268 lines
31 KiB
XML
268 lines
31 KiB
XML
<document ID-CLB-Dataset="23915" ID-DOI="http://dx.doi.org/10.3897/zookeys.913.49044" ID-GBIF-Dataset="76c0fd43-d2e1-49a4-a744-ab443358a3e0" ID-PMC="PMC7044250" ID-Pensoft-Pub="1313-2970-913-89" ID-Pensoft-UUID="0C62A6C200345036B831DC92F02CE385" ID-PubMed="32132850" ID-ZooBank="1007BEAD14F34DCD93FBFA7CA35A042B" ModsDocID="1313-2970-913-89" checkinTime="1582137858263" checkinUser="pensoft" docAuthor="Cumming, Royce T., Bank, Sarah, Le Tirant, Stephane & Bradler, Sven" docDate="2020" docId="082B66391E465B769481AD93FB1891A4" docLanguage="en" docName="ZooKeys 913: 89-126" docOrigin="ZooKeys 913" docSource="http://dx.doi.org/10.3897/zookeys.913.49044" docTitle="Phyllium jacobsoni Rehn & Rehn 1934" docType="treatment" docVersion="6" id="0C62A6C200345036B831DC92F02CE385" lastPageNumber="89" masterDocId="0C62A6C200345036B831DC92F02CE385" masterDocTitle="Notes on the leaf insects of the genus Phyllium of Sumatra and Java, Indonesia, including the description of two new species with purple coxae (Phasmatodea, Phylliidae)" masterLastPageNumber="126" masterPageNumber="89" pageNumber="89" updateTime="1732751135636" updateUser="ExternalLinkService">
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<mods:titleInfo id="07FDCDAEBEDFCD59C6229F56E12378F2">
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<mods:title id="2E29339B4572AA47DAA1710885BBDCE8">Notes on the leaf insects of the genus Phyllium of Sumatra and Java, Indonesia, including the description of two new species with purple coxae (Phasmatodea, Phylliidae)</mods:title>
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<mods:namePart id="C26E082AE9455A62134352E7C1D51198">Cumming, Royce T.</mods:namePart>
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<mods:namePart id="A4B5B377DF1AFF66281C1EE2AEEC00E7">Bank, Sarah</mods:namePart>
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<mods:name id="FA3067201E53F7076EFF82E292F04DA4" type="personal">
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<mods:namePart id="CA9D3ACA8C9257286ED24EDE02E94417">Le Tirant, Stephane</mods:namePart>
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<mods:name id="D3D377DAEBF7BC9A7A1514303611DE67" type="personal">
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<mods:roleTerm id="89DF643B04927BFE013C287AFA2EBFDA">Author</mods:roleTerm>
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<mods:namePart id="BF17506B18A87F2C836B07287736F59B">Bradler, Sven</mods:namePart>
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<mods:typeOfResource id="8B371E5269F368D969FEE34DE3E4ED41">text</mods:typeOfResource>
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<mods:title id="464F8DA61DF226C2F95BA7211D22AACB">ZooKeys</mods:title>
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<mods:date id="4572002A5ABC138C03B28469C08D78B9">2020</mods:date>
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<mods:detail id="DB2385E722830F56489E8C97EB072ED2" type="volume">
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<mods:number id="E2A579DDA5E3D621FD98F6E78B44D2A5">913</mods:number>
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<mods:start id="D0432AC903BA4AA6686A745B7099C2AB">89</mods:start>
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<mods:url id="B5A6A8B731B23C9D607C86B2DCAB4F81">http://dx.doi.org/10.3897/zookeys.913.49044</mods:url>
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<mods:classification id="CF633C38C930BA3B22D0E96C1A6D8416">journal article</mods:classification>
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<mods:identifier id="20C36AB0E537F6A500078FBECCBBCE76" type="DOI">http://dx.doi.org/10.3897/zookeys.913.49044</mods:identifier>
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<mods:identifier id="5FC6E855705DE71EC7C42EFFB43371CE" type="Pensoft-Pub">1313-2970-913-89</mods:identifier>
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<treatment id="082B66391E465B769481AD93FB1891A4" ID-GBIF-Taxon="162174119" LSID="urn:lsid:plazi:treatment:082B66391E465B769481AD93FB1891A4" httpUri="http://treatment.plazi.org/id/082B66391E465B769481AD93FB1891A4" lastPageNumber="89" pageId="0" pageNumber="89" scope_family="Phylliidae" scope_order="Phasmatodea">
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<subSubSection id="C04289BA8CC10F500A4C202CD9AE8248" pageId="0" pageNumber="89" type="nomenclature">
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<paragraph id="BD046DC915969DF20FB98BDF145EDDDE" pageId="0" pageNumber="89">
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<taxonomicName id="771A2B8562CF37DFD14B0D0F5D2F0D24" ID-CoL="8PHLD" LSID="082B6639-1E46-5B76-9481-AD93FB1891A4" authority="Rehn & Rehn, 1934" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">Phyllium (Phyllium) jacobsoni Rehn & Rehn, 1934</taxonomicName>
