSection>
<subSubSection id="F6B211C39E489328A3A8F825FCBCDDB4" box="[199,894,1985,2010]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="BE1742489E489328A3A8F825FCBCDDB4" blockId="17.[151,1436,1445,2010]" box="[199,894,1985,2010]" pageId="17" pageNumber="18">
<emphasis id="8CDC9E5A9E489328A3A8F825FEA2DDB7" bold="true" box="[199,352,1985,2010]" inLineHeading="true" pageId="17" pageNumber="18" reason="1">Distribution.</emphasis>
Currently only known from
<collectingCountry id="C6BF02D89E489328A1CCF825FCC2DDB4" box="[675,768,1985,2009]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
(
<figureCitation id="26935ECD9E489328A060F825FCB3DDB4" box="[783,881,1985,2009]" captionStart="FIGURE 128" captionStartId="88.[151,250,2001,2024]" captionTargetBox="[283,1304,469,1981]" captionTargetId="figure-67@88.[283,1304,469,1981]" captionTargetPageId="88" captionText="FIGURE 128. Geographical distributions of Epimetopus species." pageId="17" pageNumber="18">Fig. 128</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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