147 lines
18 KiB
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147 lines
18 KiB
XML
<document id="13070BCF8F077A837B4651A2F861EE14" ID-DOI="10.5281/zenodo.5392741" ID-ISSN="1638-9387" ID-Zenodo-Dep="5392741" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630317360611" checkinUser="marcus" docAuthor="Seifert, Bernhard" docDate="2000" docId="03BB87B2FF8DF17D4EF8FBC8FEF4FA05" docLanguage="en" docName="Zoosystema.2000.22.3.517-568.pdf" docOrigin="Zoosystema 22 (3)" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Formica mesasiatica Dlussky 1964" docType="treatment" docVersion="4" lastPageNumber="533" masterDocId="FF82FFCAFF82F16D4E19FFECFF83FFC7" masterDocTitle="A taxonomic revision of the ant subgenus Coptoformica Mueller, 1923 (Hymenoptera, Formicidae)" masterLastPageNumber="568" masterPageNumber="517" pageNumber="532" updateTime="1699224723493" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="7B3194F9D14791A910511662D690E63D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="ABA8EC21B05E605FB1B00CCBD1052028">
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<mods:title id="1523158698BF22D21AB1FBAF60407C28">A taxonomic revision of the ant subgenus Coptoformica Mueller, 1923 (Hymenoptera, Formicidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2C6D29E00B6F68F9045989745F1F26D3" type="personal">
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<mods:role id="CB65AEE74C7D86DB2174A80154A84A50">
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<mods:namePart id="3916E17C461CB5B109C69E07E446E20C">Seifert, Bernhard</mods:namePart>
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<mods:affiliation id="2371476F8E2892FC3AD54F8B3213544F">Staatliches Museum für Naturkunde Goerlitz, PSF 300154, D- 02806 Goerlitz (Allemagne).</mods:affiliation>
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<mods:typeOfResource id="5EF23A65302CA3C9929B751D3843F9A1">text</mods:typeOfResource>
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<mods:title id="C7F967F2BC6B4A62FBB00AD1B8F481F2">Zoosystema</mods:title>
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<mods:part id="DD7A51CC5BD5A5F22A8E5B5AF30FDFE3">
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<mods:date id="3B548DBE51EB22A51D27879282E90DE3">2000</mods:date>
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<mods:number id="58BE278EF513F3B783EFACAB582227F4">22</mods:number>
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<mods:detail id="9587EC83A8E1C8117ED3E0A8920C89AE" type="issue">
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<mods:number id="7A6DB0A151BD45CAEACCBDC1FC6AAD9D">3</mods:number>
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<mods:classification id="B5D6991C78A24D1B9979E4AD2C780967">journal article</mods:classification>
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<mods:identifier id="140838E01120A95B73547927D6EC23B9" type="DOI">10.5281/zenodo.5392741</mods:identifier>
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<mods:identifier id="FF2A878250C4B8F6A4719E04FC873273" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="DD95DCDF63E40E1AB9EFC8B734B00DFA" type="Zenodo-Dep">5392741</mods:identifier>
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<treatment id="03BB87B2FF8DF17D4EF8FBC8FEF4FA05" ID-DOI="http://doi.org/10.5281/zenodo.5469119" ID-Zenodo-Dep="5469119" LSID="urn:lsid:plazi:treatment:03BB87B2FF8DF17D4EF8FBC8FEF4FA05" httpUri="http://treatment.plazi.org/id/03BB87B2FF8DF17D4EF8FBC8FEF4FA05" lastPageId="16" lastPageNumber="533" pageId="15" pageNumber="532">
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<subSubSection id="C308652FFF8DF1624EF8FBC8FDE2FBF8" box="[225,609,1060,1087]" pageId="15" pageNumber="532" type="nomenclature">
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<paragraph id="8BAD36A4FF8DF1624EF8FBC8FDE2FBF8" blockId="15.[225,609,1060,1087]" box="[225,609,1060,1087]" pageId="15" pageNumber="532">
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<heading id="D0E581C8FF8DF1624EF8FBC8FDE2FBF8" box="[225,609,1060,1087]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="532" reason="2">
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<taxonomicName id="4C124D27FF8DF1624EF8FBC8FDE2FBF8" authority="Dlussky, 1964" authorityName="Dlussky" authorityYear="1964" box="[225,609,1060,1087]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="532" phylum="Arthropoda" rank="species" species="mesasiatica">
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<emphasis id="B966EAB6FF8DF1624EF8FBC8FE3EFBF9" bold="true" box="[225,445,1060,1086]" italics="true" pageId="15" pageNumber="532">Formica mesasiatica</emphasis>
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Dlussky, 1964
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C308652FFF8DF1624E8EFBB1FE3DFA81" pageId="15" pageNumber="532" type="materials_examined">
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<paragraph id="8BAD36A4FF8DF1624E8EFBB1FEC9FB57" blockId="15.[151,684,1117,1350]" pageId="15" pageNumber="532">
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<typeStatus id="54A98806FF8DF1624E8EFBB1FF49FBB4" box="[151,202,1117,1140]" pageId="15" pageNumber="532">TYPE</typeStatus>
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LOCALITY. — Talasskiy Alatau and Zailiyskiy Alatau, Tyanshan.
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</paragraph>
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<paragraph id="8BAD36A4FF8DF1624E8EFB73FE71FB7F" blockId="15.[151,684,1117,1350]" box="[151,498,1183,1208]" pageId="15" pageNumber="532">
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TYPE MATERIAL. —
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<typeStatus id="54A98806FF8DF1624F41FB73FE2EFB70" box="[344,429,1183,1207]" pageId="15" pageNumber="532" type="paratype">Paratype</typeStatus>
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(
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<collectionCode id="ED03AE61FF8DF1624FA2FB4CFE65FB7F" box="[443,486,1184,1208]" country="0" name="Museum of the Earth, Polish Academy of Sciences" pageId="15" pageNumber="532">MZ</collectionCode>
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).
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</paragraph>
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<paragraph id="8BAD36A4FF8DF1624E8EFB25FE3DFA81" blockId="15.[151,684,1117,1350]" pageId="15" pageNumber="532">
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GEOGRAPHIC ORIGIN OF THE MATERIAL STUDIED. — Numerically evaluated were 11 different samples with
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<specimenCount id="9D14FD2DFF8DF1624E8EFB17FE85FAD4" box="[151,262,1275,1299]" pageId="15" pageNumber="532" type="worker">18 workers</specimenCount>
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and 8 queens from the Tyanshan mountains of S
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<collectingCountry id="F3057634FF8DF1624EE4FAF9FEEEFAEA" box="[253,365,1301,1325]" name="Kazakhstan" pageId="15" pageNumber="532">Kazakhstan</collectingCountry>
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(7) and Kyrghystan (19). Total number of specimens seen 70.
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</paragraph>
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</subSubSection>
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<subSubSection id="C308652FFF8DF17D4E8EFA6BFF79FBE5" lastPageId="16" lastPageNumber="533" pageId="15" pageNumber="532" type="description">
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<paragraph id="8BAD36A4FF8DF1624E8EFA6BFEABFA58" blockId="15.[151,684,1415,1666]" box="[151,296,1415,1441]" pageId="15" pageNumber="532">DESCRIPTION</paragraph>
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<paragraph id="8BAD36A4FF8DF1624E8EFA44FF67FA05" blockId="15.[151,684,1415,1666]" box="[151,228,1448,1474]" pageId="15" pageNumber="532">
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<emphasis id="B966EAB6FF8DF1624E8EFA44FF67FA05" box="[151,228,1448,1474]" italics="true" pageId="15" pageNumber="532">Worker</emphasis>
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</paragraph>
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<paragraph id="8BAD36A4FF8DF1624E8EFA2BFC52FCE5" blockId="15.[151,684,1415,1666]" lastBlockId="15.[714,1247,167,802]" pageId="15" pageNumber="532">
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Very large (CL 1468 ± 61, 1359-1582; CW 1414 ± 61, 1321-1535). Head shape as in
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<taxonomicName id="4C124D27FF8DF1624C7EFA04FD28F9C6" authorityName="Nylander" authorityYear="1846" box="[615,683,1512,1537]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="532" phylum="Arthropoda" rank="species" species="exsecta">
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<emphasis id="B966EAB6FF8DF1624C7EFA04FD28F9C6" box="[615,683,1512,1537]" italics="true" pageId="15" pageNumber="532">exsecta</emphasis>
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</taxonomicName>
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(CL/CW 1.038 ± 0.014, 1.002-1.061). Scape long (SL/CL 1.027 ± 0.019, 0.989-1.059). Clypeus also in median and posterior portions with standing setae (ClySet 4.22 ± 0.43,4-5). Lateral semierect setae in the ocellar triangle always present (OceSet 100%). Eye hairs always strongly developed, often hook-shaped (EyeHL 35.7 ± 4.6, 30-47). Pubescence in the occellar triangle always very dense (sqrtPDF 3.99 ± 0.22, 3.58-4.41). Occipital corners in contrast to the
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<taxonomicName id="4C124D27FF8DF1624CD3FE84FC90FE46" authorityName="Nylander" authorityYear="1846" box="[714,787,360,385]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="532" phylum="Arthropoda" rank="species" species="exsecta">
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<emphasis id="B966EAB6FF8DF1624CD3FE84FC90FE46" box="[714,787,360,385]" italics="true" pageId="15" pageNumber="532">exsecta</emphasis>
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</taxonomicName>
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standard with almost appressed pubescence. Craniad profile of forecoxae with few semierect setae (nCOXA 4.58 ± 0.90, 3.0-5.5). Dorsal pronotum and propodeum occasionally, lateral metapleuron and ventrolateral propodeum always with few standing setae (nMET 3.61 ± 1.26, 1.5-6.0). Outer edge of hind tibial flexor side conspicuously hairy (nHTFL 9.50 ± 0.99, 8.0-11.0), with two size classes of setae, and subdecumbent pubescence. Semierect setae on gaster tergites beginning on the first tergite (TERG 1.00 ± 0.0). Pubescence distance on first gaster tergite very low (sqrtPDG 4.43 ± 0.65, 3.72-6.14).
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</paragraph>
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<paragraph id="8BAD36A4FF8DF1624CD3FCABFC8DFCA6" blockId="15.[714,1247,839,1666]" box="[714,782,839,865]" pageId="15" pageNumber="532">
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<emphasis id="B966EAB6FF8DF1624CD3FCABFC8DFCA6" box="[714,782,839,865]" italics="true" pageId="15" pageNumber="532">Queen</emphasis>
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</paragraph>
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<paragraph id="8BAD36A4FF8DF17D4CD3FC84FE1DFEE5" blockId="15.[714,1247,839,1666]" lastBlockId="16.[151,684,167,290]" lastPageId="16" lastPageNumber="533" pageId="15" pageNumber="532">
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As large as
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<taxonomicName id="4C124D27FF8DF1624D52FC84FC11FC46" authorityName="Nylander" authorityYear="1846" box="[843,914,872,897]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="532" phylum="Arthropoda" rank="species" species="exsecta">
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<emphasis id="B966EAB6FF8DF1624D52FC84FC11FC46" box="[843,914,872,897]" italics="true" pageId="15" pageNumber="532">exsecta</emphasis>
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</taxonomicName>
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(CL 1642 ± 33, 1585-1687; CW 1721 ± 22, 1697-1765; ML 2949 ± 79, 2831-3040). Head broad (CL/CW 0.955 ± 0.023, 0.926-0.985), scape significantly longer than in
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<taxonomicName id="4C124D27FF8DF1624D31FC04FCF2FBC6" authorityName="Nylander" authorityYear="1846" box="[808,881,1000,1025]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="532" phylum="Arthropoda" rank="species" species="exsecta">
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<emphasis id="B966EAB6FF8DF1624D31FC04FCF2FBC6" box="[808,881,1000,1025]" italics="true" pageId="15" pageNumber="532">exsecta</emphasis>
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</taxonomicName>
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(SL/CL 0.994 ± 0.023, 0.957- 1.024). Clypeus also in posterior portions with standing setae. Lateral semierect setae in the ocellar triangle usually present. Eye hairs always long and numerous, often hook-shaped (EyeHL 49.2 ± 6.2, 40-55). Pubescence in the occellar triangle always very dense (sqrtPDF 3.56 ± 0.20, 3.33-3.85). Occipital corners of head with decumbent or subdecumbent hairs (OccHD 38.5 ± 9.3, 31-59); queens with appressed hairs are reported to occur (Dlussky 1967). Brilliance of dorsal head surface low, weakly sculptured surfaces dominate (GLANZ 1.41 ± 0.38, 1.0-2.0). Craniad profile of forecoxae with semierect setae (nCOXA 8.06 ± 1.52, 6.0-11.0). Promesonotum always with standing setae that clearly differ from semierect pubescence (MnHL 195.3 ± 29.2, 152- 233). Outer edge of the hind tibial flexor side conspicuously hairy (nHTFL 9.43 ± 2.87, 6.0- 15.0), with two size classes of setae and subdecumbent pubescence. Semierect setae on gaster tergites always beginning on the first tergite (TERG 1.00 ± 0.00). Pubescence distance on first gaster tergite constantly very low (sqrtPDG 3.56 ± 0.20, 3.33-3.85).
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</paragraph>
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<paragraph id="8BAD36A4FF92F17D4E8EFEABFE56FE98" blockId="16.[151,684,327,1058]" box="[151,469,327,353]" pageId="16" pageNumber="533">TAXONOMIC COMMENTS AND</paragraph>
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<paragraph id="8BAD36A4FF92F17D4E8EFE80FE22FEB8" blockId="16.[151,684,327,1058]" box="[151,417,364,383]" pageId="16" pageNumber="533">DIFFERENTIAL DIAGNOSIS</paragraph>
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<paragraph id="8BAD36A4FF92F17D4E8EFE64FF79FBE5" blockId="16.[151,684,327,1058]" pageId="16" pageNumber="533">
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<taxonomicName id="4C124D27FF92F17D4E8EFE64FEF6FE66" authorityName="Dlussky" authorityYear="1964" box="[151,373,392,417]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="533" phylum="Arthropoda" rank="species" species="mesasiatica">
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<emphasis id="B966EAB6FF92F17D4E8EFE64FEF6FE66" box="[151,373,392,417]" italics="true" pageId="16" pageNumber="533">Formica mesasiatica</emphasis>
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</taxonomicName>
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is so far the only reported
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<taxonomicName id="4C124D27FF92F17D4E8EFE4BFE27FE05" box="[151,420,423,450]" class="Insecta" family="Formicidae" genus="Coptoformica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="533" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B966EAB6FF92F17D4E8EFE4BFEB9FE06" box="[151,314,423,449]" italics="true" pageId="16" pageNumber="533">Coptoformica</emphasis>
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species
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</taxonomicName>
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from Middle Asia (Tyanshan and N Pamir). This population seems to be fully isolated from the Palaearctic range of
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<taxonomicName id="4C124D27FF92F17D4E8EFDE4FF5EFDE6" authorityName="Nylander" authorityYear="1846" box="[151,221,520,545]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="533" phylum="Arthropoda" rank="species" species="exsecta">
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<emphasis id="B966EAB6FF92F17D4E8EFDE4FF5EFDE6" box="[151,221,520,545]" italics="true" pageId="16" pageNumber="533">exsecta</emphasis>
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</taxonomicName>
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. The description presented here is based upon investigation of a
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<typeStatus id="54A98806FF92F17D4F84FDCBFE7AFD86" box="[413,505,551,577]" pageId="16" pageNumber="533" type="paratype">paratype</typeStatus>
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queen stored in MZ, labelled: “Zailijskij Ala-Tau, verch. r. Almatin, 18.7.58” and the investigation of further 10 samples with workers and queens collected by R. Schultz in the Kazakhian and Kyrghysian Tyanshan.
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<taxonomicName id="4C124D27FF92F17D4F09FD24FE22FD26" authorityName="Dlussky" authorityYear="1964" box="[272,417,712,737]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="533" phylum="Arthropoda" rank="species" species="mesasiatica">
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<emphasis id="B966EAB6FF92F17D4F09FD24FE22FD26" box="[272,417,712,737]" italics="true" pageId="16" pageNumber="533">F. mesasiatica</emphasis>
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</taxonomicName>
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shows highly significant (p <0.0001) differences to
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<taxonomicName id="4C124D27FF92F17D4FC9FD04FD95FCC6" authorityName="Nylander" authorityYear="1846" box="[464,534,744,769]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="533" phylum="Arthropoda" rank="species" species="exsecta">
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<emphasis id="B966EAB6FF92F17D4FC9FD04FD95FCC6" box="[464,534,744,769]" italics="true" pageId="16" pageNumber="533">exsecta</emphasis>
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</taxonomicName>
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in sqrtPDG, sqrtPDH, and nCOXA of queens and workers and in SL/CL of queens (
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<tableCitation id="C690031FFF92F17D4FB7FCC4FD8CFC85" box="[430,527,808,834]" captionStart="TABLE" captionStartId="41.[151,162,166,183]" captionText="TABLE 6. — Morphometric data of worker individuals of species with mean EyeHL> 17 µm. Given are arithmetic mean ± standard deviation [lower extreme, upper extreme]; n, number of evaluated specimens." pageId="16" pageNumber="533">Tables 6</tableCitation>
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;
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<tableCitation id="C690031FFF92F17D4C05FCC4FDA8FC85" box="[540,555,808,834]" captionStart="TABLE" captionStartId="45.[151,162,166,183]" captionText="TABLE 8. — Morphometric data of queen individuals of rather large species with mean EyeHL> 20 µm. Given are arithmetic mean ± standard deviation [lower extreme, upper extreme]; n, number of evaluated specimens. The prediction for manchu is derived from queen data given by Agosti (1989) and from correlation analysis of worker and queen data in Coptoformica." pageId="16" pageNumber="533">8</tableCitation>
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). However, it cannot be completely separated from
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<taxonomicName id="4C124D27FF92F17D4C5FFCA4FD09FCA6" authorityName="Nylander" authorityYear="1846" box="[582,650,840,865]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="533" phylum="Arthropoda" rank="species" species="exsecta">
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<emphasis id="B966EAB6FF92F17D4C5FFCA4FD09FCA6" box="[582,650,840,865]" italics="true" pageId="16" pageNumber="533">exsecta</emphasis>
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</taxonomicName>
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by discriminant functions even on the basis of nest sample means. Because of the very peculiar zoogeographic situation
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<taxonomicName id="4C124D27FF92F17D4F62FC44FE6DFC06" authorityName="Dlussky" authorityYear="1964" box="[379,494,936,961]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="533" phylum="Arthropoda" rank="species" species="mesasiatica">
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<emphasis id="B966EAB6FF92F17D4F62FC44FE6DFC06" box="[379,494,936,961]" italics="true" pageId="16" pageNumber="533">mesasiatica</emphasis>
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</taxonomicName>
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is treated here as species. Complete isolation and bottle-necking events could have caused a loss in phenotype diversity.
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</paragraph>
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</subSubSection>
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<subSubSection id="C308652FFF92F17D4E8EFBABFEF4FA05" pageId="16" pageNumber="533" type="biology_ecology">
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<paragraph id="8BAD36A4FF92F17D4E8EFBABFE48FB98" blockId="16.[151,684,1095,1474]" box="[151,459,1095,1121]" pageId="16" pageNumber="533">BIOLOGY AND DISTRIBUTION</paragraph>
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<paragraph id="8BAD36A4FF92F17D4E8EFB84FEF4FA05" blockId="16.[151,684,1095,1474]" pageId="16" pageNumber="533">
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The distribution is apparently limited to the mountain areas of the Tyanshan and N Pamir where the species is found on steppe-like or semidry grasslands and woodland clearings between
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<quantity id="4CEA9B41FF92F17D4E8EFB04FEB1FAC6" box="[151,306,1255,1282]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.7" metricValueMin="1.3" pageId="16" pageNumber="533" unit="m" value="2000.0" valueMax="2700.0" valueMin="1300.0">1300-2700 m</quantity>
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(Dlussky 1967; Tarbinsky 1976; Schultz pers. comm.). The highest densities were observed in grasslands with bushes. Monodomous nests as well as polycalic colonies were observed. Nuptial flights are delayed with growing elevation and were observed between June 29 and July 30.
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</paragraph>
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</subSubSection>
|
|
</treatment>
|
|
</document> |