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<figureCitation id="A7A9E6C665A459BFDA7D30E33AD14288" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phyllium jacobsoni live captive bred individuals, photos courtesy of Bruno Kneubuehler. A Female B male C male ventral view of coxae D female ventral view of coxae E freshly hatched nymph." figureDoi="10.3897/zookeys.913.49044.figure5" httpUri="https://binary.pensoft.net/fig/381029" pageId="0" pageNumber="89">Figure 5A-E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DFCE92F46C3A9D5C3617CE2D72840F27" pageId="0" pageNumber="89" type="distribution">
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<paragraph id="6B524A3A97543B6631430A8565D5DF5B" pageId="0" pageNumber="89">Distribution.</paragraph>
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<paragraph id="D013D905A657DE7AD29FCCD7783A0E7C" pageId="0" pageNumber="89">
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Indonesia: Eastern Java (Nongkodjadjar [type locality]); Eastern Java, Mt. Argopuro; Western Java, Mt. Halimun (Figure
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<figureCitation id="184DB9A19727A0E17E41F61046B43218" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution map for Phyllium giganteum (Borneo, Peninsular Malaysia, Sumatra, and Java), Phyllium hausleithneri (Peninsular Malaysia), Phyllium nisus sp. nov. (Sumatra), and Phyllium gardabagusi sp. nov. (Java) and Phyllium jacobsoni (Java)." figureDoi="10.3897/zookeys.913.49044.figure4" httpUri="https://binary.pensoft.net/fig/381028" pageId="0" pageNumber="89">4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="708A0569F799F85E4266B100C4B6F589" pageId="0" pageNumber="89" type="discussion">
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<paragraph id="B6CCCC2F019EE408A67A66A371455064" pageId="0" pageNumber="89">Discussion.</paragraph>
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<paragraph id="E30A67C75F6CA9DEFD4A28CFDB214E91" pageId="0" pageNumber="89">
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With the description of the very morphologically similar
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<taxonomicName id="122574068A30BA4501E9AB9D9CC47FAC" authorityName="Cumming & Bank & Le Tirant & Bradler" authorityYear="2020" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium gardabagusi" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="gardabagusi">
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<emphasis id="4C757BCC2210B614CA62E38EEC15E937" italics="true" pageId="0" pageNumber="89">Phyllium gardabagusi</emphasis>
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</taxonomicName>
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sp. nov., also from Java, the numerous localities noted within
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<bibRefCitation id="4B2B31EBF28D819E40DBA28BF91449AE" DOI="https://doi.org/10.11646/zootaxa.2322.1.1" author="Hennemann, FH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="89" pagination="1 - 83" refId="B21" refString="Hennemann, FH, Conle, OV, Gottardo, M, Bresseel, J, 2009. On certain species of the genus Phyllium Illiger, 1798, with proposals for an intra-generic systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini). Zootaxa 2322: 1 - 83, DOI: https://doi.org/10.11646/zootaxa.2322.1.1" title="On certain species of the genus Phyllium Illiger, 1798, with proposals for an intra-generic systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini)." url="https://doi.org/10.11646/zootaxa.2322.1.1" volume="2322" year="2009">Hennemann et al. (2009)</bibRefCitation>
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cannot be confirmed as
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<taxonomicName id="AFE70205C88E8E8618EF55496A96CA71" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
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<emphasis id="E521CE64015E0D388FFAB327489DA018" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
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</taxonomicName>
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. This is because the only clear morphological feature to differentiate between these two populations is the coxae color, which can fade in specimens which were not well preserved. Therefore, we only list the localities where we have been able to confirm specimens as
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<taxonomicName id="55C7ACC2CB0F644C31395CE02DC2D364" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
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<emphasis id="27F0A89A887CE67D64F1BAD186A4B195" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
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</taxonomicName>
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. Nevertheless, with this species confirmed from both West and East Java, it is likely the species is found throughout the island.
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</paragraph>
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</subSubSection>
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<subSubSection id="DEE3F5DA9BDBE6EB52AA36A37E4FACBB" pageId="0" pageNumber="89" type="differentiation">
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<paragraph id="1DE332D2D87BDD9C1089728D400F8CCC" pageId="0" pageNumber="89">Differentiation.</paragraph>
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<paragraph id="CBA3D66950ED93DCB863AA9DBB5BEAF1" pageId="0" pageNumber="89">
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This is the only species in the clade that can easily be differentiated morphologically from other clade members. The coxae color in
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<taxonomicName id="B912BC1610BF5C5B8E0F273C0DBC8E85" lsidName="P. jacobsoni" pageId="0" pageNumber="89" rank="species" species="jacobsoni">
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<emphasis id="738353CB1C15B88A0C1CC922312D8A9F" italics="true" pageId="0" pageNumber="89">P. jacobsoni</emphasis>
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</taxonomicName>
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is white (see Figure
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<figureCitation id="30D7A69BCD24044866B61F419226465A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phyllium jacobsoni live captive bred individuals, photos courtesy of Bruno Kneubuehler. A Female B male C male ventral view of coxae D female ventral view of coxae E freshly hatched nymph." figureDoi="10.3897/zookeys.913.49044.figure5" httpUri="https://binary.pensoft.net/fig/381029" pageId="0" pageNumber="89">5C, D</figureCitation>
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) compared to a distinct and dark purple in the other three clade members (Table
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<tableCitation id="B24840A4A254E41A45A32E8FEC4E6FE3" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological features compared between the members of clade B. Key: a As noted in Hennemann et al. (2009). b 75 mm for Phyllium jacobsoni is noted from personal communication with Bruno Kneubuehler who bred P. jacobsoni from the type locality of Nongkodjadjar, East Java. c Gro ̈ sser (2008) lists P. hausleithneri as having 44 - 46 teeth, and Hennemann et al. (2009) note 44 - 48 teeth. d As noted in Cumming et al. (2018). e Only counted on paratype specimens Coll RC 16 - 203 and Coll RC 18 - 418." httpUri="http://table.plazi.org/id/8210CFDE3A5AD3EB26F3ED347064E54E" pageId="0" pageNumber="89" tableUuid="8210CFDE3A5AD3EB26F3ED347064E54E">1</tableCitation>
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). This purple color is easier to view in females and can only be faintly seen on very well preserved or live males. In any discolored male it is impossible to differentiate the species by looking at the coxae because the white and faint purple color do not preserve well and cannot be differentiated on poor quality specimens. It is fortunate that the holotype for
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<taxonomicName id="2AB8ABDFEBBF60151AF3572E189CAF63" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
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<emphasis id="E913BDA834179E9AD7080275957C579E" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
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</taxonomicName>
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is a female specimen as it can clearly be seen on the holotype female that this was not the sympatric species
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<taxonomicName id="474E214D8C1C1302E60D610AAAA3C18C" authorityName="Cumming & Bank & Le Tirant & Bradler" authorityYear="2020" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium gardabagusi" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="gardabagusi">
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<emphasis id="96B87BC97BDB8C677B0F4732FEABF46E" italics="true" pageId="0" pageNumber="89">Phyllium gardabagusi</emphasis>
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</taxonomicName>
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sp. nov. with purple coxae and is instead a specimen with white coxae which defines
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<taxonomicName id="9F6C75E735C4A76E33552958F9607540" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
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<emphasis id="3D0E10BB15292A5F2261000E99D03B23" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="37861BA3E2E50275D3691DAC1E204534" pageId="0" pageNumber="89">
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On average
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<taxonomicName id="E44B21626774F21BD26E301349B7526E" lsidName="P. jacobsoni" pageId="0" pageNumber="89" rank="species" species="jacobsoni">
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<emphasis id="E60C7A43EDFBA76EB71587289750FD55" italics="true" pageId="0" pageNumber="89">P. jacobsoni</emphasis>
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</taxonomicName>
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tend to be smaller individuals, but as with the other species in the clade there were significant outliers that made the range of sizes overlap with the other species significantly (Table
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<tableCitation id="DE57C891B035D7EC7D7B47AC9FCB4904" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological features compared between the members of clade B. Key: a As noted in Hennemann et al. (2009). b 75 mm for Phyllium jacobsoni is noted from personal communication with Bruno Kneubuehler who bred P. jacobsoni from the type locality of Nongkodjadjar, East Java. c Gro ̈ sser (2008) lists P. hausleithneri as having 44 - 46 teeth, and Hennemann et al. (2009) note 44 - 48 teeth. d As noted in Cumming et al. (2018). e Only counted on paratype specimens Coll RC 16 - 203 and Coll RC 18 - 418." httpUri="http://table.plazi.org/id/8210CFDE3A5AD3EB26F3ED347064E54E" pageId="0" pageNumber="89" tableUuid="8210CFDE3A5AD3EB26F3ED347064E54E">1</tableCitation>
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).
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<taxonomicName id="8220071A1B03DEE28015844A2535A9CA" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
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<emphasis id="0CF98248815DA10A8B7E7FFC1D2E341E" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
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</taxonomicName>
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newly hatched nymphs (Figure
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<figureCitation id="43E411FFCBF35F2D21B0873E413501D9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phyllium jacobsoni live captive bred individuals, photos courtesy of Bruno Kneubuehler. A Female B male C male ventral view of coxae D female ventral view of coxae E freshly hatched nymph." figureDoi="10.3897/zookeys.913.49044.figure5" httpUri="https://binary.pensoft.net/fig/381029" pageId="0" pageNumber="89">5E</figureCitation>
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) appear to be the most variable in coloration of the four clade B members.
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<taxonomicName id="3D8F9D96EDB724F874086525E6515AFB" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
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<emphasis id="6120A28F4DD98D566EF6BBDF246C8797" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
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</taxonomicName>
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overall coloration ranges from reddish brown to dark brown (in the other three members dark brown to black), and on
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<taxonomicName id="5CAB37A1FD7D8A5DC45C3DC7963D0B35" lsidName="P. jacobsoni" pageId="0" pageNumber="89" rank="species" species="jacobsoni">
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<emphasis id="CFE5B7783D53AB547BA34F2C4E751029" italics="true" pageId="0" pageNumber="89">P. jacobsoni</emphasis>
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</taxonomicName>
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the white transverse band on the meso- and metafemora can be a solid line (like in the other three clade B species) or can occasionally be a broken white line (but this is a less common form).
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</paragraph>
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<caption id="90D22DB39D6D7784B695281C49750455" doi="10.3897/zookeys.913.49044.figure5" httpUri="https://binary.pensoft.net/fig/381029" pageId="0" pageNumber="89" start="Figure 5" startId="F5">
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<paragraph id="8CB08500FF1BA1D3E4CF0E2EDA7B849C" pageId="0" pageNumber="89">
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<emphasis id="210F949B5176454FA7F5AD9B65458631" bold="true" pageId="0" pageNumber="89">Figure 5.</emphasis>
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<taxonomicName id="0CACE0636493AC4076A13A0B0158B321" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
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<emphasis id="A6DBDB964BCB9B952CF1FC16952830B4" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
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</taxonomicName>
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live captive bred individuals, photos courtesy of Bruno
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<normalizedToken id="F02BC1FB3635220899161F59CF059847" originalValue="Kneubühler">Kneubuehler</normalizedToken>
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.
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<emphasis id="AF3AE35BEE17F2A6F9FA389EF8A19F75" bold="true" pageId="0" pageNumber="89">A</emphasis>
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Female
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<emphasis id="FFDB9B9539C944F0233B73D6F22C7893" bold="true" pageId="0" pageNumber="89">B</emphasis>
|
|
male
|
|
<emphasis id="24DA8B36F18CA8CC88FA7035A7EF34EC" bold="true" pageId="0" pageNumber="89">C</emphasis>
|
|
male ventral view of coxae
|
|
<emphasis id="59A5AF6DEEAC61E2850A359EA174994C" bold="true" pageId="0" pageNumber="89">D</emphasis>
|
|
female ventral view of coxae
|
|
<emphasis id="B81EA6E9EA98F10156408F1D5FB496BE" bold="true" pageId="0" pageNumber="89">E</emphasis>
|
|
freshly hatched nymph.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="8210CFDE3A5AD3EB26F3ED347064E54E" ID-Table-UUID="8210CFDE3A5AD3EB26F3ED347064E54E" httpUri="http://table.plazi.org/id/8210CFDE3A5AD3EB26F3ED347064E54E" pageId="0" pageNumber="89" start="Table 1" startId="T1">
|
|
<paragraph id="11FDE4DE8976B294A848344F1BFC7307" pageId="0" pageNumber="89">
|
|
<emphasis id="CA788B483F60BE0A1AE13B73E5C53EAE" bold="true" pageId="0" pageNumber="89">Table 1.</emphasis>
|
|
Morphological features compared between the members of clade B. Key: a As noted in
|
|
<bibRefCitation id="505464897D1378F53C77B246442C1EE9" DOI="https://doi.org/10.11646/zootaxa.2322.1.1" author="Hennemann, FH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="89" pagination="1 - 83" refId="B21" refString="Hennemann, FH, Conle, OV, Gottardo, M, Bresseel, J, 2009. On certain species of the genus Phyllium Illiger, 1798, with proposals for an intra-generic systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini). Zootaxa 2322: 1 - 83, DOI: https://doi.org/10.11646/zootaxa.2322.1.1" title="On certain species of the genus Phyllium Illiger, 1798, with proposals for an intra-generic systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini)." url="https://doi.org/10.11646/zootaxa.2322.1.1" volume="2322" year="2009">Hennemann et al. (2009)</bibRefCitation>
|
|
. b 75 mm for
|
|
<taxonomicName id="B44EA7BE7405292B2900BF5D3142070C" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
|
|
<emphasis id="687D1F8465F436A8C1824CA64C47655F" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
|
|
</taxonomicName>
|
|
is noted from personal communication with Bruno
|
|
<normalizedToken id="8BADF26B731465207A8AB31D1130BB3E" originalValue="Kneubühler">Kneubuehler</normalizedToken>
|
|
who bred
|
|
<taxonomicName id="A001D47D892252CE9542496490190665" lsidName="P. jacobsoni" pageId="0" pageNumber="89" rank="species" species="jacobsoni">
|
|
<emphasis id="5D5AB565B73C5CE76841F03E22A2CBDB" italics="true" pageId="0" pageNumber="89">P. jacobsoni</emphasis>
|
|
</taxonomicName>
|
|
from the type locality of Nongkodjadjar, East Java. c
|
|
<bibRefCitation id="F70F61A08DDCEBBB9FFEC76D595521AB" author="Gro ̈ sser, D" journalOrPublisher="2 nd Edition, Chimaira, Frankfurt am Main" pageId="0" pageNumber="89" refId="B17" refString="Grösser, D, 2008. Wandelnde Blätter. Ein Katalog aller bisher beschriebenen Phylliinae-Arten und deren Eier mit drei Neubeschreibungen. 2nd Edition, Chimaira, Frankfurt am Main" title="Wandelnde Bla ̈ tter. Ein Katalog aller bisher beschriebenen Phylliinae-Arten und deren Eier mit drei Neubeschreibungen." year="2008">
|
|
Grö
|
|
<normalizedToken id="A06566E7CE086F7424E6A9B6B7F52B6C" originalValue="ßer">sser</normalizedToken>
|
|
(2008)
|
|
</bibRefCitation>
|
|
lists
|
|
<taxonomicName id="4562C8086CEF313890CE69685B8B5BA4" lsidName="P. hausleithneri" pageId="0" pageNumber="89" rank="species" species="hausleithneri">
|
|
<emphasis id="7C988D8EF160DA09F8287006E39AA291" italics="true" pageId="0" pageNumber="89">P. hausleithneri</emphasis>
|
|
</taxonomicName>
|
|
as having 44-46 teeth, and
|
|
<bibRefCitation id="7CABD6164ECA34BB253B2C142A6DE499" DOI="https://doi.org/10.11646/zootaxa.2322.1.1" author="Hennemann, FH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="89" pagination="1 - 83" refId="B21" refString="Hennemann, FH, Conle, OV, Gottardo, M, Bresseel, J, 2009. On certain species of the genus Phyllium Illiger, 1798, with proposals for an intra-generic systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini). Zootaxa 2322: 1 - 83, DOI: https://doi.org/10.11646/zootaxa.2322.1.1" title="On certain species of the genus Phyllium Illiger, 1798, with proposals for an intra-generic systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini)." url="https://doi.org/10.11646/zootaxa.2322.1.1" volume="2322" year="2009">Hennemann et al. (2009)</bibRefCitation>
|
|
note 44-48 teeth. d As noted in
|
|
<bibRefCitation id="3672ABED49851943DAAD51575C870AD5" author="Cumming, RT" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="89" pagination="1 - 9" publicationUrl="https://journals.flvc.org/mundi/article/view/107790" refId="B12" refString="Cumming, RT, Valero Riquelme, P, Teemsma, SN, 2018. Description of Phyllium (Phyllium) conlei, new species, and a first look at the Phylliidae (Phasmatodea) of the Lesser Sunda Islands, Indonesia. Insecta Mundi 0677: 1 - 9, https://journals.flvc.org/mundi/article/view/107790" title="Description of Phyllium (Phyllium) conlei, new species, and a first look at the Phylliidae (Phasmatodea) of the Lesser Sunda Islands, Indonesia." url="https://journals.flvc.org/mundi/article/view/107790" volume="0677" year="2018">Cumming et al. (2018)</bibRefCitation>
|
|
. e Only counted on paratype specimens Coll RC 16-203 and Coll RC 18-418.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="72CD836ADA1F1C44B56BA23CAAEC2465" pageId="0" pageNumber="89">
|
|
<table id="8BE58E956D2D8BEBA956AA37DCE1FB28" pageId="0" pageNumber="89">
|
|
<tr id="B515DA8016916EE34CB8CF2D0C3B69E5" pageId="0" pageNumber="89">
|
|
<th id="FD5B73001D2F24DD548E28F8FD3AFBA9" colspan="1" pageId="0" pageNumber="89" rowspan="1">Feature</th>
|
|
<th id="7FAB597192B7918F9B7976145720900B" colspan="1" pageId="0" pageNumber="89" rowspan="1">
|
|
<taxonomicName id="338174E09F5432DFA35A453FA042629B" authorityName="Rehn & Rehn" authorityYear="1934" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium jacobsoni" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="jacobsoni">
|
|
<emphasis id="9574CED67752F467CF4CB6F850A1EF56" italics="true" pageId="0" pageNumber="89">Phyllium jacobsoni</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<th id="FAA7E2B9E740ED9F93F43BC488D75689" colspan="1" pageId="0" pageNumber="89" rowspan="1">
|
|
<taxonomicName id="C85759E9C3DD479D1B8CE59C39AC623E" authorityName="Brock" authorityYear="1999" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium hausleithneri" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="hausleithneri">
|
|
<emphasis id="76E81EA502E80B0B166FF69EE37CED20" italics="true" pageId="0" pageNumber="89">Phyllium hausleithneri</emphasis>
|
|
</taxonomicName>
|
|
</th>
|
|
<th id="4EDF520012BF1844DD513B57C2281312" colspan="1" pageId="0" pageNumber="89" rowspan="1">
|
|
<taxonomicName id="886E0082D35A6143624425BC48231D2A" authorityName="Cumming & Bank & Le Tirant & Bradler" authorityYear="2020" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium nisus" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="nisus">
|
|
<emphasis id="86D965668A5FCF4AB980337B47E45374" italics="true" pageId="0" pageNumber="89">Phyllium nisus</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</th>
|
|
<th id="C26150F61D3357CC19FC9D510BA31DA8" colspan="1" pageId="0" pageNumber="89" rowspan="1">
|
|
<taxonomicName id="75AEE70C6FC8B95D0AD3DA2A4EDE7E4D" authorityName="Cumming & Bank & Le Tirant & Bradler" authorityYear="2020" class="Insecta" family="Phylliidae" genus="Phyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllium gardabagusi" order="Phasmida" pageId="0" pageNumber="89" phylum="Arthropoda" rank="species" species="gardabagusi">
|
|
<emphasis id="66D4FF7C878D8D3385C0D0A036F09E46" italics="true" pageId="0" pageNumber="89">Phyllium gardabagusi</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</th>
|
|
</tr>
|
|
<tr id="3750505551F2D54865F6C907B334887E" pageId="0" pageNumber="89">
|
|
<td id="2917DAF4DE8EF14192373AFDD9BF2478" colspan="1" pageId="0" pageNumber="89" rowspan="1">Distribution</td>
|
|
<td id="C1A0A129394224C08B526E98FBFD2BD8" colspan="1" pageId="0" pageNumber="89" rowspan="1">Java</td>
|
|
<td id="4534E546B0927680F0DB958D5B9AE841" colspan="1" pageId="0" pageNumber="89" rowspan="1">Peninsular Malaysia</td>
|
|
<td id="75ED24CE7F589928B6EE6CF5F3791D1F" colspan="1" pageId="0" pageNumber="89" rowspan="1">Sumatra</td>
|
|
<td id="9C5E9828AC3D286AE056149F745B3650" colspan="1" pageId="0" pageNumber="89" rowspan="1">Java</td>
|
|
</tr>
|
|
<tr id="43157D4B19ED6301A5ED3246DDE0A01E" pageId="0" pageNumber="89">
|
|
<td id="516CD298A3827094D42BF5C78CF414AC" colspan="1" pageId="0" pageNumber="89" rowspan="1">Female length [mm]</td>
|
|
<td id="9554E93A55E76EA07F3B39F4E1FA5B1F" colspan="1" pageId="0" pageNumber="89" rowspan="1">63.5-75.0a/b</td>
|
|
<td id="56F800B87422F3DC2D222BB9D2686D0E" colspan="1" pageId="0" pageNumber="89" rowspan="1">74.6-82.8</td>
|
|
<td id="446DEF8AF9C475D63CB39791D290371C" colspan="1" pageId="0" pageNumber="89" rowspan="1">70.3-79.3</td>
|
|
<td id="DEDCB6E7BC0300AA9F6F3B9F85CA31C5" colspan="1" pageId="0" pageNumber="89" rowspan="1">68.4-77.3</td>
|
|
</tr>
|
|
<tr id="95BDC2221F1EFD11F38342DA8C43DFA7" pageId="0" pageNumber="89">
|
|
<td id="E8297C3793235538AA875C16971BD6EF" colspan="1" pageId="0" pageNumber="89" rowspan="1">Male length [mm]</td>
|
|
<td id="374ED873B03E250730B7B5EBA4A85BA0" colspan="1" pageId="0" pageNumber="89" rowspan="1">42.5-56.5a</td>
|
|
<td id="708D654DD81046DFF7B086C5E7F2B8E7" colspan="1" pageId="0" pageNumber="89" rowspan="1">55.8-57.8</td>
|
|
<td id="58D582E1B6C879E5DF07AE8867C577AB" colspan="1" pageId="0" pageNumber="89" rowspan="1">51.4-56.1</td>
|
|
<td id="0E2E609FDBFF9367B43C6DA6D2FF4904" colspan="1" pageId="0" pageNumber="89" rowspan="1">50.0-50.5</td>
|
|
</tr>
|
|
<tr id="47A7781BE87B28FD6ACD239460D9FB68" pageId="0" pageNumber="89">
|
|
<td id="F6F9551C7B7565593C8284EAD562198F" colspan="1" pageId="0" pageNumber="89" rowspan="1">
|
|
Number of teeth on the
|
|
<emphasis id="32A69D4D1EFC46001932BFABEB77E276" italics="true" pageId="0" pageNumber="89">pars stridens</emphasis>
|
|
of antennomere III in females
|
|
</td>
|
|
<td id="BA667432F0825456AC86928076CF2207" colspan="1" pageId="0" pageNumber="89" rowspan="1">40a</td>
|
|
<td id="FB1B8A40A7C308B4C79571DA8A011234" colspan="1" pageId="0" pageNumber="89" rowspan="1">44-48c</td>
|
|
<td id="32AF603EEB8E6A752B527449CD90D4A6" colspan="1" pageId="0" pageNumber="89" rowspan="1">37-44d</td>
|
|
<td id="37888A5499112C33881DE583FE684544" colspan="1" pageId="0" pageNumber="89" rowspan="1">34-39e</td>
|
|
</tr>
|
|
<tr id="727C085CBFC177A006080DE33B52DE29" pageId="0" pageNumber="89">
|
|
<td id="CD6E1B6998FED8ADD087F25D624A25DB" colspan="1" pageId="0" pageNumber="89" rowspan="1">Egg color</td>
|
|
<td id="B4005A9AAFFA3BF9EFA98F263FCAA8DA" colspan="1" pageId="0" pageNumber="89" rowspan="1">Tan to medium brown</td>
|
|
<td id="C7042B9FC39DC0AC301F559FBB60CFCC" colspan="1" pageId="0" pageNumber="89" rowspan="1">Dark, rich brown</td>
|
|
<td id="8E902D94A9583242FA8A84387E32FBE6" colspan="1" pageId="0" pageNumber="89" rowspan="1">Pale tan to medium brown</td>
|
|
<td id="49E9970A9D2D5AE1C1F8454EC940335C" colspan="1" pageId="0" pageNumber="89" rowspan="1">Pale brown to medium brown</td>
|
|
</tr>
|
|
<tr id="F7A80271F85B034720806050528E4FEC" pageId="0" pageNumber="89">
|
|
<td id="D3C1A6373D6A616BD050AEA6097611D7" colspan="1" pageId="0" pageNumber="89" rowspan="1">Coxae color</td>
|
|
<td id="D4648041C59BEB4F6EC2741593D7B1D9" colspan="1" pageId="0" pageNumber="89" rowspan="1">White</td>
|
|
<td id="21CC8EA720E5FFB15E2A30B42EEC8CEE" colspan="1" pageId="0" pageNumber="89" rowspan="1">Purple</td>
|
|
<td id="2077C9C6670B9A8F2A7F80A45FE64D03" colspan="1" pageId="0" pageNumber="89" rowspan="1">Purple</td>
|
|
<td id="686F36214D7757B08C1B240F89AA81D1" colspan="1" pageId="0" pageNumber="89" rowspan="1">Purple</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |