161 lines
17 KiB
XML
161 lines
17 KiB
XML
<document id="1032CB7D7EC1BFFA2CAB6854AEF5F5A5" ID-ISSN="11755334" ID-ZooBank="C1FA90AF-1C31-45D6-9CB6-C7D3058E501C" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629820076988" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2012" docId="3601F35E9E09936BA3F8F8E4FBFFDBA2" docLanguage="en" docName="zt03531p095.pdf" docOrigin="Zootaxa 3531" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Epimetopus tridens Perkins 2012, new species" docType="treatment" docVersion="2" lastPageNumber="83" masterDocId="CA388B269E599339A36FFFE4FFC2DA6D" masterDocTitle="3531" masterLastPageNumber="95" masterPageNumber="1" pageNumber="81" updateTime="1699116377101" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F79AEA0F3EC27512C304AFA05570881A">3531</mods:title>
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<mods:roleTerm id="237335BABBC90DE2975201AE565C9322">Author</mods:roleTerm>
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<mods:namePart id="5F1D6B5DD399F6EC7B05BBBF2C67579B">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="1A53D480483F01E3CB36284FFADD8D11">text</mods:typeOfResource>
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<mods:title id="2ABFC742F395FF467D8481CCE22B142F">Zootaxa</mods:title>
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<mods:part id="EFF477164A61483135813C9608B22650">
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<mods:date id="D0E62E517E0316DA850C0920B5C27CC3">2012</mods:date>
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<mods:number id="20F3E0E2010918CD5F5F7C56A6E5CFBD">2012-10-30</mods:number>
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<mods:number id="8019B88CB25F56310EFD1F4E6EA169E5">3531</mods:number>
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<mods:classification id="5ECEA3B81574AF0C840C3DC9E4909744">journal article</mods:classification>
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<mods:identifier id="79E0B4E165C333A73330103E2D55A042" type="ISSN">11755334</mods:identifier>
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</mods:mods>
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<treatment id="3601F35E9E09936BA3F8F8E4FBFFDBA2" LSID="urn:lsid:plazi:treatment:3601F35E9E09936BA3F8F8E4FBFFDBA2" httpUri="http://treatment.plazi.org/id/3601F35E9E09936BA3F8F8E4FBFFDBA2" lastPageId="82" lastPageNumber="83" pageId="80" pageNumber="81">
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<subSubSection id="F6B211C39E099369A3F8F8E4FDDDDD77" box="[151,543,1792,1818]" pageId="80" pageNumber="81" type="nomenclature">
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<paragraph id="BE1742489E099369A3F8F8E4FDDDDD77" blockId="80.[151,675,1792,1851]" box="[151,543,1792,1818]" pageId="80" pageNumber="81">
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<heading id="E55FF5249E099369A3F8F8E4FDDDDD77" bold="true" box="[151,543,1792,1818]" fontSize="11" level="1" pageId="80" pageNumber="81" reason="1">
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<emphasis id="8CDC9E5A9E099369A3F8F8E4FDDDDD77" bold="true" box="[151,543,1792,1818]" pageId="80" pageNumber="81">
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<taxonomicName id="79A839CB9E099369A3F8F8E4FE42DD77" ID-CoL="6FTXG" authority="Perkins, 2012" authorityName="Perkins" authorityYear="2012" box="[151,384,1792,1818]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="tridens" status="sp. nov.">
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<emphasis id="8CDC9E5A9E099369A3F8F8E4FE42DD77" bold="true" box="[151,384,1792,1818]" italics="true" pageId="80" pageNumber="81">Epimetopus tridens</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="97EF23219E099369A2E0F8E4FDDDDD77" box="[399,543,1792,1818]" pageId="80" pageNumber="81" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F6B211C39E099369A3F8F8C7FD61DD56" box="[151,675,1827,1851]" pageId="80" pageNumber="81" type="description">
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<paragraph id="BE1742489E099369A3F8F8C7FD61DD56" blockId="80.[151,675,1792,1851]" box="[151,675,1827,1851]" pageId="80" pageNumber="81">
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<figureCitation id="26935ECD9E099369A3F8F8C7FF3CDD56" box="[151,254,1827,1851]" captionStart="FIGURE 118" captionStartId="81.[151,250,1908,1931]" captionTargetBox="[177,1421,428,1887]" captionTargetId="figure-179@81.[151,1436,413,1893]" captionTargetPageId="81" captionText="FIGURE 118. Epimetopus tridens holotype, dorsal and lateral habitus, and perpendicular to pronotum." pageId="80" pageNumber="81">Figs. 118</figureCitation>
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(habitus), 123 (aedeagus), 126 (map)
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</paragraph>
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</subSubSection>
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<subSubSection id="F6B211C39E099369A3F8F888FB46DDCA" pageId="80" pageNumber="81" type="materials_examined">
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<paragraph id="BE1742489E099369A3F8F888FB46DDCA" blockId="80.[151,1437,1899,2031]" pageId="80" pageNumber="81">
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<emphasis id="8CDC9E5A9E099369A3F8F888FE8ADDE9" bold="true" box="[151,328,1899,1924]" pageId="80" pageNumber="81">Type Material.</emphasis>
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<materialsCitation id="0EC048159E099369A220F888FD45DDCA" collectingDate="1992-10-17" collectionCode="MSP" country="Brazil" latitude="-22.733334" location="Parque do Estado Rio Galharada" longLatPrecision="1258" longitude="-45.6" municipality="Campos do Jordao" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
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<emphasis id="8CDC9E5A9E099369A220F888FE7EDDE9" bold="true" box="[335,444,1900,1924]" pageId="80" pageNumber="81">
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<typeStatus id="6113FCEA9E099369A220F888FE7EDDE9" box="[335,444,1900,1924]" pageId="80" pageNumber="81" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):
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<collectingCountry id="C6BF02D89E099369A173F88FFDA6DDEE" box="[540,612,1899,1923]" name="Brazil" pageId="80" pageNumber="81">Brazil</collectingCountry>
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:
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<collectingRegion id="7C6C8CAA9E099369A11FF88FFD21DDEE" box="[624,739,1899,1923]" country="Brazil" name="Sao Paulo" pageId="80" pageNumber="81">Sao Paulo</collectingRegion>
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,
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<collectingMunicipality id="5E73D8329E099369A180F88FFC01DDEE" box="[751,963,1899,1923]" pageId="80" pageNumber="81">Campos do Jordão</collectingMunicipality>
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,
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<location id="BB7714939E099369A0A0F888FA83DDEE" LSID="urn:lsid:plazi:treatment:3601F35E9E09936BA3F8F8E4FBFFDBA2:BB7714939E099369A0A0F888FA83DDEE" box="[975,1345,1899,1923]" country="Brazil" latitude="-22.733334" longLatPrecision="1258" longitude="-45.6" municipality="Campos do Jordao" name="Parque do Estado Rio Galharada" pageId="80" pageNumber="81" stateProvince="Sao Paulo">Parque do Estado Rio Galharada</location>
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,
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<geoCoordinate id="DB9C248F9E099369A623F88FFF6ADDCA" degrees="22" direction="south" minutes="44" orientation="latitude" pageId="80" pageNumber="81" precision="925" value="-22.733334">22° 44' S</geoCoordinate>
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,
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<geoCoordinate id="DB9C248F9E099369A3DBF86BFEE4DDCA" box="[180,294,1935,1959]" degrees="45" direction="west" minutes="36" orientation="longitude" pageId="80" pageNumber="81" precision="925" value="-45.6">45° 36' W</geoCoordinate>
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,
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<date id="CA1664889E099369A25FF86BFE61DDCA" box="[304,419,1935,1959]" pageId="80" pageNumber="81" value="1992-10-17">
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<collectingDate id="DA529D609E099369A25FF86BFE61DDCA" box="[304,419,1935,1959]" pageId="80" pageNumber="81" value="1992-10-17">17 x 1992</collectingDate>
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</date>
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, Exp. MZSP (
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<collectionCode id="D8B9DA8D9E099369A12DF86BFD42DDCA" box="[578,640,1935,1959]" pageId="80" pageNumber="81">MSP</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="0EC048159E099369A1FDF874FB42DDCA" box="[658,1152,1935,1960]" collectingDate="1992-10-17" collectionCode="MSP" country="Brazil" latitude="-22.733334" location="Parque do Estado Rio Galharada" longLatPrecision="1258" longitude="-45.6" municipality="Campos do Jordao" pageId="80" pageNumber="81" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
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<emphasis id="8CDC9E5A9E099369A1FDF874FCD1DDC5" bold="true" box="[658,787,1936,1960]" pageId="80" pageNumber="81">
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<typeStatus id="6113FCEA9E099369A1FDF874FCCCDDC5" box="[658,782,1936,1960]" pageId="80" pageNumber="81" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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Same data as holotype (3
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<collectionCode id="D8B9DA8D9E099369A754F86BFBBADDCA" box="[1083,1144,1935,1959]" pageId="80" pageNumber="81">MSP</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6B211C39E099368A3A8F857FC5FDB12" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81" type="diagnosis">
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<paragraph id="BE1742489E099368A3A8F857FC5FDB12" blockId="80.[151,1437,1899,2031]" lastBlockId="81.[151,1436,151,383]" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81">
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<emphasis id="8CDC9E5A9E099369A3A8F857FE1BDDA1" bold="true" box="[199,473,1971,1996]" pageId="80" pageNumber="81">Differential Diagnosis.</emphasis>
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This distinctive species (
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<figureCitation id="26935ECD9E099369A07EF857FCB5DDA6" box="[785,887,1971,1995]" captionStart="FIGURE 118" captionStartId="81.[151,250,1908,1931]" captionTargetBox="[177,1421,428,1887]" captionTargetId="figure-179@81.[151,1436,413,1893]" captionTargetPageId="81" captionText="FIGURE 118. Epimetopus tridens holotype, dorsal and lateral habitus, and perpendicular to pronotum." pageId="80" pageNumber="81">Fig. 118</figureCitation>
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) is differentiated from other members of the
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<emphasis id="8CDC9E5A9E099369A3F8F832FEC4DD83" box="[151,262,2006,2030]" italics="true" pageId="80" pageNumber="81">Trogoides</emphasis>
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group by several characters: 1) the longer and more sharply rounded pronotal hood; 2) the anteriorly confluent pronotal hood carinae; 3) the very small, almost absent, secondary pronotal lateral lobes, 4) the relatively deep pronotal depressions; and 5) the comparatively small elytral punctures. All carinae are well developed, and the carinae of the pronotal hood are confluent and raised anteriorly, thereby differing from all other members of the
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<emphasis id="8CDC9E5A9E089368A3F8FEE4FEC4DB75" box="[151,262,256,280]" italics="true" pageId="81" pageNumber="82">Trogoides</emphasis>
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group. Also, males have an upturned clypeus, a character not present in the others species in the group. The aedeagus (
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<figureCitation id="26935ECD9E089368A225FEC6FE72DB57" box="[330,432,290,314]" captionStart="FIGURES 120–123" captionStartId="83.[151,264,1697,1720]" captionTargetBox="[178,1410,182,1665]" captionTargetId="figure-93@83.[164,1424,169,1681]" captionTargetPageId="83" captionText="FIGURES 120–123. Epimetopus, aedeagi of holotypes. 120. E. tuberculatus. 121. E. venezuelensis. 122. E. fimbriatus. 123. E. tridens." pageId="81" pageNumber="82">Fig. 123</figureCitation>
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), which has a very long and slender median lobe, and slender parameres, cannot be confused with any other known
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<taxonomicName id="79A839CB9E089368A17DFEA2FD54DB30" authorityName=": Orchymont" authorityYear="1933" box="[530,662,326,349]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">
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<emphasis id="8CDC9E5A9E089368A17DFEA2FD54DB30" box="[530,662,326,349]" italics="true" pageId="81" pageNumber="82">Epimetopus</emphasis>
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</taxonomicName>
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. It is similar in general plan to that of
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<taxonomicName id="79A839CB9E089368A706FEA2FB3DDB30" authorityName="Perkins" authorityYear="2012" box="[1129,1279,325,349]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="fimbriatus">
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<emphasis id="8CDC9E5A9E089368A706FEA2FB3DDB30" box="[1129,1279,325,349]" italics="true" pageId="81" pageNumber="82">E. fimbriatus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="26935ECD9E089368A67DFEA1FAB5DB30" box="[1298,1399,325,349]" captionStart="FIGURES 120–123" captionStartId="83.[151,264,1697,1720]" captionTargetBox="[178,1410,182,1665]" captionTargetId="figure-93@83.[164,1424,169,1681]" captionTargetPageId="83" captionText="FIGURES 120–123. Epimetopus, aedeagi of holotypes. 120. E. tuberculatus. 121. E. venezuelensis. 122. E. fimbriatus. 123. E. tridens." pageId="81" pageNumber="82">Fig. 122</figureCitation>
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), a species with well developed secondary lateral lobes of the pronotum.
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</paragraph>
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</subSubSection>
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<caption id="EAD712C09E089368A3F8F890FB69DDE6" box="[151,1195,1908,1931]" pageId="81" pageNumber="82" startId="81.[151,250,1908,1931]" targetBox="[177,1421,428,1887]" targetPageId="81">
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||
<paragraph id="BE1742489E089368A3F8F890FB69DDE6" blockId="81.[151,1195,1908,1931]" box="[151,1195,1908,1931]" pageId="81" pageNumber="82">
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<emphasis id="8CDC9E5A9E089368A3F8F890FEE6DDE6" bold="true" box="[151,292,1908,1931]" pageId="81" pageNumber="82">FIGURE 118</emphasis>
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.
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<taxonomicName id="79A839CB9E089368A25EF892FE37DDE6" authorityName="Perkins" authorityYear="2012" box="[305,501,1909,1931]" class="Insecta" family="Epimetopidae" genus="Epimetopus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="tridens">
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<emphasis id="8CDC9E5A9E089368A25EF892FE37DDE6" box="[305,501,1909,1931]" italics="true" pageId="81" pageNumber="82">Epimetopus tridens</emphasis>
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</taxonomicName>
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holotype, dorsal and lateral habitus, and perpendicular to pronotum.
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</paragraph>
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</caption>
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<subSubSection id="F6B211C39E08936BA39AF854FAE4DBEA" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="description">
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||
<paragraph id="BE1742489E08936BA39AF854FAE4DBEA" blockId="81.[151,1436,1968,2029]" lastBlockId="82.[151,1436,151,464]" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
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||
<emphasis id="8CDC9E5A9E089368A39AF854FE44DDA4" box="[245,390,1968,1993]" inLineHeading="true" pageId="81" pageNumber="82" reason="1">
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<emphasis id="8CDC9E5A9E089368A39AF854FEBDDDA4" bold="true" box="[245,383,1968,1993]" pageId="81" pageNumber="82">Description</emphasis>
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.
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</emphasis>
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Size:
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<typeStatus id="6113FCEA9E089368A2A3F855FDECDDA4" box="[460,558,1969,1993]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length from anterior margin of pronotum to elytral apices) 3.16/1.63; head (width) 0.82; pronotum 1.26/1.21; elytra 2.08/1.63. Habitus as illustrated (
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||
<figureCitation id="26935ECD9E089368A784F831FA8FDD80" box="[1259,1357,2005,2029]" captionStart="FIGURE 118" captionStartId="81.[151,250,1908,1931]" captionTargetBox="[177,1421,428,1887]" captionTargetId="figure-179@81.[151,1436,413,1893]" captionTargetPageId="81" captionText="FIGURE 118. Epimetopus tridens holotype, dorsal and lateral habitus, and perpendicular to pronotum." pageId="81" pageNumber="82">Fig. 118</figureCitation>
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). Head dark brown to piceous, maxillary palpi light brown; dorsum dark brown to reddish brown with lighter V-shaped fascia at about midlength of elytra; venter and coxae reddish brown, remainder of legs ochraceous. Head (male) with clypeus upturned. Pronotum with hood carinae straight, confluent and raised anteriorly; secondary lateral lobe very small, almost absent; all ridges well raised, depressions deep. Elytron with four strong granulate carinae, third interrupted anteriorly for distance of ca. 4–5 punctures; punctures round or oval, not large, each with small granule at anterior margin, non-carinate intervals without granules. Protibiae slightly emarginate on inner margin in distal 1/2, outer margin arcuate. Metaventral depression moderately wide, ca. 14 granules along base, shallow.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F6B211C39E0B936BA3A8FE70FB8ADBC1" box="[199,1096,404,428]" pageId="82" pageNumber="83" type="etymology">
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||
<paragraph id="BE1742489E0B936BA3A8FE70FB8ADBC1" blockId="82.[151,1436,151,464]" box="[199,1096,404,428]" pageId="82" pageNumber="83">
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||
<emphasis id="8CDC9E5A9E0B936BA3A8FE70FE8FDBC1" bold="true" box="[199,333,404,428]" inLineHeading="true" pageId="82" pageNumber="83" reason="1">Etymology.</emphasis>
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Named in reference to the trident-like median lobe of the aedeagus.
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</paragraph>
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</subSubSection>
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<subSubSection id="F6B211C39E0B936BA3A8FE53FBFFDBA2" box="[199,1085,439,464]" pageId="82" pageNumber="83" type="distribution">
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||
<paragraph id="BE1742489E0B936BA3A8FE53FBFFDBA2" blockId="82.[151,1436,151,464]" box="[199,1085,439,464]" pageId="82" pageNumber="83">
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||
<emphasis id="8CDC9E5A9E0B936BA3A8FE53FEA2DBBD" bold="true" box="[199,352,439,464]" inLineHeading="true" pageId="82" pageNumber="83" reason="1">Distribution.</emphasis>
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Currently known only from the
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||
<typeStatus id="6113FCEA9E0B936BA1A3FE5CFD3EDBBD" box="[716,764,440,464]" pageId="82" pageNumber="83">type</typeStatus>
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locality in
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<collectingCountry id="C6BF02D89E0B936BA014FE53FC7DDBA2" box="[891,959,439,463]" name="Brazil" pageId="82" pageNumber="83">Brazil</collectingCountry>
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||
(
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||
<figureCitation id="26935ECD9E0B936BA0A1FE53FBF3DBA2" box="[974,1073,439,463]" captionStart="FIGURE 126" captionStartId="86.[151,250,1870,1893]" captionTargetBox="[283,1304,337,1849]" captionTargetId="figure-105@86.[283,1304,337,1849]" captionTargetPageId="86" captionText="FIGURE 126. Geographical distributions of Epimetopus species." pageId="82" pageNumber="83">Fig. 126</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |