treatments-xml/data/77/4A/DF/774ADF2BD617FF8AFEA58492FBF4FB15.xml

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<mods:title id="50C84EF3A317CBC3EBB26D0E05DF993B">Damaeus lupus n. sp. (Acarina), a new oribatid mite species from the Carpathian wilderness (Tatra Mountains, Slovakia).</mods:title>
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<heading id="A414D951D617FF86FEA58492F866F9F7" bold="true" box="[292,629,1519,1544]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="280" reason="2">
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<taxonomicName id="38E315BED617FF86FEA58492F827F9F8" ID-CoL="8T8GZ" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[292,564,1519,1544]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="280" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD617FF86FEA58492F827F9F8" bold="true" box="[292,564,1519,1544]" italics="true" pageId="2" pageNumber="280">Damaeus (s. str.) lupus</emphasis>
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<paragraph id="FF5C6E3DD617FF86FF1E8744F96FFAB8" blockId="2.[159,762,1574,1949]" lastBlockId="2.[826,1429,311,1352]" pageId="2" pageNumber="280">
Diagnosis — Large sized (around 
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) 
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<emphasis id="CD97B22FD617FF86FF1E8728FB10F99E" box="[159,259,1610,1633]" italics="true" pageId="2" pageNumber="280">Damaeus</emphasis>
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with distinct costular structures on prodorsum, arched translamellar line (
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) and axially interrupted anterior prodorsal tectum (
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) bearing a distinct tubercle. Rostrum with distinct rostral protuberance (
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). Usual set of prodorsal tubercles (
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) present, well developed. Propodolateral apophysis 
<emphasis id="CD97B22FD617FF86FE9F867EFB3DF8CC" box="[286,302,1820,1843]" italics="true" pageId="2" pageNumber="280">P</emphasis>
present, not particularly large, but still pedotectum-like. Carina 
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anterior to acetabulum I developed as angular or tubercular anterolateral protuberance (
<emphasis id="CD97B22FD617FF86FEFF86E6FB8DF863" box="[382,414,1924,1948]" italics="true" pageId="2" pageNumber="280">alp</emphasis>
), carina 
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on posterior end of propodolateral area angular, sometimes weaker developed. Bothridia shifted anteriad and laterad, far from anterior border of notogaster. Sensillus long, setiform, distally slightly attenuated, with rugose surface. Parastigmatic apophyses 
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and 
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well developed, blunt, of subequal length, both visible in dorsal view, 
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sharply pointed, 
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blunt. Epimeral area with complex set of ridges and tubercles, tubercles 
<emphasis id="CD97B22FD617FF86FB90832DFE29FD98" box="[1041,1082,591,615]" italics="true" pageId="2" pageNumber="280">E2a</emphasis>
and 
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weakly developed, 
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and 
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medium sized, tubercular, 
<emphasis id="CD97B22FD617FF86FA8C8311FF3EFD75" box="[1293,1325,627,650]" italics="true" pageId="2" pageNumber="280">Vp</emphasis>
particularly large. Ventrolateral ridge absent or only indistinct and interrupted, additional two tubercles (
<emphasis id="CD97B22FD617FF86FCC283BEF962FD0C" box="[835,881,732,755]" italics="true" pageId="2" pageNumber="280">VLa</emphasis>
, 
<emphasis id="CD97B22FD617FF86FC0083BEF9B2FD0C" box="[897,929,732,755]" italics="true" pageId="2" pageNumber="280">Ua</emphasis>
) present, 
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shifted laterally and pointing laterad. Anterior border of notogaster with 1-2 arched ridges between spinae adnatae. Spinae adnatae small, triangular, difficult to observe. Distance of insertions 
<emphasis id="CD97B22FD617FF86FBA9820BFE20FC7F" box="[1064,1075,873,896]" italics="true" pageId="2" pageNumber="280">c</emphasis>
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- 
<emphasis id="CD97B22FD617FF86FBC7820BFE42FC7F" box="[1094,1105,873,896]" italics="true" pageId="2" pageNumber="280">c</emphasis>
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subequal to 
<emphasis id="CD97B22FD617FF86FA82820BFF1DFC7F" box="[1283,1294,873,896]" italics="true" pageId="2" pageNumber="280">c</emphasis>
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- 
<emphasis id="CD97B22FD617FF86FAA0820BFF3FFC7F" box="[1313,1324,873,896]" italics="true" pageId="2" pageNumber="280">c</emphasis>
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. Setae 
<emphasis id="CD97B22FD617FF86FCBB82EEF956FC5C" box="[826,837,908,931]" italics="true" pageId="2" pageNumber="280">c</emphasis>
<subScript id="63676C78D617FF86FCC482F6F95DFC5A" attach="left" box="[837,846,916,933]" fontSize="7" pageId="2" pageNumber="280">1</subScript>
directed anteriad, 
<emphasis id="CD97B22FD617FF86FBB782EEFE52FC5C" box="[1078,1089,908,931]" italics="true" pageId="2" pageNumber="280">c</emphasis>
<subScript id="63676C78D617FF86FBC082F6FE59FC5A" attach="left" box="[1089,1098,916,933]" fontSize="7" pageId="2" pageNumber="280">2</subScript>
laterad, both pairs slightly curved and longest of all notogastral setae. Remaining notogastral setae more or less curved and directed laterad, lateroposteriad or posteriad. Legs moderately long, both leg I and leg IV longer than body. Leg setae rather short with few exceptions, seta 
<emphasis id="CD97B22FD617FF86FCF0853FF99CFB8A" box="[881,911,1117,1141]" italics="true" pageId="2" pageNumber="280">ft&quot;</emphasis>
of leg IV the longest, both genual and tibial solenidia particularly short – never longer than half-length of respective leg segment. Larva with very long notogastral setae 
<emphasis id="CD97B22FD617FF86FB0A85A5FEB0FB20" box="[1163,1187,1223,1247]" italics="true" pageId="2" pageNumber="280">c2</emphasis>
, 
<emphasis id="CD97B22FD617FF86FB3485A5FED9FB20" box="[1205,1226,1223,1247]" italics="true" pageId="2" pageNumber="280">lp</emphasis>
and 
<emphasis id="CD97B22FD617FF86FA8B85A5FF36FB20" box="[1290,1317,1223,1247]" italics="true" pageId="2" pageNumber="280">h1</emphasis>
(particularly 
<emphasis id="CD97B22FD617FF86FCF68588F99FFAFD" box="[887,908,1258,1282]" italics="true" pageId="2" pageNumber="280">lp</emphasis>
which is longer than whole body) and with very strong, thick, arched and distinctly rugose setae 
<emphasis id="CD97B22FD617FF86FCE28452F966FAB7" box="[867,885,1328,1352]" italics="true" pageId="2" pageNumber="280">le</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="B7F93DB6D617FF89FCE18407FB91F8B5" lastPageId="13" lastPageNumber="291" pageId="2" pageNumber="280" type="description">
<paragraph id="FF5C6E3DD617FF86FCE18407FF73F9D3" blockId="2.[826,1429,1380,1580]" pageId="2" pageNumber="280">
Description of adult — 
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. 
<emphasis id="CD97B22FD617FF86FA8F8407FF9DFA83" box="[1294,1422,1381,1404]" italics="true" pageId="2" pageNumber="280">Dimensions</emphasis>
. Total body length of 
<typeStatus id="2058D09FD617FF86FBB684EAFE8CFA5F" box="[1079,1183,1416,1440]" pageId="2" pageNumber="280" type="holotype">holotype</typeStatus>
1050 (
<typeStatus id="2058D09FD617FF86FB6F84EAFF70FA5F" box="[1262,1379,1416,1440]" pageId="2" pageNumber="280" type="paratype">paratypes</typeStatus>
950 – 1150), ventral length 980 (830 – 1040), maximum width of notogaster 670 (620 – 670), prodorsum length 345 (330 – 390), maximum prodorsum width 445 (410 – 450). For other measures see 
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.
</paragraph>
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Integument — Body colour dark, almost black, with reddish brown colour tones appearing after maceration in lactic acid. Body surface covered by layer of cerotegument, mostly granular or amorphous. Cerotegument granules fine, subequal in size over whole body including legs except distal part of tarsi and leg setae. Integument under cerotegument with fine punctate microstructure, well visible particularly on posterior part of ventral plate. All studied individuals carried nymphal scalps, with attached fine soil and debris, creating soil particle resembling camouflage cover similarly as in some 
<taxonomicName id="38E315BED616FF87FEA18480FB49FA05" authorityName="von Heyden" authorityYear="1826" box="[288,346,1506,1530]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD616FF87FEA18480FB49FA05" box="[288,346,1506,1530]" italics="true" pageId="3" pageNumber="281">Belba</emphasis>
</taxonomicName>
species.
</paragraph>
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<paragraph id="FF5C6E3DD616FF87FF1E8472F8E5FAA4" blockId="3.[159,1429,1296,1371]" pageId="3" pageNumber="281">
FIGURE 1: 
<taxonomicName id="38E315BED616FF87FE878473FB9EFADC" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[262,397,1296,1315]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="281" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD616FF87FE878473FB9EFADC" box="[262,397,1296,1315]" italics="true" pageId="3" pageNumber="281">Damaeus lupus</emphasis>
</taxonomicName>
<emphasis id="CD97B22FD616FF87FE128472FBDAFADC" bold="true" box="[403,457,1296,1315]" pageId="3" pageNumber="281">
<taxonomicNameLabel id="D6A40F54D616FF87FE128472FBDAFADC" box="[403,457,1296,1315]" pageId="3" pageNumber="281" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(adult, paratype): A – dorsal view, B – ventral view. Acronyms: 
<emphasis id="CD97B22FD616FF87FBB28472FE5EFADC" box="[1075,1101,1296,1315]" italics="true" pageId="3" pageNumber="281">alp</emphasis>
– anterolateral protuberance, 
<emphasis id="CD97B22FD616FF87FAE88472FF96FADC" box="[1385,1413,1296,1315]" italics="true" pageId="3" pageNumber="281">abr</emphasis>
– anterobothridial ridge, 
<emphasis id="CD97B22FD616FF87FE1A844EFBA7FAC0" box="[411,436,1324,1343]" italics="true" pageId="3" pageNumber="281">dis</emphasis>
– discidium, 
<emphasis id="CD97B22FD616FF87FDB3844EF85CFAC0" box="[562,591,1324,1343]" italics="true" pageId="3" pageNumber="281">pbr</emphasis>
– postbothridial ridge, 
<emphasis id="CD97B22FD616FF87FCAA844EF954FAC1" box="[811,839,1324,1342]" italics="true" pageId="3" pageNumber="281">rop</emphasis>
– rostral protuberance, 
<emphasis id="CD97B22FD616FF87FBA4844EFE5AFAC1" box="[1061,1097,1324,1342]" italics="true" pageId="3" pageNumber="281">s. a.</emphasis>
– spina adnatum, 
<emphasis id="CD97B22FD616FF87FB77844EFF10FAC0" box="[1270,1283,1324,1343]" italics="true" pageId="3" pageNumber="281">tl</emphasis>
– translamellar line, 
<emphasis id="CD97B22FD616FF87FF6F842AFB15FAA4" box="[238,262,1352,1371]" italics="true" pageId="3" pageNumber="281">tpf</emphasis>
– tectum of podocephalic fossa (scale bar = 200 µm).
</paragraph>
</caption>
<paragraph id="FF5C6E3DD616FF8DFF448744F884FE01" blockId="3.[159,762,1574,1948]" lastBlockId="9.[159,762,311,511]" lastPageId="9" lastPageNumber="287" pageId="3" pageNumber="281">
Prodorsum — 
<figureCitation id="67D872B8D616FF87FE098744F806F9C1" box="[392,533,1574,1598]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="3" pageNumber="281">Figures 1A</figureCitation>
, 
<figureCitation id="67D872B8D616FF87FDAC8744F87BF9C1" box="[557,616,1574,1598]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="3" pageNumber="281">2A,B</figureCitation>
, 
<figureCitation id="67D872B8D616FF87FDFE8744F89EF9C1" box="[639,653,1574,1598]" captionStart="FIGURE" captionStartId="6.[159,171,1818,1837]" captionTargetBox="[227,1361,349,1785]" captionTargetId="figure-2@6.[227,1361,349,1785]" captionTargetPageId="6" captionText="FIGURE 4: Damaeus lupus n. sp. (adult, paratype): A – lateral view of the body with legs, B – dorsal view of the body, C – sensillus and bothridium, D – spina adnatum, E – rostrum and gnathosoma from lateral view, F – anogenital region, lateral view. Acronyms: bo – bothridium, chel – chelicere, pp – palps, s. ad. – spina adnatum." figureDoi="http://doi.org/10.5281/zenodo.4667412" httpUri="https://zenodo.org/record/4667412/files/figure.png" pageId="3" pageNumber="281">4</figureCitation>
A-C, 4E. Broadly triangular or slightly pentagonal, with rather weakly developed propodolateral apophysis 
<emphasis id="CD97B22FD616FF87FF1E87F2FABCF958" box="[159,175,1680,1703]" italics="true" pageId="3" pageNumber="281">P</emphasis>
, appearing as a blunt perpendicular projection. Carinae 
<emphasis id="CD97B22FD616FF87FE8487D0FB34F935" box="[261,295,1714,1738]" italics="true" pageId="3" pageNumber="281">kpI</emphasis>
and 
<emphasis id="CD97B22FD616FF87FEE987D0FB80F935" box="[360,403,1714,1738]" italics="true" pageId="3" pageNumber="281">kpII</emphasis>
anterior to acetabulum I and II, respectively, well developed, 
<emphasis id="CD97B22FD616FF87FD9487B7F824F912" box="[533,567,1749,1773]" italics="true" pageId="3" pageNumber="281">kpI</emphasis>
forming distinct triangular projection directed anterolaterad (anterolateral protuberance, 
<emphasis id="CD97B22FD616FF87FE168679FBA4F8CC" box="[407,439,1819,1843]" italics="true" pageId="3" pageNumber="281">alp</emphasis>
, 
<figureCitation id="67D872B8D616FF87FE458679F800F8CC" box="[452,531,1819,1843]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="3" pageNumber="281">Fig 1A</figureCitation>
). Rostrum (
<figureCitation id="67D872B8D616FF87FD1B8679FAA3F8A9" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="3" pageNumber="281">Figs 2A, B</figureCitation>
) of complicated structure, rounded in outline, but with prominent rostral protuberance (
<emphasis id="CD97B22FD616FF87FDEB8600F89EF886" box="[618,653,1890,1913]" italics="true" pageId="3" pageNumber="281">rop</emphasis>
) and 1-2 carinae, visible as simple lines (sub-)parallel to rostral margin in dorsal view. Usual set of three pairs of tubercles (
<emphasis id="CD97B22FD616FF87FC5984A2FE94FA28" box="[984,1159,1471,1495]" italics="true" pageId="3" pageNumber="281">Ba, Bp, and Da</emphasis>
) present. Tubercle 
<emphasis id="CD97B22FD616FF87FAF884A2FF86FA28" box="[1401,1429,1472,1495]" italics="true" pageId="3" pageNumber="281">Ba</emphasis>
rather small, placed on posterolateral angle of distinct and strongly developed postbothridial ridge (
<emphasis id="CD97B22FD616FF87FCC2874AF974F9BF" box="[835,871,1576,1600]" italics="true" pageId="3" pageNumber="281">pbr</emphasis>
), quite far behind bothridium and laterally, so that 
<emphasis id="CD97B22FD616FF87FCF58729F98BF99C" box="[884,920,1611,1635]" italics="true" pageId="3" pageNumber="281">pbr</emphasis>
appears almost rectangular curved. Tubercle 
<emphasis id="CD97B22FD616FF87FC0D870DF9B9F979" box="[908,938,1647,1670]" italics="true" pageId="3" pageNumber="281">Bp</emphasis>
largest of prodorsal tubercles (
<figureCitation id="67D872B8D616FF87FAAF870CFF9CF979" box="[1326,1423,1646,1670]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="3" pageNumber="281">Figs 1A</figureCitation>
, 
<figureCitation id="67D872B8D616FF87FCBB87F0F949F955" box="[826,858,1682,1706]" captionStart="FIGURE" captionStartId="6.[159,171,1818,1837]" captionTargetBox="[227,1361,349,1785]" captionTargetId="figure-2@6.[227,1361,349,1785]" captionTargetPageId="6" captionText="FIGURE 4: Damaeus lupus n. sp. (adult, paratype): A – lateral view of the body with legs, B – dorsal view of the body, C – sensillus and bothridium, D – spina adnatum, E – rostrum and gnathosoma from lateral view, F – anogenital region, lateral view. Acronyms: bo – bothridium, chel – chelicere, pp – palps, s. ad. – spina adnatum." figureDoi="http://doi.org/10.5281/zenodo.4667412" httpUri="https://zenodo.org/record/4667412/files/figure.png" pageId="3" pageNumber="281">4B</figureCitation>
), blunt, positioned even slightly more lateral than 
<emphasis id="CD97B22FD616FF87FCF687D7F980F933" box="[887,915,1717,1740]" italics="true" pageId="3" pageNumber="281">Ba</emphasis>
. Tubercle 
<emphasis id="CD97B22FD616FF87FB9287D7FE20F933" box="[1043,1075,1717,1740]" italics="true" pageId="3" pageNumber="281">Da</emphasis>
small but distinct, cuticle anteriad rugose with indistinct longitudinal thickenings, which nevertheless do not create a distinct interbothridial ridge. Tubercles 
<emphasis id="CD97B22FD616FF87FB2A867CFEDEF8CA" box="[1195,1229,1822,1845]" italics="true" pageId="3" pageNumber="281">Dp</emphasis>
absent. Central part of prodorsum elevated, forming flat prodorsal protuberance. Bothridial protuberance anteriorly bordered by distinct and quite long antebothridial ridge (
<emphasis id="CD97B22FD61DFF8CFEB384C1FB47FA44" box="[306,340,1443,1467]" italics="true" pageId="8" pageNumber="286">abr</emphasis>
), distal ends of the two ridges however not reaching each other, leaving narrow gap in the middle of prodorsum. Anteroprodorsal tectum (
<emphasis id="CD97B22FD61DFF8CFF5E876FFB12F9DB" box="[223,257,1549,1572]" italics="true" pageId="8" pageNumber="286">apt</emphasis>
) visible as weakly developed line between protuberances 
<emphasis id="CD97B22FD61DFF8CFECE874DFB7CF9B8" box="[335,367,1583,1607]" italics="true" pageId="8" pageNumber="286">alp</emphasis>
, broadly interrupted in the middle, with strong tubercular projection near protuberance 
<emphasis id="CD97B22FD61DFF8CFE808717FB32F972" box="[257,289,1653,1677]" italics="true" pageId="8" pageNumber="286">alp</emphasis>
on each side. Arched translamellar line (
<emphasis id="CD97B22FD61DFF8CFF2987FAFAABF94F" box="[168,184,1688,1712]" italics="true" pageId="8" pageNumber="286">tl</emphasis>
) well developed. Parastigmatic apophyses visible in dorsal view but rather short, not very prominent. 
<emphasis id="CD97B22FD61DFF8CFF6C87BDFB1BF909" box="[237,264,1759,1782]" italics="true" pageId="8" pageNumber="286">Sa</emphasis>
smaller, spiniform, with acute tip pointing slightly posterolaterad, 
<emphasis id="CD97B22FD61DFF8CFE738660F81DF8E6" box="[498,526,1794,1817]" italics="true" pageId="8" pageNumber="286">Sp</emphasis>
broadly tubercular and blunt, oriented anteriolaterad. Rostral seta (
<emphasis id="CD97B22FD61DFF8CFD5A8647F8E2F8C3" box="[731,753,1829,1852]" italics="true" pageId="8" pageNumber="286">ro</emphasis>
) inserted on small tubercle positioned slightly behind short, oblique rostral costula (
<emphasis id="CD97B22FD61DFF8CFDDA8609F895F87D" box="[603,646,1899,1922]" italics="true" pageId="8" pageNumber="286">rcos</emphasis>
, 
<figureCitation id="67D872B8D61DFF8CFD1B8609F8FDF87C" box="[666,750,1899,1923]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="8" pageNumber="286">Fig 2B</figureCitation>
). Lamellar seta (
<emphasis id="CD97B22FD61DFF8CFC6F84E2FE13FA67" box="[1006,1024,1408,1432]" italics="true" pageId="8" pageNumber="286">le</emphasis>
) inserted on more prominent tubercle at distal end of sclerotised ridge (
<emphasis id="CD97B22FD61DFF8CFA9C84C6FF59FA44" box="[1309,1354,1444,1467]" italics="true" pageId="8" pageNumber="286">acos</emphasis>
, anterior lamellar costula, 
<figureCitation id="67D872B8D61DFF8CFBB884A4FE9FFA21" box="[1081,1164,1478,1502]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="8" pageNumber="286">Fig 2A</figureCitation>
). Prodorsal setae (
<figureCitation id="67D872B8D61DFF8CFAE484A4F997F9FE" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="8" pageNumber="286">Figures 2</figureCitation>
A-C) differ in length, lamellar seta (
<emphasis id="CD97B22FD61DFF8CFACA848BFF4EF9FE" box="[1355,1373,1513,1537]" italics="true" pageId="8" pageNumber="286">le</emphasis>
) being the longest, followed by rostral (
<emphasis id="CD97B22FD61DFF8CFB66876FFEEEF9DB" box="[1255,1277,1549,1572]" italics="true" pageId="8" pageNumber="286">ro</emphasis>
), interlamellar (
<emphasis id="CD97B22FD61DFF8CFCEB8752F993F9B8" box="[874,896,1584,1607]" italics="true" pageId="8" pageNumber="286">in</emphasis>
) and exobothridial (
<emphasis id="CD97B22FD61DFF8CFBF58752FE9FF9B8" box="[1140,1164,1584,1607]" italics="true" pageId="8" pageNumber="286">ex</emphasis>
) setae. All setae more or less curved, setiform, with rather blunt tip, 
<emphasis id="CD97B22FD61DFF8CFA028730FF86F995" box="[1411,1429,1618,1642]" italics="true" pageId="8" pageNumber="286">le</emphasis>
and 
<emphasis id="CD97B22FD61DFF8CFCEE8714F996F972" box="[879,901,1654,1677]" italics="true" pageId="8" pageNumber="286">in</emphasis>
with rough surface. Sensillus (
<figureCitation id="67D872B8D61DFF8CFB7D8717FF5DF972" box="[1276,1358,1653,1677]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="8" pageNumber="286">Fig 2C</figureCitation>
, Plate 1C) long, setiform, hardly attenuated distally, with rough surface, protruding well (by more than half of its length) beyond margin of prodorsum. Bothridium funnel-like, its’ asymmetric expanded rim with blunt anterolateral angle. Bothridium positioned unusually for 
<taxonomicName id="38E315BED61DFF8CFBB0862AFE86F8A0" authorityName="C. L. Koch" authorityYear="1835" box="[1073,1173,1864,1887]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="286" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD61DFF8CFBB0862AFE86F8A0" box="[1073,1173,1864,1887]" italics="true" pageId="8" pageNumber="286">Damaeus</emphasis>
</taxonomicName>
, shifted rather far laterad and slightly anteriad, more or less in the middle of the line connecting tips of protuberance 
<emphasis id="CD97B22FD61CFF8DFD588055F8EAFEB0" box="[729,761,311,335]" italics="true" pageId="9" pageNumber="287">alp</emphasis>
and tubercle 
<emphasis id="CD97B22FD61CFF8DFEB98039FB47FE8D" box="[312,340,347,370]" italics="true" pageId="9" pageNumber="287">Ba</emphasis>
. Three areas with less distinct muscle sigillae present: pair anterior to distal ends of anterolateral ridges and single one axially in interbothridial area between and anteriad to tubercles 
<emphasis id="CD97B22FD61CFF8DFF1E8085FAACFE01" box="[159,191,487,510]" italics="true" pageId="9" pageNumber="287">Da</emphasis>
; sigillae quite small and not very dense.
</paragraph>
<caption id="AB9C3EB5D611FF80FF1E86F3FB25F7EC" ID-DOI="http://doi.org/10.5281/zenodo.4667404" ID-Zenodo-Dep="4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="4" pageNumber="282" startId="4.[159,171,1937,1956]" targetBox="[308,1323,351,1756]" targetPageId="4">
<paragraph id="FF5C6E3DD611FF80FF1E86F3FB25F7EC" blockId="4.[159,1429,1937,1984]" lastBlockId="4.[159,1429,1992,2067]" pageId="4" pageNumber="282">
FIGURE 2: Details of 
<taxonomicName id="38E315BED611FF80FEE886F3FBE3F85B" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[361,496,1937,1956]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="282" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD611FF80FEE886F3FBE3F85B" box="[361,496,1937,1956]" italics="true" pageId="4" pageNumber="282">Damaeus lupus</emphasis>
</taxonomicName>
<emphasis id="CD97B22FD611FF80FE7686F3F83DF85B" bold="true" box="[503,558,1937,1956]" pageId="4" pageNumber="282">
<taxonomicNameLabel id="D6A40F54D611FF80FE7686F3F83DF85B" box="[503,558,1937,1956]" pageId="4" pageNumber="282" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: 
<emphasis id="CD97B22FD611FF80FDCA86AAF874F824" box="[587,615,1992,2011]" italics="true" pageId="4" pageNumber="282">abr</emphasis>
– anterobothridial ridge, 
<emphasis id="CD97B22FD611FF80FCDA86ABF96CF824" box="[859,895,1993,2011]" italics="true" pageId="4" pageNumber="282">acos</emphasis>
– anterior lamellar costula, 
<emphasis id="CD97B22FD611FF80FB0B86ABFEB9F824" box="[1162,1194,1993,2011]" italics="true" pageId="4" pageNumber="282">anx</emphasis>
– insertion of additional anal seta, 
<emphasis id="CD97B22FD611FF80FE9C8686FB23F808" box="[285,304,2020,2039]" italics="true" pageId="4" pageNumber="282">bo</emphasis>
– bothridium, 
<emphasis id="CD97B22FD611FF80FE3C8686FBC9F808" box="[445,474,2020,2039]" italics="true" pageId="4" pageNumber="282">pbr</emphasis>
– postbothridial ridge, 
<emphasis id="CD97B22FD611FF80FD378686F8CBF809" box="[694,728,2020,2038]" italics="true" pageId="4" pageNumber="282">rcos</emphasis>
– rostral costula, 
<emphasis id="CD97B22FD611FF80FCFC8686F98AF809" box="[893,921,2020,2038]" italics="true" pageId="4" pageNumber="282">rop</emphasis>
– rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm).
</paragraph>
</caption>
<caption id="AB9C3EB5D610FF81FF1E8655FE3EF87D" ID-DOI="http://doi.org/10.5281/zenodo.4667408" ID-Zenodo-Dep="4667408" httpUri="https://zenodo.org/record/4667408/files/figure.png" pageId="5" pageNumber="283" startId="5.[159,171,1847,1866]" targetBox="[286,1314,355,1768]" targetPageId="5">
<paragraph id="FF5C6E3DD610FF81FF1E8655FE3EF87D" blockId="5.[159,1429,1847,1922]" pageId="5" pageNumber="283">
FIGURE 3: Legs of 
<taxonomicName id="38E315BED610FF81FED5865AFBC8F8B5" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[340,475,1847,1866]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="283" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD610FF81FED5865AFBC8F8B5" box="[340,475,1847,1866]" italics="true" pageId="5" pageNumber="283">Damaeus lupus</emphasis>
</taxonomicName>
<emphasis id="CD97B22FD610FF81FE638655F809F8B5" bold="true" box="[482,538,1847,1866]" pageId="5" pageNumber="283">
<taxonomicNameLabel id="D6A40F54D610FF81FE638655F809F8B5" box="[482,538,1847,1866]" pageId="5" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(adult, paratype): A – femur and genu of leg I, B – genu, tarsus and tibia of leg I, C – femur and genu of leg II, D – tibia and tarsus of leg II, E – trochanter, femur and genu of leg III, F – genu, tibia and tarsus of leg III, G – trochanter, femur and genu of leg IV, H – genu, tibia and tarsus of leg IV (scale bar = 200 µm).
</paragraph>
</caption>
<caption id="AB9C3EB5D613FF82FF1E8678F918F89B" ID-DOI="http://doi.org/10.5281/zenodo.4667412" ID-Zenodo-Dep="4667412" httpUri="https://zenodo.org/record/4667412/files/figure.png" pageId="6" pageNumber="284" startId="6.[159,171,1818,1837]" targetBox="[227,1361,349,1785]" targetPageId="6">
<paragraph id="FF5C6E3DD613FF82FF1E8678F918F89B" blockId="6.[159,1429,1817,1892]" pageId="6" pageNumber="284">
FIGURE 4: 
<taxonomicName id="38E315BED613FF82FE878678FB9FF8D3" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[262,396,1817,1836]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="284" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD613FF82FE878678FB9FF8D3" box="[262,396,1817,1836]" italics="true" pageId="6" pageNumber="284">Damaeus lupus</emphasis>
</taxonomicName>
<emphasis id="CD97B22FD613FF82FE13867BFBD4F8D3" bold="true" box="[402,455,1817,1836]" pageId="6" pageNumber="284">
<taxonomicNameLabel id="D6A40F54D613FF82FE13867BFBD4F8D3" box="[402,455,1817,1836]" pageId="6" pageNumber="284" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(adult, paratype): A – lateral view of the body with legs, B – dorsal view of the body, C – sensillus and bothridium, D – spina adnatum, E – rostrum and gnathosoma from lateral view, F – anogenital region, lateral view. Acronyms: 
<emphasis id="CD97B22FD613FF82FAF08657FF97F8B7" box="[1393,1412,1845,1864]" italics="true" pageId="6" pageNumber="284">bo</emphasis>
– bothridium, 
<emphasis id="CD97B22FD613FF82FEB68633FB4BF89B" box="[311,344,1873,1892]" italics="true" pageId="6" pageNumber="284">chel</emphasis>
– chelicere, 
<emphasis id="CD97B22FD613FF82FE4A8633FBF3F89C" box="[459,480,1873,1891]" italics="true" pageId="6" pageNumber="284">pp</emphasis>
– palps, 
<emphasis id="CD97B22FD613FF82FDB28633F871F89B" box="[563,610,1873,1892]" italics="true" pageId="6" pageNumber="284">s. ad.</emphasis>
– spina adnatum.
</paragraph>
</caption>
<caption id="AB9C3EB5D612FF83FF1E8670FAEEF8BE" ID-DOI="http://doi.org/10.5281/zenodo.4667414" ID-Zenodo-Dep="4667414" httpUri="https://zenodo.org/record/4667414/files/figure.png" pageId="7" pageNumber="285" startId="7.[159,171,1810,1829]" targetBox="[227,1361,385,1778]" targetPageId="7">
<paragraph id="FF5C6E3DD612FF83FF1E8670FAEEF8BE" blockId="7.[159,1429,1809,1857]" pageId="7" pageNumber="285">
FIGURE 5: 
<taxonomicName id="38E315BED612FF83FE848670FB99F8DB" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[261,394,1809,1828]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="285" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD612FF83FE848670FB99F8DB" box="[261,394,1809,1828]" italics="true" pageId="7" pageNumber="285">Damaeus lupus</emphasis>
</taxonomicName>
<emphasis id="CD97B22FD612FF83FE118670FBD7F8DA" bold="true" box="[400,452,1810,1829]" pageId="7" pageNumber="285">
<taxonomicNameLabel id="D6A40F54D612FF83FE118670FBD7F8DA" box="[400,452,1810,1829]" pageId="7" pageNumber="285" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(adult, paratype): A – genu and tibia of leg I, B – tarsus of leg I, C – tibia and tarsus of leg II, D – tibia of leg IV.
</paragraph>
</caption>
<caption id="AB9C3EB5D61DFF8CFF1E8056F891FEB9" box="[159,642,307,327]" pageId="8" pageNumber="286">
<paragraph id="FF5C6E3DD61DFF8CFF1E8056F891FEB9" blockId="8.[159,1087,307,354]" box="[159,642,307,327]" pageId="8" pageNumber="286">
TABLE 1: Selected measures of 
<taxonomicName id="38E315BED61DFF8CFE438056F85BFEB9" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[450,584,307,326]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="286" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD61DFF8CFE438056F85BFEB9" box="[450,584,307,326]" italics="true" pageId="8" pageNumber="286">Damaeus lupus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6A40F54D61DFF8CFDCF8051F891FEB9" box="[590,642,307,326]" pageId="8" pageNumber="286" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="FF5C6E3DD61DFF8CFF40802DFE2CFE9D" blockId="8.[159,1087,307,354]" box="[193,1087,335,354]" pageId="8" pageNumber="286">
Values correspond to 
<typeStatus id="2058D09FD61DFF8CFE0D802DFBF3FE9D" box="[396,480,335,354]" pageId="8" pageNumber="286" type="paratype">paratype</typeStatus>
individual studied in detail and to larva. Measures given in µm.
</paragraph>
<paragraph id="FF5C6E3DD61DFF8CFF4580EFFF7DFAC9" pageId="8" pageNumber="286">
<table id="8DE39C9DD61D007BFF4580EFFF7DFAC7" box="[196,1390,397,1336]" gridcols="6" gridrows="29" pageId="8" pageNumber="286">
<tr id="41D36C7FD61D007BFF4580EFFF7DFE5E" box="[196,1390,397,417]" gridrow="0" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF4580EFFF7DFE5E" box="[196,1390,397,417]" colspan="6" colspanRight="5" gridcol="0" gridrow="0" pageId="8" pageNumber="286">
<emphasis id="CD97B22FD61DFF8CFF4580EFFB70FE5E" bold="true" box="[196,355,397,417]" pageId="8" pageNumber="286">adult (paratype)</emphasis>
</th>
</tr>
<tr id="41D36C7FD61D007BFF4580CCFF7DFE3C" box="[196,1390,430,451]" gridrow="1" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF4580CCFBAFFE3C" box="[196,444,430,451]" gridcol="0" gridrow="1" pageId="8" pageNumber="286">body – total length</th>
<td id="02020503D61D007BFD9F80CCF859FE3C" box="[542,586,430,451]" gridcol="1" gridrow="1" pageId="8" pageNumber="286">1003</td>
<td id="02020503D61D007BFD1380CCF932FE3C" box="[658,801,430,451]" gridcol="2" gridrow="1" pageId="8" pageNumber="286">sensillus</td>
<td id="02020503D61D007BFCE980CCF9A6FE3C" box="[872,949,430,451]" gridcol="3" gridrow="1" pageId="8" pageNumber="286">298–300</td>
<td id="02020503D61D007BFB8280CCFEF6FE3C" box="[1027,1253,430,451]" gridcol="4" gridrow="1" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF80CCFE2CFE3E" box="[1070,1087,430,449]" italics="true" pageId="8" pageNumber="286">l”</emphasis>
on genu I
</td>
<td id="02020503D61D007BFAA080CCFF7DFE3C" box="[1313,1390,430,451]" gridcol="5" gridrow="1" pageId="8" pageNumber="286">58</td>
</tr>
<tr id="41D36C7FD61D007BFF4580B2FF7DFE1B" box="[196,1390,464,484]" gridrow="2" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF4580B2FBAFFE1B" box="[196,444,464,484]" gridcol="0" gridrow="2" pageId="8" pageNumber="286">body – ventral length</th>
<td id="02020503D61D007BFD9F80B2F859FE1B" box="[542,586,464,484]" gridcol="1" gridrow="2" pageId="8" pageNumber="286">831</td>
<td id="02020503D61D007BFD1380B2F932FE1B" box="[658,801,464,484]" gridcol="2" gridrow="2" pageId="8" pageNumber="286">spina adnatum</td>
<td id="02020503D61D007BFCE980B2F9A6FE1B" box="[872,949,464,484]" gridcol="3" gridrow="2" pageId="8" pageNumber="286">18–19</td>
<td id="02020503D61D007BFB8280B2FEF6FE1B" box="[1027,1253,464,484]" gridcol="4" gridrow="2" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF80B2FE29FE1C" box="[1070,1082,464,483]" italics="true" pageId="8" pageNumber="286">l’</emphasis>
on genu I
</td>
<td id="02020503D61D007BFAA080B2FF7DFE1B" box="[1313,1390,464,484]" gridcol="5" gridrow="2" pageId="8" pageNumber="286">39</td>
</tr>
<tr id="41D36C7FD61D007BFF45808CFF7DFDFC" box="[196,1390,494,515]" gridrow="3" pageId="8" pageNumber="286" rowspan-2="1" rowspan-3="1">
<th id="02020503D61D007BFF45808CFBAFFDFC" box="[196,444,494,515]" gridcol="0" gridrow="3" pageId="8" pageNumber="286">body – maximum width</th>
<td id="02020503D61D007BFD9F808CF859FDFC" box="[542,586,494,515]" gridcol="1" gridrow="3" pageId="8" pageNumber="286">670</td>
<td id="02020503D61D007BFB82808CFEF6FDFC" box="[1027,1253,494,515]" gridcol="4" gridrow="3" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF808CFE2AFDFE" box="[1070,1081,494,513]" italics="true" pageId="8" pageNumber="286">d</emphasis>
on genu I
</td>
<td id="02020503D61D007BFAA0808CFF7DFDFC" box="[1313,1390,494,515]" gridcol="5" gridrow="3" pageId="8" pageNumber="286">30</td>
</tr>
<tr id="41D36C7FD61D007BFF45836DFF7DFDDB" box="[196,1390,527,548]" gridrow="4" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF45836DFBAFFDDB" box="[196,444,527,548]" gridcol="0" gridrow="4" pageId="8" pageNumber="286">prodorsum – length</th>
<td id="02020503D61D007BFD9F836DF859FDDB" box="[542,586,527,548]" gridcol="1" gridrow="4" pageId="8" pageNumber="286">345</td>
<td id="02020503D61D007BFD13836DF932FDDB" box="[658,801,527,548]" gridcol="2" gridrow="4" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8373F8DDFDDC" box="[700,718,529,547]" italics="true" pageId="8" pageNumber="286">ro</emphasis>
</td>
<td id="02020503D61D007BFCE9836DF9A6FDDB" box="[872,949,527,548]" gridcol="3" gridrow="4" pageId="8" pageNumber="286">100</td>
<td id="02020503D61D007BFB82836DFEF6FDDB" box="[1027,1253,527,548]" gridcol="4" gridrow="4" pageId="8" pageNumber="286">
solenidion 
<emphasis id="CD97B22FD61DFF8CFBEC8372FE6AFDDD" box="[1133,1145,528,546]" italics="true" pageId="8" pageNumber="286">σ</emphasis>
on genu I
</td>
<td id="02020503D61D007BFAA0836DFF7DFDDB" box="[1313,1390,527,548]" gridcol="5" gridrow="4" pageId="8" pageNumber="286">42</td>
</tr>
<tr id="41D36C7FD61D007BFF458352FF7DFDBA" box="[196,1390,560,581]" gridrow="5" pageId="8" pageNumber="286" rowspan-4="1" rowspan-5="1">
<th id="02020503D61D007BFF458352FBAFFDBA" box="[196,444,560,581]" gridcol="0" gridrow="5" pageId="8" pageNumber="286">prodorsum – width</th>
<td id="02020503D61D007BFD9F8352F859FDBA" box="[542,586,560,581]" gridcol="1" gridrow="5" pageId="8" pageNumber="286">410</td>
<td id="02020503D61D007BFD138352F932FDBA" box="[658,801,560,581]" gridcol="2" gridrow="5" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8350F8D9FDBA" box="[700,714,562,581]" italics="true" pageId="8" pageNumber="286">le</emphasis>
</td>
<td id="02020503D61D007BFCE98352F9A6FDBA" box="[872,949,560,581]" gridcol="3" gridrow="5" pageId="8" pageNumber="286">139</td>
</tr>
<tr id="41D36C7FD61D007BFF458333FF7DFD99" box="[196,1390,593,614]" gridrow="6" pageId="8" pageNumber="286" rowspan-0="1" rowspan-1="1">
<td id="02020503D61D007BFD138333F932FD99" box="[658,801,593,614]" gridcol="2" gridrow="6" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8331F8DDFD9A" box="[700,718,595,613]" italics="true" pageId="8" pageNumber="286">in</emphasis>
</td>
<td id="02020503D61D007BFCE98333F9A6FD99" box="[872,949,593,614]" gridcol="3" gridrow="6" pageId="8" pageNumber="286">94</td>
<td id="02020503D61D007BFB828333FEF6FD99" box="[1027,1253,593,614]" gridcol="4" gridrow="6" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF8331FE29FD99" box="[1070,1082,595,614]" italics="true" pageId="8" pageNumber="286">l’</emphasis>
of tibia I
</td>
<td id="02020503D61D007BFAA08333FF7DFD99" box="[1313,1390,593,614]" gridcol="5" gridrow="6" pageId="8" pageNumber="286">131</td>
</tr>
<tr id="41D36C7FD61D007BFF458310FF7DFD78" box="[196,1390,626,647]" gridrow="7" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF458310FBAFFD78" box="[196,444,626,647]" gridcol="0" gridrow="7" pageId="8" pageNumber="286">femur I</th>
<td id="02020503D61D007BFD9F8310F859FD78" box="[542,586,626,647]" gridcol="1" gridrow="7" pageId="8" pageNumber="286">342</td>
<td id="02020503D61D007BFD138310F932FD78" box="[658,801,626,647]" gridcol="2" gridrow="7" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8316F8DCFD79" box="[700,719,628,646]" italics="true" pageId="8" pageNumber="286">ex</emphasis>
</td>
<td id="02020503D61D007BFCE98310F9A6FD78" box="[872,949,626,647]" gridcol="3" gridrow="7" pageId="8" pageNumber="286">51</td>
<td id="02020503D61D007BFB828310FEF6FD78" box="[1027,1253,626,647]" gridcol="4" gridrow="7" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF8310FE57FD7A" box="[1070,1092,626,645]" italics="true" pageId="8" pageNumber="286">v”</emphasis>
of tibia I
</td>
<td id="02020503D61D007BFAA08310FF7DFD78" box="[1313,1390,626,647]" gridcol="5" gridrow="7" pageId="8" pageNumber="286">112–115</td>
</tr>
<tr id="41D36C7FD61D007BFF4583F1FF7DFD55" box="[196,1390,659,682]" gridrow="8" pageId="8" pageNumber="286" rowspan-2="1" rowspan-3="1">
<th id="02020503D61D007BFF4583F1FBAFFD55" box="[196,444,659,682]" gridcol="0" gridrow="8" pageId="8" pageNumber="286">genu I</th>
<td id="02020503D61D007BFD9F83F1F859FD55" box="[542,586,659,682]" gridcol="1" gridrow="8" pageId="8" pageNumber="286">119</td>
<td id="02020503D61D007BFB8283F1FEF6FD55" box="[1027,1253,659,682]" gridcol="4" gridrow="8" pageId="8" pageNumber="286">
solenidion 
<emphasis id="CD97B22FD61DFF8CFBEC83F7FE6FFD58" box="[1133,1148,661,679]" italics="true" pageId="8" pageNumber="286">φ</emphasis>
<emphasis id="CD97B22FD61DFF8CFB0083FCFE9BFD55" box="[1153,1160,670,682]" italics="true" pageId="8" pageNumber="286">1</emphasis>
of tibia I
</td>
<td id="02020503D61D007BFAA083F1FF7DFD55" box="[1313,1390,659,682]" gridcol="5" gridrow="8" pageId="8" pageNumber="286">112</td>
</tr>
<tr id="41D36C7FD61D007BFF4583D6FF7DFD34" box="[196,1390,692,715]" gridrow="9" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF4583D6FBAFFD34" box="[196,444,692,715]" gridcol="0" gridrow="9" pageId="8" pageNumber="286">tibia I</th>
<td id="02020503D61D007BFD9F83D6F859FD34" box="[542,586,692,715]" gridcol="1" gridrow="9" pageId="8" pageNumber="286">229</td>
<td id="02020503D61D007BFD1383D6F932FD34" box="[658,801,692,715]" gridcol="2" gridrow="9" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D83D4F8DCFD36" box="[700,719,694,713]" italics="true" pageId="8" pageNumber="286">c1</emphasis>
</td>
<td id="02020503D61D007BFCE983D6F9A6FD34" box="[872,949,692,715]" gridcol="3" gridrow="9" pageId="8" pageNumber="286">138–144</td>
<td id="02020503D61D007BFB8283D6FEF6FD34" box="[1027,1253,692,715]" gridcol="4" gridrow="9" pageId="8" pageNumber="286">
solenidion 
<emphasis id="CD97B22FD61DFF8CFBEC83D4FE6FFD37" box="[1133,1148,694,712]" italics="true" pageId="8" pageNumber="286">φ</emphasis>
<emphasis id="CD97B22FD61DFF8CFB0083DDFE9BFD34" box="[1153,1160,703,715]" italics="true" pageId="8" pageNumber="286">2</emphasis>
of tibia I
</td>
<td id="02020503D61D007BFAA083D6FF7DFD34" box="[1313,1390,692,715]" gridcol="5" gridrow="9" pageId="8" pageNumber="286">36</td>
</tr>
<tr id="41D36C7FD61D007BFF4583B7FF7DFD15" box="[196,1390,725,746]" gridrow="10" pageId="8" pageNumber="286" rowspan-4="1" rowspan-5="1">
<th id="02020503D61D007BFF4583B7FBAFFD15" box="[196,444,725,746]" gridcol="0" gridrow="10" pageId="8" pageNumber="286">tarsus I</th>
<td id="02020503D61D007BFD9F83B7F859FD15" box="[542,586,725,746]" gridcol="1" gridrow="10" pageId="8" pageNumber="286">340</td>
<td id="02020503D61D007BFD1383B7F932FD15" box="[658,801,725,746]" gridcol="2" gridrow="10" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D83B5F8DCFD15" box="[700,719,727,746]" italics="true" pageId="8" pageNumber="286">c2</emphasis>
</td>
<td id="02020503D61D007BFCE983B7F9A6FD15" box="[872,949,725,746]" gridcol="3" gridrow="10" pageId="8" pageNumber="286">97–99</td>
</tr>
<tr id="41D36C7FD61D007BFF458394FF7DFCF4" box="[196,1390,758,779]" gridrow="11" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF458394FBAFFCF4" box="[196,444,758,779]" gridcol="0" gridrow="11" pageId="8" pageNumber="286">leg I total length</th>
<td id="02020503D61D007BFD9F8394F859FCF4" box="[542,586,758,779]" gridcol="1" gridrow="11" pageId="8" pageNumber="286">1030</td>
<td id="02020503D61D007BFD138394F932FCF4" box="[658,801,758,779]" gridcol="2" gridrow="11" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8395F8DFFCF5" box="[700,716,759,778]" italics="true" pageId="8" pageNumber="286">la</emphasis>
</td>
<td id="02020503D61D007BFCE98394F9A6FCF4" box="[872,949,758,779]" gridcol="3" gridrow="11" pageId="8" pageNumber="286">78–81</td>
<td id="02020503D61D007BFB828394FEF6FCF4" box="[1027,1253,758,779]" gridcol="4" gridrow="11" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF8394FE2DFCF7" box="[1070,1086,758,776]" italics="true" pageId="8" pageNumber="286">tc</emphasis>
on tarsus I
</td>
<td id="02020503D61D007BFAA08394FF7DFCF4" box="[1313,1390,758,779]" gridcol="5" gridrow="11" pageId="8" pageNumber="286">130</td>
</tr>
<tr id="41D36C7FD61D007BFF458275FF7DFCD3" box="[196,1390,791,812]" gridrow="12" pageId="8" pageNumber="286" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<td id="02020503D61D007BFD138275F932FCD3" box="[658,801,791,812]" gridcol="2" gridrow="12" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D827AF8C0FCD4" box="[700,723,792,811]" italics="true" pageId="8" pageNumber="286">lm</emphasis>
</td>
<td id="02020503D61D007BFCE98275F9A6FCD3" box="[872,949,791,812]" gridcol="3" gridrow="12" pageId="8" pageNumber="286">67–75</td>
</tr>
<tr id="41D36C7FD61D007BFF45825AFF7DFCB2" box="[196,1390,824,845]" gridrow="13" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF45825AFBAFFCB2" box="[196,444,824,845]" gridcol="0" gridrow="13" pageId="8" pageNumber="286">trochanter IV</th>
<td id="02020503D61D007BFD9F825AF859FCB2" box="[542,586,824,845]" gridcol="1" gridrow="13" pageId="8" pageNumber="286">216</td>
<td id="02020503D61D007BFD13825AF932FCB2" box="[658,801,824,845]" gridcol="2" gridrow="13" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D825BF8DEFCB3" box="[700,717,825,844]" italics="true" pageId="8" pageNumber="286">lp</emphasis>
</td>
<td id="02020503D61D007BFCE9825AF9A6FCB2" box="[872,949,824,845]" gridcol="3" gridrow="13" pageId="8" pageNumber="286">73</td>
<td id="02020503D61D007BFB82825AFEF6FCB2" box="[1027,1253,824,845]" gridcol="4" gridrow="13" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF825AFE29FCB4" box="[1070,1082,824,843]" italics="true" pageId="8" pageNumber="286">l’</emphasis>
of tibia IV
</td>
<td id="02020503D61D007BFAA0825AFF7DFCB2" box="[1313,1390,824,845]" gridcol="5" gridrow="13" pageId="8" pageNumber="286">112</td>
</tr>
<tr id="41D36C7FD61D007BFF45823BFF7DFC91" box="[196,1390,857,878]" gridrow="14" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF45823BFBAFFC91" box="[196,444,857,878]" gridcol="0" gridrow="14" pageId="8" pageNumber="286">femur IV</th>
<td id="02020503D61D007BFD9F823BF859FC91" box="[542,586,857,878]" gridcol="1" gridrow="14" pageId="8" pageNumber="286">264</td>
<td id="02020503D61D007BFD13823BF932FC91" box="[658,801,857,878]" gridcol="2" gridrow="14" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8238F8C1FC92" box="[700,722,858,877]" italics="true" pageId="8" pageNumber="286">h3</emphasis>
</td>
<td id="02020503D61D007BFCE9823BF9A6FC91" box="[872,949,857,878]" gridcol="3" gridrow="14" pageId="8" pageNumber="286">75–88</td>
<td id="02020503D61D007BFB82823BFEF6FC91" box="[1027,1253,857,878]" gridcol="4" gridrow="14" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF8239FE2CFC91" box="[1070,1087,859,878]" italics="true" pageId="8" pageNumber="286">v’</emphasis>
of tibia IV
</td>
<td id="02020503D61D007BFAA0823BFF7DFC91" box="[1313,1390,857,878]" gridcol="5" gridrow="14" pageId="8" pageNumber="286">107–117</td>
</tr>
<tr id="41D36C7FD61D007BFF458218FF7DFC70" box="[196,1390,890,911]" gridrow="15" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF458218FBAFFC70" box="[196,444,890,911]" gridcol="0" gridrow="15" pageId="8" pageNumber="286">genu IV</th>
<td id="02020503D61D007BFD9F8218F859FC70" box="[542,586,890,911]" gridcol="1" gridrow="15" pageId="8" pageNumber="286">109</td>
<td id="02020503D61D007BFD138218F932FC70" box="[658,801,890,911]" gridcol="2" gridrow="15" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8219F8C1FC71" box="[700,722,891,910]" italics="true" pageId="8" pageNumber="286">h2</emphasis>
</td>
<td id="02020503D61D007BFCE98218F9A6FC70" box="[872,949,890,911]" gridcol="3" gridrow="15" pageId="8" pageNumber="286">70</td>
<td id="02020503D61D007BFB828218FEF6FC70" box="[1027,1253,890,911]" gridcol="4" gridrow="15" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF8218FE57FC72" box="[1070,1092,890,909]" italics="true" pageId="8" pageNumber="286">v”</emphasis>
of tibia IV
</td>
<td id="02020503D61D007BFAA08218FF7DFC70" box="[1313,1390,890,911]" gridcol="5" gridrow="15" pageId="8" pageNumber="286">72</td>
</tr>
<tr id="41D36C7FD61D007BFF4582FEFF7DFC4F" box="[196,1390,924,944]" gridrow="16" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF4582FEFBAFFC4F" box="[196,444,924,944]" gridcol="0" gridrow="16" pageId="8" pageNumber="286">tibia IV</th>
<td id="02020503D61D007BFD9F82FEF859FC4F" box="[542,586,924,944]" gridcol="1" gridrow="16" pageId="8" pageNumber="286">222</td>
<td id="02020503D61D007BFD1382FEF932FC4F" box="[658,801,924,944]" gridcol="2" gridrow="16" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D82FEF8C1FC50" box="[700,722,924,943]" italics="true" pageId="8" pageNumber="286">h1</emphasis>
</td>
<td id="02020503D61D007BFCE982FEF9A6FC4F" box="[872,949,924,944]" gridcol="3" gridrow="16" pageId="8" pageNumber="286">68</td>
<td id="02020503D61D007BFB8282FEFEF6FC4F" box="[1027,1253,924,944]" gridcol="4" gridrow="16" pageId="8" pageNumber="286">
solenidion 
<emphasis id="CD97B22FD61DFF8CFBEC82FFFE6FFC50" box="[1133,1148,925,943]" italics="true" pageId="8" pageNumber="286">φ</emphasis>
of tibia IV
</td>
<td id="02020503D61D007BFAA082FEFF7DFC4F" box="[1313,1390,924,944]" gridcol="5" gridrow="16" pageId="8" pageNumber="286">69</td>
</tr>
<tr id="41D36C7FD61D007BFF4582D8FF7DFC30" box="[196,1390,954,975]" gridrow="17" pageId="8" pageNumber="286" rowspan-4="1" rowspan-5="1">
<th id="02020503D61D007BFF4582D8FBAFFC30" box="[196,444,954,975]" gridcol="0" gridrow="17" pageId="8" pageNumber="286">tarsus IV</th>
<td id="02020503D61D007BFD9F82D8F859FC30" box="[542,586,954,975]" gridcol="1" gridrow="17" pageId="8" pageNumber="286">379</td>
<td id="02020503D61D007BFD1382D8F932FC30" box="[658,801,954,975]" gridcol="2" gridrow="17" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D82DEF8C1FC30" box="[700,722,956,975]" italics="true" pageId="8" pageNumber="286">p1</emphasis>
</td>
<td id="02020503D61D007BFCE982D8F9A6FC30" box="[872,949,954,975]" gridcol="3" gridrow="17" pageId="8" pageNumber="286">58</td>
</tr>
<tr id="41D36C7FD61D007BFF4582B9FF7DFC0F" box="[196,1390,987,1008]" gridrow="18" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF4582B9FBAFFC0F" box="[196,444,987,1008]" gridcol="0" gridrow="18" pageId="8" pageNumber="286">leg IV total length</th>
<td id="02020503D61D007BFD9F82B9F859FC0F" box="[542,586,987,1008]" gridcol="1" gridrow="18" pageId="8" pageNumber="286">1190</td>
<td id="02020503D61D007BFD1382B9F932FC0F" box="[658,801,987,1008]" gridcol="2" gridrow="18" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D82BFF8C1FC0F" box="[700,722,989,1008]" italics="true" pageId="8" pageNumber="286">p2</emphasis>
</td>
<td id="02020503D61D007BFCE982B9F9A6FC0F" box="[872,949,987,1008]" gridcol="3" gridrow="18" pageId="8" pageNumber="286">38 – 45</td>
<td id="02020503D61D007BFB8282B9FEF6FC0F" box="[1027,1253,987,1008]" gridcol="4" gridrow="18" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF82B9FE55FC11" box="[1070,1094,987,1006]" italics="true" pageId="8" pageNumber="286">ft”</emphasis>
of tarsus IV
</td>
<td id="02020503D61D007BFAA082B9FF7DFC0F" box="[1313,1390,987,1008]" gridcol="5" gridrow="18" pageId="8" pageNumber="286">138</td>
</tr>
<tr id="41D36C7FD61D007BFF45829EFF7DFBEE" box="[196,1390,1020,1041]" gridrow="19" pageId="8" pageNumber="286" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1">
<td id="02020503D61D007BFD13829EF932FBEE" box="[658,801,1020,1041]" gridcol="2" gridrow="19" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D829CF8C1FBEE" box="[700,722,1022,1041]" italics="true" pageId="8" pageNumber="286">p3</emphasis>
</td>
<td id="02020503D61D007BFCE9829EF9A6FBEE" box="[872,949,1020,1041]" gridcol="3" gridrow="19" pageId="8" pageNumber="286">31 – 45</td>
</tr>
<tr id="41D36C7FD61D007BFF45857FFF7DFBCD" box="[196,1390,1053,1074]" gridrow="20" pageId="8" pageNumber="286" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="02020503D61D007BFF45857FFBAFFBCD" box="[196,444,1053,1074]" gridcol="0" gridrow="20" pageId="8" pageNumber="286">ratio leg I/ventral length</th>
<td id="02020503D61D007BFD9F857FF859FBCD" box="[542,586,1053,1074]" gridcol="1" gridrow="20" pageId="8" pageNumber="286">1,24</td>
</tr>
<tr id="41D36C7FD61D007BFF45855CFF7DFBAC" box="[196,1390,1086,1107]" gridrow="21" pageId="8" pageNumber="286" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="02020503D61D007BFF45855CFBAFFBAC" box="[196,444,1086,1107]" gridcol="0" gridrow="21" pageId="8" pageNumber="286">ratio leg I/total length</th>
<td id="02020503D61D007BFD9F855CF859FBAC" box="[542,586,1086,1107]" gridcol="1" gridrow="21" pageId="8" pageNumber="286">1,03</td>
</tr>
<tr id="41D36C7FD61D007BFF45853DFF7DFB8C" box="[196,1390,1119,1139]" gridrow="22" pageId="8" pageNumber="286" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="02020503D61D007BFF45853DFBAFFB8C" box="[196,444,1119,1139]" gridcol="0" gridrow="22" pageId="8" pageNumber="286">ratio leg IV/ventral length</th>
<td id="02020503D61D007BFD9F853DF859FB8C" box="[542,586,1119,1139]" gridcol="1" gridrow="22" pageId="8" pageNumber="286">1,43</td>
</tr>
<tr id="41D36C7FD61D007BFF4585E2FF7DFB6B" box="[196,1390,1152,1172]" gridrow="23" pageId="8" pageNumber="286" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="02020503D61D007BFF4585E2FBAFFB6B" box="[196,444,1152,1172]" gridcol="0" gridrow="23" pageId="8" pageNumber="286">ratio leg IV/total length</th>
<td id="02020503D61D007BFD9F85E2F859FB6B" box="[542,586,1152,1172]" gridcol="1" gridrow="23" pageId="8" pageNumber="286">1,19</td>
</tr>
<tr id="41D36C7FD61D007BFF4585C2FF7DFB4B" box="[196,1390,1184,1204]" gridrow="24" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF4585C2FF7DFB4B" box="[196,1390,1184,1204]" colspan="6" colspanRight="5" gridcol="0" gridrow="24" pageId="8" pageNumber="286">
<emphasis id="CD97B22FD61DFF8CFF4585C2FAE6FB4B" bold="true" box="[196,245,1184,1204]" pageId="8" pageNumber="286">larva</emphasis>
</th>
</tr>
<tr id="41D36C7FD61D007BFF4585A2FF7DFB2A" box="[196,1390,1216,1237]" gridrow="25" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF4585A2FBAFFB2A" box="[196,444,1216,1237]" gridcol="0" gridrow="25" pageId="8" pageNumber="286">body length</th>
<td id="02020503D61D007BFD9F85A2F859FB2A" box="[542,586,1216,1237]" gridcol="1" gridrow="25" pageId="8" pageNumber="286">430</td>
<td id="02020503D61D007BFD1385A2F932FB2A" box="[658,801,1216,1237]" gridcol="2" gridrow="25" pageId="8" pageNumber="286">sensillus</td>
<td id="02020503D61D007BFCE985A2F9A6FB2A" box="[872,949,1216,1237]" gridcol="3" gridrow="25" pageId="8" pageNumber="286">180</td>
<td id="02020503D61D007BFB8285A2FEF6FB2A" box="[1027,1253,1216,1237]" gridcol="4" gridrow="25" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF85A2FE52FB2C" box="[1070,1089,1216,1235]" italics="true" pageId="8" pageNumber="286">c1</emphasis>
</td>
<td id="02020503D61D007BFAA085A2FF7DFB2A" box="[1313,1390,1216,1237]" gridcol="5" gridrow="25" pageId="8" pageNumber="286">185</td>
</tr>
<tr id="41D36C7FD61D007BFF458583FF7DFB09" box="[196,1390,1249,1270]" gridrow="26" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF458583FBAFFB09" box="[196,444,1249,1270]" gridcol="0" gridrow="26" pageId="8" pageNumber="286">body width</th>
<td id="02020503D61D007BFD9F8583F859FB09" box="[542,586,1249,1270]" gridcol="1" gridrow="26" pageId="8" pageNumber="286">240</td>
<td id="02020503D61D007BFD138583F932FB09" box="[658,801,1249,1270]" gridcol="2" gridrow="26" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8581F8DDFB0A" box="[700,718,1251,1269]" italics="true" pageId="8" pageNumber="286">ro</emphasis>
</td>
<td id="02020503D61D007BFCE98583F9A6FB09" box="[872,949,1249,1270]" gridcol="3" gridrow="26" pageId="8" pageNumber="286">48</td>
<td id="02020503D61D007BFB828583FEF6FB09" box="[1027,1253,1249,1270]" gridcol="4" gridrow="26" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF8583FE52FB0B" box="[1070,1089,1249,1268]" italics="true" pageId="8" pageNumber="286">c2</emphasis>
</td>
<td id="02020503D61D007BFAA08583FF7DFB09" box="[1313,1390,1249,1270]" gridcol="5" gridrow="26" pageId="8" pageNumber="286">282</td>
</tr>
<tr id="41D36C7FD61D007BFF458460FF7DFAE8" box="[196,1390,1282,1303]" gridrow="27" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF458460FBAFFAE8" box="[196,444,1282,1303]" gridcol="0" gridrow="27" pageId="8" pageNumber="286">prodorsum length</th>
<td id="02020503D61D007BFD9F8460F859FAE8" box="[542,586,1282,1303]" gridcol="1" gridrow="27" pageId="8" pageNumber="286">161</td>
<td id="02020503D61D007BFD138460F932FAE8" box="[658,801,1282,1303]" gridcol="2" gridrow="27" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8466F8D9FAE8" box="[700,714,1284,1303]" italics="true" pageId="8" pageNumber="286">le</emphasis>
</td>
<td id="02020503D61D007BFCE98460F9A6FAE8" box="[872,949,1282,1303]" gridcol="3" gridrow="27" pageId="8" pageNumber="286">123</td>
<td id="02020503D61D007BFB828460FEF6FAE8" box="[1027,1253,1282,1303]" gridcol="4" gridrow="27" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF8460FE57FAEA" box="[1070,1092,1282,1301]" italics="true" pageId="8" pageNumber="286">h1</emphasis>
</td>
<td id="02020503D61D007BFAA08460FF7DFAE8" box="[1313,1390,1282,1303]" gridcol="5" gridrow="27" pageId="8" pageNumber="286">277</td>
</tr>
<tr id="41D36C7FD61D007BFF458441FF7DFAC7" box="[196,1390,1315,1336]" gridrow="28" pageId="8" pageNumber="286">
<th id="02020503D61D007BFF458441FBAFFAC7" box="[196,444,1315,1336]" gridcol="0" gridrow="28" pageId="8" pageNumber="286">prodorsum width</th>
<td id="02020503D61D007BFD9F8441F859FAC7" box="[542,586,1315,1336]" gridcol="1" gridrow="28" pageId="8" pageNumber="286">178</td>
<td id="02020503D61D007BFD138441F932FAC7" box="[658,801,1315,1336]" gridcol="2" gridrow="28" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFD3D8447F8DDFAC8" box="[700,718,1317,1335]" italics="true" pageId="8" pageNumber="286">in</emphasis>
</td>
<td id="02020503D61D007BFCE98441F9A6FAC7" box="[872,949,1315,1336]" gridcol="3" gridrow="28" pageId="8" pageNumber="286">55</td>
<td id="02020503D61D007BFB828441FEF6FAC7" box="[1027,1253,1315,1336]" gridcol="4" gridrow="28" pageId="8" pageNumber="286">
seta 
<emphasis id="CD97B22FD61DFF8CFBAF8441FE2CFAC9" box="[1070,1087,1315,1334]" italics="true" pageId="8" pageNumber="286">lp</emphasis>
</td>
<td id="02020503D61D007BFAA08441FF7DFAC7" box="[1313,1390,1315,1336]" gridcol="5" gridrow="28" pageId="8" pageNumber="286">461</td>
</tr>
</table>
</paragraph>
<paragraph id="FF5C6E3DD61CFF8DFF44837EFB0CF924" blockId="9.[159,762,539,1755]" pageId="9" pageNumber="287">
Notogaster — 
<figureCitation id="67D872B8D61CFF8DFE098379F805FDCC" box="[392,534,539,563]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="9" pageNumber="287">Figures 1A</figureCitation>
, 
<figureCitation id="67D872B8D61CFF8DFDAE8379F895FDCC" box="[559,646,539,563]" captionStart="FIGURE" captionStartId="6.[159,171,1818,1837]" captionTargetBox="[227,1361,349,1785]" captionTargetId="figure-2@6.[227,1361,349,1785]" captionTargetPageId="6" captionText="FIGURE 4: Damaeus lupus n. sp. (adult, paratype): A – lateral view of the body with legs, B – dorsal view of the body, C – sensillus and bothridium, D – spina adnatum, E – rostrum and gnathosoma from lateral view, F – anogenital region, lateral view. Acronyms: bo – bothridium, chel – chelicere, pp – palps, s. ad. – spina adnatum." figureDoi="http://doi.org/10.5281/zenodo.4667412" httpUri="https://zenodo.org/record/4667412/files/figure.png" pageId="9" pageNumber="287">4A,B,D</figureCitation>
. Broadly ovate, narrowed anteriorly. Spinae adnatae (
<emphasis id="CD97B22FD61CFF8DFD41835CF8E2FDA9" box="[704,753,574,598]" italics="true" pageId="9" pageNumber="287">s.ad.</emphasis>
) small, short and triangular (
<figureCitation id="67D872B8D61CFF8DFD808303F870FD86" box="[513,611,609,633]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="9" pageNumber="287">Figs 2D</figureCitation>
, 
<figureCitation id="67D872B8D61CFF8DFDF58303F88AFD86" box="[628,665,609,633]" captionStart="FIGURE" captionStartId="6.[159,171,1818,1837]" captionTargetBox="[227,1361,349,1785]" captionTargetId="figure-2@6.[227,1361,349,1785]" captionTargetPageId="6" captionText="FIGURE 4: Damaeus lupus n. sp. (adult, paratype): A – lateral view of the body with legs, B – dorsal view of the body, C – sensillus and bothridium, D – spina adnatum, E – rostrum and gnathosoma from lateral view, F – anogenital region, lateral view. Acronyms: bo – bothridium, chel – chelicere, pp – palps, s. ad. – spina adnatum." figureDoi="http://doi.org/10.5281/zenodo.4667412" httpUri="https://zenodo.org/record/4667412/files/figure.png" pageId="9" pageNumber="287">4D</figureCitation>
), pointing anteriad or slightly anterolaterad. Two distinct, arched transversal folds or ridges present at notogaster anterior edge, between 
<emphasis id="CD97B22FD61CFF8DFDC583A8F866FD1D" box="[580,629,714,738]" italics="true" pageId="9" pageNumber="287">s.ad.</emphasis>
and anterior to 
<emphasis id="CD97B22FD61CFF8DFF71838CFB1AFCF9" box="[240,265,750,774]" italics="true" pageId="9" pageNumber="287">c1</emphasis>
insertions. Notogastral setae (
<figureCitation id="67D872B8D61CFF8DFDF0838FF8E7FCF9" box="[625,756,749,774]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="9" pageNumber="287">Figures 1A</figureCitation>
, 
<figureCitation id="67D872B8D61CFF8DFF1E8273FAD1FCD6" box="[159,194,785,809]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="9" pageNumber="287">2D</figureCitation>
) of different size and shape, with rough surface except setae of row 
<emphasis id="CD97B22FD61CFF8DFE1D8256FBBAFCB4" box="[412,425,820,843]" italics="true" pageId="9" pageNumber="287">p</emphasis>
, in principle with decreasing size from anterior to posterior ones. Setae 
<emphasis id="CD97B22FD61CFF8DFD658235F8FCFC91" box="[740,751,855,878]" italics="true" pageId="9" pageNumber="287">c</emphasis>
<subScript id="63676C78D61CFF8DFD6E8202F8EBFC8E" attach="left" box="[751,760,864,881]" fontSize="7" pageId="9" pageNumber="287">1</subScript>
and 
<emphasis id="CD97B22FD61CFF8DFF578218FAF2FC6E" box="[214,225,890,913]" italics="true" pageId="9" pageNumber="287">c</emphasis>
<subScript id="63676C78D61CFF8DFF6182E1FAFAFC6B" attach="left" box="[224,233,899,916]" fontSize="7" pageId="9" pageNumber="287">2</subScript>
longest, subequal in length, 
<emphasis id="CD97B22FD61CFF8DFDCC8218F84BFC6E" box="[589,600,890,913]" italics="true" pageId="9" pageNumber="287">c</emphasis>
<subScript id="63676C78D61CFF8DFDD982E1F872FC6B" attach="left" box="[600,609,899,916]" fontSize="7" pageId="9" pageNumber="287">1</subScript>
oriented anteriad, appearing slightly bent in dorsal view but distinctly curved upwards in lateral view (
<figureCitation id="67D872B8D61CFF8DFD2D82A2FAD0FC04" captionStart="FIGURE" captionStartId="6.[159,171,1818,1837]" captionTargetBox="[227,1361,349,1785]" captionTargetId="figure-2@6.[227,1361,349,1785]" captionTargetPageId="6" captionText="FIGURE 4: Damaeus lupus n. sp. (adult, paratype): A – lateral view of the body with legs, B – dorsal view of the body, C – sensillus and bothridium, D – spina adnatum, E – rostrum and gnathosoma from lateral view, F – anogenital region, lateral view. Acronyms: bo – bothridium, chel – chelicere, pp – palps, s. ad. – spina adnatum." figureDoi="http://doi.org/10.5281/zenodo.4667412" httpUri="https://zenodo.org/record/4667412/files/figure.png" pageId="9" pageNumber="287">Figure 4A</figureCitation>
), 
<emphasis id="CD97B22FD61CFF8DFF598286FAF0FC04" box="[216,227,996,1019]" italics="true" pageId="9" pageNumber="287">c</emphasis>
<subScript id="63676C78D61CFF8DFF62828EFAFFFC02" attach="left" box="[227,236,1004,1021]" fontSize="7" pageId="9" pageNumber="287">2</subScript>
slightly S-shaped, pointing more or less laterad, both distally attenuating and pointed. Distance of insertions 
<emphasis id="CD97B22FD61CFF8DFE018548FB98FBBE" box="[384,395,1066,1089]" italics="true" pageId="9" pageNumber="287">c</emphasis>
<subScript id="63676C78D61CFF8DFE0B8551FB80FBBB" attach="left" box="[394,403,1075,1092]" fontSize="7" pageId="9" pageNumber="287">1</subScript>
- 
<emphasis id="CD97B22FD61CFF8DFE1F8548FBBAFBBE" box="[414,425,1066,1089]" italics="true" pageId="9" pageNumber="287">c</emphasis>
<subScript id="63676C78D61CFF8DFE288551FBA1FBBB" attach="left" box="[425,434,1075,1092]" fontSize="7" pageId="9" pageNumber="287">1</subScript>
slightly less or subequal to 
<emphasis id="CD97B22FD61CFF8DFF1E852FFAB9FB9B" box="[159,170,1101,1124]" italics="true" pageId="9" pageNumber="287">c</emphasis>
<subScript id="63676C78D61CFF8DFF2B8534FAA0FB98" attach="left" box="[170,179,1110,1127]" fontSize="7" pageId="9" pageNumber="287">1</subScript>
- 
<emphasis id="CD97B22FD61CFF8DFF3C852FFADBFB9B" box="[189,200,1101,1124]" italics="true" pageId="9" pageNumber="287">c</emphasis>
<subScript id="63676C78D61CFF8DFF498534FAC2FB98" attach="left" box="[200,209,1110,1127]" fontSize="7" pageId="9" pageNumber="287">2</subScript>
. Seta 
<emphasis id="CD97B22FD61CFF8DFE9C852EFB23FB9B" box="[285,304,1100,1124]" italics="true" pageId="9" pageNumber="287">la</emphasis>
strongly curved, erect, attenuated distally with tips oriented anteriad. Setae 
<emphasis id="CD97B22FD61CFF8DFD08850DF8B6FB78" box="[649,677,1135,1159]" italics="true" pageId="9" pageNumber="287">lm</emphasis>
, 
<emphasis id="CD97B22FD61CFF8DFD39850DF8DEFB78" box="[696,717,1135,1159]" italics="true" pageId="9" pageNumber="287">lp</emphasis>
, 
<emphasis id="CD97B22FD61CFF8DFD5E850DF8E9FB78" box="[735,762,1135,1159]" italics="true" pageId="9" pageNumber="287">h2</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFF5785F0FAE3FB55" box="[214,240,1170,1194]" italics="true" pageId="9" pageNumber="287">h3</emphasis>
of similar shape, shorter and more robust, less bent or almost straight, almost spiniform with shorter and less acute tips, pointing laterad or posterolaterad (
<emphasis id="CD97B22FD61CFF8DFEAC8599FB5AFAEC" box="[301,329,1275,1299]" italics="true" pageId="9" pageNumber="287">lm</emphasis>
) or more or less posteriad, or slightly posterolaterad. Setae 
<emphasis id="CD97B22FD61CFF8DFE1B847CFBA6FAC9" box="[410,437,1310,1334]" italics="true" pageId="9" pageNumber="287">h1</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFE6A847DFBEBFAC9" box="[491,504,1311,1334]" italics="true" pageId="9" pageNumber="287">p</emphasis>
<subScript id="63676C78D61CFF8DFE78844AF811FAC6" attach="left" box="[505,514,1320,1337]" fontSize="7" pageId="9" pageNumber="287">1</subScript>
- 
<emphasis id="CD97B22FD61CFF8DFD8D847DF80AFAC9" box="[524,537,1311,1334]" italics="true" pageId="9" pageNumber="287">p</emphasis>
<subScript id="63676C78D61CFF8DFD98844AF831FAC6" attach="left" box="[537,546,1320,1337]" fontSize="7" pageId="9" pageNumber="287">3</subScript>
closer to posterior border of notogaster so that their insertions are not visible from dorsal view, finer and smoother than other notogastral setae, curved at their base, more or less straight distally. Setae 
<emphasis id="CD97B22FD61CFF8DFE7884CEF815FA3C" box="[505,518,1452,1475]" italics="true" pageId="9" pageNumber="287">p</emphasis>
<subScript id="63676C78D61CFF8DFD8784D6F81CFA3A" attach="left" box="[518,527,1460,1477]" fontSize="7" pageId="9" pageNumber="287">2</subScript>
and 
<emphasis id="CD97B22FD61CFF8DFDCD84CEF84AFA3C" box="[588,601,1452,1475]" italics="true" pageId="9" pageNumber="287">p</emphasis>
<subScript id="63676C78D61CFF8DFDD884D6F871FA3A" attach="left" box="[601,610,1460,1477]" fontSize="7" pageId="9" pageNumber="287">3</subScript>
shorter than other notogastral setae, directed laterad or even anteriolaterad. Circumgastric row of small and fine muscle sigillae present, larger sigillae present between 
<emphasis id="CD97B22FD61CFF8DFF6F8755FB19F9B0" box="[238,266,1591,1615]" italics="true" pageId="9" pageNumber="287">lm</emphasis>
insertions, 4 smaller groups or individual spots present in central part of notogaster. Common set of lyrifissures and notogastral gland openings present in usual position for 
<taxonomicName id="38E315BED61CFF8DFDAA87C3F89CF947" authorityName="C. L. Koch" authorityYear="1835" box="[555,655,1697,1720]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD61CFF8DFDAA87C3F89CF947" box="[555,655,1697,1720]" italics="true" pageId="9" pageNumber="287">Damaeus</emphasis>
</taxonomicName>
, difficult to observe.
</paragraph>
<paragraph id="FF5C6E3DD61CFF8DFF44879AFEB5FE8D" blockId="9.[159,762,1784,1949]" lastBlockId="9.[826,1428,311,370]" pageId="9" pageNumber="287">
Gnathosoma — 
<figureCitation id="67D872B8D61CFF8DFE15879AF808F8EF" box="[404,539,1784,1808]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="9" pageNumber="287">Figures 1B</figureCitation>
, 
<figureCitation id="67D872B8D61CFF8DFDAE879AF85CF8EF" box="[559,591,1784,1808]" captionStart="FIGURE" captionStartId="6.[159,171,1818,1837]" captionTargetBox="[227,1361,349,1785]" captionTargetId="figure-2@6.[227,1361,349,1785]" captionTargetPageId="6" captionText="FIGURE 4: Damaeus lupus n. sp. (adult, paratype): A – lateral view of the body with legs, B – dorsal view of the body, C – sensillus and bothridium, D – spina adnatum, E – rostrum and gnathosoma from lateral view, F – anogenital region, lateral view. Acronyms: bo – bothridium, chel – chelicere, pp – palps, s. ad. – spina adnatum." figureDoi="http://doi.org/10.5281/zenodo.4667412" httpUri="https://zenodo.org/record/4667412/files/figure.png" pageId="9" pageNumber="287">4E</figureCitation>
. As usual of 
<taxonomicName id="38E315BED61CFF8DFF1E867EFB10F8CC" authorityName="C. L. Koch" authorityYear="1835" box="[159,259,1820,1843]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="287" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD61CFF8DFF1E867EFB10F8CC" box="[159,259,1820,1843]" italics="true" pageId="9" pageNumber="287">Damaeus</emphasis>
</taxonomicName>
. Mentum relatively broad, slightly shorter than its width, its surface posteriad rugose. Setae 
<emphasis id="CD97B22FD61CFF8DFF4F8603FAC8F886" box="[206,219,1889,1913]" italics="true" pageId="9" pageNumber="287">h</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFEA08600FB25F886" box="[289,310,1890,1913]" italics="true" pageId="9" pageNumber="287">m</emphasis>
of medium length, subequal, seta 
<emphasis id="CD97B22FD61CFF8DFD6F8600F8E9F886" box="[750,762,1890,1913]" italics="true" pageId="9" pageNumber="287">a</emphasis>
shorter and finer. Palp setation 0-2-1-3-9(1), palp tarsal solenidion straight, blunt distally, adhered to surface of segment (
<figureCitation id="67D872B8D61CFF8DFBA58038FE89FE8D" box="[1060,1178,346,370]" captionStart="FIGURE" captionStartId="6.[159,171,1818,1837]" captionTargetBox="[227,1361,349,1785]" captionTargetId="figure-2@6.[227,1361,349,1785]" captionTargetPageId="6" captionText="FIGURE 4: Damaeus lupus n. sp. (adult, paratype): A – lateral view of the body with legs, B – dorsal view of the body, C – sensillus and bothridium, D – spina adnatum, E – rostrum and gnathosoma from lateral view, F – anogenital region, lateral view. Acronyms: bo – bothridium, chel – chelicere, pp – palps, s. ad. – spina adnatum." figureDoi="http://doi.org/10.5281/zenodo.4667412" httpUri="https://zenodo.org/record/4667412/files/figure.png" pageId="9" pageNumber="287">Figure 4E</figureCitation>
).
</paragraph>
<paragraph id="FF5C6E3DD61CFF8EFCE180C3F8B2FCB3" blockId="9.[826,1429,417,1949]" lastBlockId="10.[159,762,715,1949]" lastPageId="10" lastPageNumber="288" pageId="9" pageNumber="287">
Epimeral region — 
<figureCitation id="67D872B8D61CFF8DFBD580C3FEC5FE46" box="[1108,1238,417,441]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="9" pageNumber="287">Figures 1B</figureCitation>
, 
<figureCitation id="67D872B8D61CFF8DFB6680C3FF1BFE46" box="[1255,1288,417,441]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="9" pageNumber="287">2E</figureCitation>
. With deep epimeral and ventrosejugal grooves and very complex set of cuticular ridges and tubercles. Tectum of podocephalic fossa (
<emphasis id="CD97B22FD61CFF8DFBCB8368FE7BFDDD" box="[1098,1128,522,546]" italics="true" pageId="9" pageNumber="287">tpf</emphasis>
) with complicated structure, laterally with attenuated, rather sharp but only slightly protruding posterolateral projection. Mentotectum broad, collar-like, without median incision. Ridges and tubercles in epimeral area distinct, with complicated structure. Tubercles 
<emphasis id="CD97B22FD61CFF8DFAB883DBFF71FD2E" box="[1337,1378,697,721]" italics="true" pageId="9" pageNumber="287">E2a</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFCBB83BEF977FD0B" box="[826,868,732,756]" italics="true" pageId="9" pageNumber="287">E2p</emphasis>
flat and indistinct, placed on strong and curved ridges, which frame large part of epimeres I and II. These ridges sometimes with additional small angular projections laterally. Tubercles of ventrosejugal enantiophyse more prominent, 
<emphasis id="CD97B22FD61CFF8DFB5D820BFEEAFC7F" box="[1244,1273,873,896]" italics="true" pageId="9" pageNumber="287">Va</emphasis>
smaller than 
<emphasis id="CD97B22FD61CFF8DFCBB82EEF949FC5C" box="[826,858,908,931]" italics="true" pageId="9" pageNumber="287">Vp</emphasis>
, 
<emphasis id="CD97B22FD61CFF8DFCEB82EEF999FC5C" box="[874,906,908,931]" italics="true" pageId="9" pageNumber="287">Vp</emphasis>
bearing insertions of epimeral setae 
<emphasis id="CD97B22FD61CFF8DFAC482E9FF4CFC5C" box="[1349,1375,907,931]" italics="true" pageId="9" pageNumber="287">3b</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFCBB82CDF940FC38" box="[826,851,943,967]" italics="true" pageId="9" pageNumber="287">4b</emphasis>
on its base. Bases of tubercles 
<emphasis id="CD97B22FD61CFF8DFB5082CDFEEFFC38" box="[1233,1276,943,967]" italics="true" pageId="9" pageNumber="287">E2p</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFABD82CDFF4AFC39" box="[1340,1369,943,966]" italics="true" pageId="9" pageNumber="287">Va</emphasis>
connected by distinct curved ridge which may be interrupted in the middle. Ventrolateral ridge broadly interrupted or transformed into irregular longitudinal rugosity in area of epimeral furrow, well developed only in its posterior (between tubercles 
<emphasis id="CD97B22FD61CFF8DFAF9853DFF86FB89" box="[1400,1429,1119,1142]" italics="true" pageId="9" pageNumber="287">Va</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFCF085E3F99FFB67" box="[881,908,1153,1176]" italics="true" pageId="9" pageNumber="287">Sa</emphasis>
as a flatly arched ridge) and anterior ends (anterior to 
<emphasis id="CD97B22FD61CFF8DFC4C85C6F9E5FB43" box="[973,1014,1188,1212]" italics="true" pageId="9" pageNumber="287">E2a</emphasis>
as strongly bent ridge with anterior part running mediad and almost reaching borders of mentotectum). Sub-parastigmatic tubercle 
<emphasis id="CD97B22FD61CFF8DFCBB846CF949FADA" box="[826,858,1294,1317]" italics="true" pageId="9" pageNumber="287">Ua</emphasis>
present lateral to ventrolateral ridge, in area between tubercles 
<emphasis id="CD97B22FD61CFF8DFB828453FE33FAB7" box="[1027,1056,1329,1352]" italics="true" pageId="9" pageNumber="287">Va</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFBE48453FE93FAB7" box="[1125,1152,1329,1352]" italics="true" pageId="9" pageNumber="287">Sa</emphasis>
, very well developed, larger than 
<emphasis id="CD97B22FD61CFF8DFC468436F9F7FA94" box="[967,996,1364,1387]" italics="true" pageId="9" pageNumber="287">Va</emphasis>
, spiniform, with acute or blunt tip. Anterior ventrolateral tubercle (
<emphasis id="CD97B22FD61CFF8DFB388415FEF4FA71" box="[1209,1255,1399,1422]" italics="true" pageId="9" pageNumber="287">VLa</emphasis>
) present anterior to 
<emphasis id="CD97B22FD61CFF8DFC1584F8F9A7FA4E" box="[916,948,1434,1457]" italics="true" pageId="9" pageNumber="287">Ua</emphasis>
, shifted more laterally so its tip could even slightly overlook margins of prodorsum, 
<emphasis id="CD97B22FD61CFF8DFAE484DFFF87FA2B" box="[1381,1428,1469,1492]" italics="true" pageId="9" pageNumber="287">VLp</emphasis>
absent. Epimeral setation with additional setae on epimeres I and II, epimeral setal formula 4-3-3-4, with setae differing in shape and size. All setae setiform, most of them smooth, only setae 
<emphasis id="CD97B22FD61CFF8DFAA4872BFF2CF99E" box="[1317,1343,1609,1633]" italics="true" pageId="9" pageNumber="287">3b</emphasis>
, 
<emphasis id="CD97B22FD61CFF8DFAD3872BFF7AF99E" box="[1362,1385,1609,1633]" italics="true" pageId="9" pageNumber="287">3c</emphasis>
, 
<emphasis id="CD97B22FD61CFF8DFAFC872BFF86F99E" box="[1405,1429,1609,1633]" italics="true" pageId="9" pageNumber="287">4c</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFCF0870EF99FF97B" box="[881,908,1644,1668]" italics="true" pageId="9" pageNumber="287">4d</emphasis>
are rough on surface and usually strongly bent. Setae 
<emphasis id="CD97B22FD61CFF8DFC4E87EDF9FBF958" box="[975,1000,1679,1703]" italics="true" pageId="9" pageNumber="287">1b</emphasis>
, 
<emphasis id="CD97B22FD61CFF8DFC7B87EDFE07F958" box="[1018,1044,1679,1703]" italics="true" pageId="9" pageNumber="287">3b</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFBD487EDFE7DF958" box="[1109,1134,1679,1703]" italics="true" pageId="9" pageNumber="287">4b</emphasis>
longest, setae 
<emphasis id="CD97B22FD61CFF8DFAA587EDFF2EF958" box="[1316,1341,1679,1703]" italics="true" pageId="9" pageNumber="287">1a</emphasis>
, 
<emphasis id="CD97B22FD61CFF8DFACE87EDFF79F958" box="[1359,1386,1679,1703]" italics="true" pageId="9" pageNumber="287">1d</emphasis>
, 
<emphasis id="CD97B22FD61CFF8DFAFD87EDFF86F958" box="[1404,1429,1679,1703]" italics="true" pageId="9" pageNumber="287">2a</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFCF087D0F998F935" box="[881,907,1714,1738]" italics="true" pageId="9" pageNumber="287">3a</emphasis>
shortest. Epimeral seta 
<emphasis id="CD97B22FD61CFF8DFB3A87D0FEC7F935" box="[1211,1236,1714,1738]" italics="true" pageId="9" pageNumber="287">1c</emphasis>
inserted anteriorly on ventrolateral ridge, setae 
<emphasis id="CD97B22FD61CFF8DFB4187B7FEC9F912" box="[1216,1242,1749,1773]" italics="true" pageId="9" pageNumber="287">3b</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFA9487B7FF3DF912" box="[1301,1326,1749,1773]" italics="true" pageId="9" pageNumber="287">4b</emphasis>
inserted at the base of tubercle 
<emphasis id="CD97B22FD61CFF8DFBDF879BFE6DF8EF" box="[1118,1150,1785,1808]" italics="true" pageId="9" pageNumber="287">Vp</emphasis>
. Additional epimeral seta 
<emphasis id="CD97B22FD61CFF8DFCF08679F99FF8CC" box="[881,908,1819,1843]" italics="true" pageId="9" pageNumber="287">1d</emphasis>
very small, inserted on distinct tubercle on the edge of the body above anterior part of ventrolateral ridge, behind acetabulum I. Additional epimeral setae 
<emphasis id="CD97B22FD61CFF8DFC7186E7FE1AF862" box="[1008,1033,1925,1949]" italics="true" pageId="9" pageNumber="287">2a</emphasis>
and 
<emphasis id="CD97B22FD61CFF8DFBC886E6FE70F863" box="[1097,1123,1924,1948]" italics="true" pageId="9" pageNumber="287">2b</emphasis>
inserted in area between tubercles 
<emphasis id="CD97B22FD61FFF8EFE9083A9FB28FD1C" box="[273,315,715,739]" italics="true" pageId="10" pageNumber="288">E2p</emphasis>
, 
<emphasis id="CD97B22FD61FFF8EFECA83AEFB78FD1C" box="[331,363,716,739]" italics="true" pageId="10" pageNumber="288">Ua</emphasis>
and 
<emphasis id="CD97B22FD61FFF8EFE2883AEFBC4FD1C" box="[425,471,716,739]" italics="true" pageId="10" pageNumber="288">VLa</emphasis>
. Muscle sigillae present in most of epimeral area. Discidium (
<emphasis id="CD97B22FD61FFF8EFDD0838CF863FCF9" box="[593,624,750,774]" italics="true" pageId="10" pageNumber="288">dis</emphasis>
) triangular, dentiform, with blunt tip, rather short, not reaching laterally as far as margins of acetabulum IV.
</paragraph>
<caption id="AB9C3EB5D61FFF8EFD948056FE0CFEB9" box="[533,1055,307,327]" pageId="10" pageNumber="288">
<paragraph id="FF5C6E3DD61FFF8EFD948056FE0CFEB9" blockId="10.[533,1055,307,327]" box="[533,1055,307,327]" pageId="10" pageNumber="288">
TABLE 2: Homology of leg setae of 
<taxonomicName id="38E315BED61FFF8EFCDE8056F9F6FEB9" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[863,997,307,326]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="288" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD61FFF8EFCDE8056F9F6FEB9" box="[863,997,307,326]" italics="true" pageId="10" pageNumber="288">Damaeus lupus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6A40F54D61FFF8EFC6A8051FE0CFEB9" box="[1003,1055,307,326]" pageId="10" pageNumber="288" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="FF5C6E3DD61FFF8EFE268012FE00FDA9" pageId="10" pageNumber="288">
<table id="8DE39C9DD61F007BFF3F8012FF87FDA7" box="[190,1428,368,600]" gridcols="5" gridrows="7" pageId="10" pageNumber="288">
<tr id="41D36C7FD61F007BFF3F8012FF87FE7F" box="[190,1428,368,384]" gridrow="0" pageId="10" pageNumber="288" rowspan-0="1">
<th id="02020503D61F007BFEB38012F82DFE7F" box="[306,574,368,384]" gridcol="1" gridrow="0" pageId="10" pageNumber="288">Leg I</th>
<th id="02020503D61F007BFDE18012F957FE7F" box="[608,836,368,384]" gridcol="2" gridrow="0" pageId="10" pageNumber="288">Leg II</th>
<th id="02020503D61F007BFCF08012FE7CFE7F" box="[881,1135,368,384]" gridcol="3" gridrow="0" pageId="10" pageNumber="288">Leg III</th>
<th id="02020503D61F007BFB028012FF87FE7F" box="[1155,1428,368,384]" gridcol="4" gridrow="0" pageId="10" pageNumber="288">Leg IV</th>
</tr>
<tr id="41D36C7FD61F007BFF3F80E8FF87FE64" box="[190,1428,394,411]" gridrow="1" pageId="10" pageNumber="288">
<th id="02020503D61F007BFF3F80E8FB00FE64" box="[190,275,394,411]" gridcol="0" gridrow="1" pageId="10" pageNumber="288">Trochanter</th>
<td id="02020503D61F007BFEB380E8F82DFE64" box="[306,574,394,411]" gridcol="1" gridrow="1" pageId="10" pageNumber="288">v’</td>
<td id="02020503D61F007BFDE180E8F957FE64" box="[608,836,394,411]" gridcol="2" gridrow="1" pageId="10" pageNumber="288">v’</td>
<td id="02020503D61F007BFCF080E8FE7CFE64" box="[881,1135,394,411]" gridcol="3" gridrow="1" pageId="10" pageNumber="288">v’, l’</td>
<td id="02020503D61F007BFB0280E8FF87FE64" box="[1155,1428,394,411]" gridcol="4" gridrow="1" pageId="10" pageNumber="288">v’</td>
</tr>
<tr id="41D36C7FD61F007BFF3F80D0FF87FE3B" box="[190,1428,434,452]" gridrow="2" pageId="10" pageNumber="288">
<th id="02020503D61F007BFF3F80D0FB00FE3B" box="[190,275,434,452]" gridcol="0" gridrow="2" pageId="10" pageNumber="288">Femur</th>
<td id="02020503D61F007BFEB380D0F82DFE3B" box="[306,574,434,452]" gridcol="1" gridrow="2" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFEEB80D0FBDEFE3D" box="[362,461,434,451]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFEEB80D0FB8BFE3C" bold="true" box="[362,408,434,451]" italics="true" pageId="10" pageNumber="288">d, bv”</emphasis>
, (l), (v
</emphasis>
<emphasis id="CD97B22FD61FFF8EFE5180D9FBC5FE3B" box="[464,470,443,452]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFE5B80D0FBE1FE3C" box="[474,498,434,451]" italics="true" pageId="10" pageNumber="288">), v</emphasis>
<emphasis id="CD97B22FD61FFF8EFE7480D9FBE8FE3B" box="[501,507,443,452]" italics="true" pageId="10" pageNumber="288">2</emphasis>
<emphasis id="CD97B22FD61FFF8EFE7F80D0F814FE3D" box="[510,519,434,450]" italics="true" pageId="10" pageNumber="288">”</emphasis>
</td>
<td id="02020503D61F007BFDE180D0F957FE3B" box="[608,836,434,452]" gridcol="2" gridrow="2" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFD1E80D0F91AFE3C" box="[671,777,434,451]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFD1E80D0F8C1FE3C" bold="true" box="[671,722,434,451]" italics="true" pageId="10" pageNumber="288">d, bv”</emphasis>
, v”, (l)
</emphasis>
</td>
<td id="02020503D61F007BFCF080D0FE7CFE3B" box="[881,1135,434,452]" gridcol="3" gridrow="2" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFC4480D0FE08FE3C" box="[965,1051,434,451]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFC4480D0F9FDFE3C" bold="true" box="[965,1006,434,451]" italics="true" pageId="10" pageNumber="288">d, ev’</emphasis>
, v’, l’
</emphasis>
</td>
<td id="02020503D61F007BFB0280D0FF87FE3B" box="[1155,1428,434,452]" gridcol="4" gridrow="2" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFB6580D0FF20FE3C" box="[1252,1331,434,451]" italics="true" pageId="10" pageNumber="288">ev’, v’, l’, d</emphasis>
</td>
</tr>
<tr id="41D36C7FD61F007BFF3F80B8FF87FE13" box="[190,1428,474,492]" gridrow="3" pageId="10" pageNumber="288">
<th id="02020503D61F007BFF3F80B8FB00FE13" box="[190,275,474,492]" gridcol="0" gridrow="3" pageId="10" pageNumber="288">Genu</th>
<td id="02020503D61F007BFEB380B8F82DFE13" box="[306,574,474,492]" gridcol="1" gridrow="3" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFE1480B8FBCEFE14" box="[405,477,474,491]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFE1480B8FBD1FE14" bold="true" box="[405,450,474,491]" italics="true" pageId="10" pageNumber="288">dσ, (l)</emphasis>
, v’
</emphasis>
</td>
<td id="02020503D61F007BFDE180B8F957FE13" box="[608,836,474,492]" gridcol="2" gridrow="3" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFD3180B8F8EAFE14" box="[688,761,474,491]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFD3180B8F8CDFE14" bold="true" box="[688,734,474,491]" italics="true" pageId="10" pageNumber="288">dσ, (l)</emphasis>
, v’
</emphasis>
</td>
<td id="02020503D61F007BFCF080B8FE7CFE13" box="[881,1135,474,492]" gridcol="3" gridrow="3" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFC5380B9FE1EFE13" box="[978,1037,475,492]" italics="true" pageId="10" pageNumber="288">v’, l’, dσ</emphasis>
</td>
<td id="02020503D61F007BFB0280B8FF87FE13" box="[1155,1428,474,492]" gridcol="4" gridrow="3" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFB7280B9FF36FE13" box="[1267,1317,475,492]" italics="true" pageId="10" pageNumber="288">v’, l’, d</emphasis>
</td>
</tr>
<tr id="41D36C7FD61F007BFF3F8361FF87FDEA" box="[190,1428,515,533]" gridrow="4" pageId="10" pageNumber="288">
<th id="02020503D61F007BFF3F8361FB00FDEA" box="[190,275,515,533]" gridcol="0" gridrow="4" pageId="10" pageNumber="288">Tibia</th>
<td id="02020503D61F007BFEB38361F82DFDEA" box="[306,574,515,533]" gridcol="1" gridrow="4" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFEE88361FB96FDEC" bold="true" box="[361,389,515,532]" italics="true" pageId="10" pageNumber="288">d φ</emphasis>
<emphasis id="CD97B22FD61FFF8EFE09836EFB9DFDEA" bold="true" box="[392,398,524,533]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFE108361FBACFDEC" bold="true" box="[401,447,515,532]" italics="true" pageId="10" pageNumber="288">, (l), v</emphasis>
<emphasis id="CD97B22FD61FFF8EFE43836EFBDBFDEA" bold="true" box="[450,456,524,533]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFE4A8366FBCEFDEB" box="[459,477,516,532]" italics="true" pageId="10" pageNumber="288">, v</emphasis>
<emphasis id="CD97B22FD61FFF8EFE60836EFBF4FDEA" box="[481,487,524,533]" italics="true" pageId="10" pageNumber="288">2</emphasis>
<emphasis id="CD97B22FD61FFF8EFE6B8366FBECFDEB" box="[490,511,516,532]" italics="true" pageId="10" pageNumber="288">, φ</emphasis>
<emphasis id="CD97B22FD61FFF8EFD83836EF81BFDEA" box="[514,520,524,533]" italics="true" pageId="10" pageNumber="288">2</emphasis>
</td>
<td id="02020503D61F007BFDE18361F957FDEA" box="[608,836,515,533]" gridcol="2" gridrow="4" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFD288361F913FDEC" box="[681,768,515,532]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFD288361F8F0FDEB" bold="true" box="[681,739,515,532]" italics="true" pageId="10" pageNumber="288">d φ, (l),</emphasis>
(v)
</emphasis>
</td>
<td id="02020503D61F007BFCF08361FE7CFDEA" box="[881,1135,515,533]" gridcol="3" gridrow="4" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFC498361FE0BFDEC" box="[968,1048,515,532]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFC498361F9E4FDEB" bold="true" box="[968,1015,515,532]" italics="true" pageId="10" pageNumber="288">d φ, l’</emphasis>
, (v)
</emphasis>
</td>
<td id="02020503D61F007BFB028361FF87FDEA" box="[1155,1428,515,533]" gridcol="4" gridrow="4" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFB6F8361FF39FDEB" box="[1262,1322,515,532]" italics="true" pageId="10" pageNumber="288">(v), l’, φ</emphasis>
</td>
</tr>
<tr id="41D36C7FD61F007BFF3F8349FF87FDC1" box="[190,1428,555,574]" gridrow="5" pageId="10" pageNumber="288">
<th id="02020503D61F007BFF3F8349FB00FDC1" box="[190,275,555,574]" gridcol="0" gridrow="5" pageId="10" pageNumber="288">Tarsus</th>
<td id="02020503D61F007BFEB38349F82DFDC1" box="[306,574,555,574]" gridcol="1" gridrow="5" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFEB38349FB61FDC3" bold="true" box="[306,370,555,572]" italics="true" pageId="10" pageNumber="288">(ft), e, ω</emphasis>
<emphasis id="CD97B22FD61FFF8EFEF78357FB6FFDC1" bold="true" box="[374,380,565,574]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFEFE834EFB84FDC3" box="[383,407,556,572]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFEFE834EFB90FDC3" bold="true" box="[383,387,556,572]" italics="true" pageId="10" pageNumber="288">,</emphasis>
ω
</emphasis>
<emphasis id="CD97B22FD61FFF8EFE1A8357FBB2FDC1" box="[411,417,565,574]" italics="true" pageId="10" pageNumber="288">2</emphasis>
<emphasis id="CD97B22FD61FFF8EFE24834EF82DFDC3" box="[421,574,556,573]" italics="true" pageId="10" pageNumber="288">
, 
<emphasis id="CD97B22FD61FFF8EFE2F834EF82DFDC3" bold="true" box="[430,574,556,573]" italics="true" pageId="10" pageNumber="288">(tc), (p), (u), (a), s,</emphasis>
</emphasis>
</td>
<td id="02020503D61F007BFDE18349F957FDC1" box="[608,836,555,574]" gridcol="2" gridrow="5" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFDE18349F89CFDC3" bold="true" box="[608,655,555,572]" italics="true" pageId="10" pageNumber="288">(ft), ω</emphasis>
<emphasis id="CD97B22FD61FFF8EFD128357F88AFDC1" bold="true" box="[659,665,565,574]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFD1D834EF957FDC3" bold="true" box="[668,836,556,573]" italics="true" pageId="10" pageNumber="288">, (tc), (p), (u), (a), s,</emphasis>
</td>
<td id="02020503D61F007BFCF08349FE7CFDC1" box="[881,1135,555,574]" gridcol="3" gridrow="5" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFCF08349FE7CFDC3" box="[881,1135,555,573]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFCF08349FE5FFDC2" bold="true" box="[881,1100,555,573]" italics="true" pageId="10" pageNumber="288">(ft), (tc), (p), (u), (a), s, (pv),</emphasis>
(it),
</emphasis>
</td>
<td id="02020503D61F007BFB028349FF87FDC1" box="[1155,1428,555,574]" gridcol="4" gridrow="5" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFB02834EFF4DFDC3" box="[1155,1374,556,573]" italics="true" pageId="10" pageNumber="288">ft”, (tc), (p), (u), (a), s, (pv), (v</emphasis>
<emphasis id="CD97B22FD61FFF8EFAE08357FF74FDC1" box="[1377,1383,565,574]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFAEA834EFF91FDC3" box="[1387,1410,556,572]" italics="true" pageId="10" pageNumber="288">), v</emphasis>
<emphasis id="CD97B22FD61FFF8EFA078357FF9FFDC1" box="[1414,1420,565,574]" italics="true" pageId="10" pageNumber="288">2</emphasis>
<emphasis id="CD97B22FD61FFF8EFA0E834EFF87FDC3" box="[1423,1428,556,572]" italics="true" pageId="10" pageNumber="288">’</emphasis>
</td>
</tr>
<tr id="41D36C7FD61F007BFF3F8324FF87FDA7" box="[190,1428,582,600]" gridrow="6" pageId="10" pageNumber="288" rowspan-0="1" rowspan-4="1">
<td id="02020503D61F007BFEB38324F82DFDA7" box="[306,574,582,600]" gridcol="1" gridrow="6" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFEDC8325FBC8FDA7" box="[349,475,583,600]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFEDC8325FBB6FDA7" bold="true" box="[349,421,583,600]" italics="true" pageId="10" pageNumber="288">(pv), (pl),</emphasis>
(it), (v
</emphasis>
<emphasis id="CD97B22FD61FFF8EFE5F832CFBF7FDA8" box="[478,484,590,599]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFE698325F807FDA7" box="[488,532,583,600]" italics="true" pageId="10" pageNumber="288">), (v2)</emphasis>
</td>
<td id="02020503D61F007BFDE18324F957FDA7" box="[608,836,582,600]" gridcol="2" gridrow="6" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFDF08324F8F1FDA8" box="[625,738,582,599]" italics="true" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFDF08324F8BFFDA8" bold="true" box="[625,684,582,599]" italics="true" pageId="10" pageNumber="288">(pv),?,</emphasis>
(it), (v
</emphasis>
<emphasis id="CD97B22FD61FFF8EFD67832CF8FFFDA8" box="[742,748,590,599]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFD6E8324F914FDA8" box="[751,775,582,599]" italics="true" pageId="10" pageNumber="288">), v</emphasis>
<emphasis id="CD97B22FD61FFF8EFC8B832CF903FDA8" box="[778,784,590,599]" italics="true" pageId="10" pageNumber="288">2</emphasis>
<emphasis id="CD97B22FD61FFF8EFC958324F93EFDA8" box="[788,813,582,599]" italics="true" pageId="10" pageNumber="288">’, ω</emphasis>
<emphasis id="CD97B22FD61FFF8EFCB0832CF924FDA8" box="[817,823,590,599]" italics="true" pageId="10" pageNumber="288">2</emphasis>
</td>
<td id="02020503D61F007BFCF08324FE7CFDA7" box="[881,1135,582,600]" gridcol="3" gridrow="6" pageId="10" pageNumber="288">
<emphasis id="CD97B22FD61FFF8EFC4C8324F9CFFDA9" box="[973,988,582,598]" italics="true" pageId="10" pageNumber="288">(v</emphasis>
<emphasis id="CD97B22FD61FFF8EFC61832CF9F5FDA8" box="[992,998,590,599]" italics="true" pageId="10" pageNumber="288">1</emphasis>
<emphasis id="CD97B22FD61FFF8EFC688324FE12FDA9" box="[1001,1025,582,598]" italics="true" pageId="10" pageNumber="288">), v</emphasis>
<emphasis id="CD97B22FD61FFF8EFB85832CFE19FDA8" box="[1028,1034,590,599]" italics="true" pageId="10" pageNumber="288">2</emphasis>
’
</td>
</tr>
</table>
</paragraph>
<paragraph id="FF5C6E3DD61FFF8EFF298315FF9DFD78" blockId="10.[168,1422,630,649]" box="[168,1422,630,649]" pageId="10" pageNumber="288">
<tableNote id="02056FB3D61FFF8EFF298315FF9DFD78" box="[168,1422,630,649]" pageId="10" pageNumber="288" targetBox="[190,1428,368,600]" targetPageId="10">Note: Larval setation is indicated in bold, underlined setae are lost in adult; question mark (?) indicates additional seta in larva which was difficult to homologize.</tableNote>
</paragraph>
<paragraph id="FF5C6E3DD61FFF8EFF448202F838F9ED" blockId="10.[159,762,715,1949]" pageId="10" pageNumber="288">
<location id="FA3C38E6D61FFF8EFF448202FBB1FC87" LSID="urn:lsid:plazi:treatment:774ADF2BD617FF8AFEA58492FBF4FB15:FA3C38E6D61FFF8EFF448202FBB1FC87" box="[197,418,864,888]" county="All" municipality="Short" name="Anogenital region" pageId="10" pageNumber="288">Anogenital region</location>
— 
<figureCitation id="67D872B8D61FFF8EFE5A8202F873FC87" box="[475,608,864,888]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="10" pageNumber="288">Figures 1B</figureCitation>
, 
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<figureCitation id="67D872B8D61FFF8EFDF58202F882FC87" box="[628,657,864,888]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="10" pageNumber="288">2F</figureCitation>
.
</specimenCount>
<location id="FA3C38E6D61FFF8EFD258202F8E9FC87" LSID="urn:lsid:plazi:treatment:774ADF2BD617FF8AFEA58492FBF4FB15:FA3C38E6D61FFF8EFD258202F8E9FC87" box="[676,762,864,888]" county="All" municipality="Short" name="Genital" pageId="10" pageNumber="288">Genital</location>
opening broader and larger than anal opening, framed by thicker cuticle. 
<location id="FA3C38E6D61FFF8EFE6082C4F829FC41" LSID="urn:lsid:plazi:treatment:774ADF2BD617FF8AFEA58492FBF4FB15:FA3C38E6D61FFF8EFE6082C4F829FC41" box="[481,570,934,958]" county="All" municipality="Short" name="Preanal" pageId="10" pageNumber="288">Preanal</location>
sclerite appearing round, in detail horseshoe-shaped. 
<collectingMunicipality id="1F38F447D61FFF8EFDFF82ABF8AEFC1E" box="[638,701,969,993]" pageId="10" pageNumber="288">Short</collectingMunicipality>
concave thickening (ridge) present caudally, posterior to anal opening. 
<collectingCounty id="163D16B1D61FFF8EFEF4856DFB8AFBD8" box="[373,409,1039,1063]" pageId="10" pageNumber="288">All</collectingCounty>
setae simple setiform and of similar length, except of anal setae in usual numbers: 
<emphasis id="CD97B22FD61FFF8EFF5E8537FB72FB92" box="[223,353,1109,1133]" italics="true" pageId="10" pageNumber="288">
<quantity id="381BC3D8D61FFF8EFF5E8537FAEFFB92" box="[223,252,1109,1133]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="10" pageNumber="288" unit="g" value="6.0">6g</quantity>
, 1ag, 
<specimenCount id="E9E5A5B4D61FFF8EFEBB8537FB72FB92" box="[314,353,1109,1133]" pageId="10" pageNumber="288" type="generic">3ad</specimenCount>
</emphasis>
. Anal plates with variable number of setae, 
<typeStatus id="2058D09FD61FFF8EFE84851AFB7EFB6F" box="[261,365,1144,1168]" pageId="10" pageNumber="288" type="holotype">holotype</typeStatus>
with additional setae, with 3 anal setae present (
<emphasis id="CD97B22FD61FFF8EFEC885FEFB62FB4B" box="[329,369,1180,1204]" italics="true" pageId="10" pageNumber="288">3an</emphasis>
); insertions of 4th anal seta (
<emphasis id="CD97B22FD61FFF8EFD4885FEF8E2FB4C" box="[713,753,1180,1203]" italics="true" pageId="10" pageNumber="288">anx</emphasis>
) observed unilaterally (
<figureCitation id="67D872B8D61FFF8EFE2785DCFBE1FB28" box="[422,498,1214,1239]" captionStart="FIGURE" captionStartId="4.[159,171,1937,1956]" captionTargetBox="[308,1323,351,1756]" captionTargetId="figure-2@4.[227,1361,302,1905]" captionTargetPageId="4" captionText="FIGURE 2: Details of Damaeus lupus n. sp. (adult, paratype): A – rostral part of prodorsum, dorsal view, B – rostral part, lateral view, C – sensillus and prodorsal setae, D – notogastral setae and spina adnatum, E – ventral ridges and tubercles in epimeral area, F – anogenital area, lateral view. Acronyms: abr – anterobothridial ridge, acos – anterior lamellar costula, anx – insertion of additional anal seta, bo – bothridium, pbr – postbothridial ridge, rcos – rostral costula, rop – rostral protuberance (scale bars: A-B, C-E = 100 µm, F = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667404" httpUri="https://zenodo.org/record/4667404/files/figure.png" pageId="10" pageNumber="288">Fig 2F</figureCitation>
). One 
<typeStatus id="2058D09FD61FFF8EFDBD85DDF8B6FB28" box="[572,677,1215,1239]" pageId="10" pageNumber="288" type="paratype">paratype</typeStatus>
unilaterally (on left valve in ventral view) with 3 setae and insertion of fourth seta as in 
<typeStatus id="2058D09FD61FFF8EFD838467F87EFAE2" box="[514,621,1285,1309]" pageId="10" pageNumber="288" type="holotype">holotype</typeStatus>
, with other anal plate bearing 2 setae. Two other 
<typeStatus id="2058D09FD61FFF8EFD04844AF8EAFABF" box="[645,761,1320,1344]" pageId="10" pageNumber="288" type="paratype">paratypes</typeStatus>
with normal number of anal setae (
<emphasis id="CD97B22FD61FFF8EFDD08429F86BFA9C" box="[593,632,1355,1379]" italics="true" pageId="10" pageNumber="288">2an</emphasis>
). Seta 
<emphasis id="CD97B22FD61FFF8EFD578429F8FCFA9C" box="[726,751,1355,1379]" italics="true" pageId="10" pageNumber="288">ad</emphasis>
<subScript id="63676C78D61FFF8EFD6E8436F8EBFA9A" attach="left" box="[751,760,1364,1381]" fontSize="7" pageId="10" pageNumber="288">1</subScript>
inserted in postanal area, seta 
<emphasis id="CD97B22FD61FFF8EFD98840CF821FA79" box="[537,562,1390,1414]" italics="true" pageId="10" pageNumber="288">ad</emphasis>
<subScript id="63676C78D61FFF8EFDB38415F828FA77" attach="left" box="[562,571,1399,1416]" fontSize="7" pageId="10" pageNumber="288">2</subScript>
and 
<emphasis id="CD97B22FD61FFF8EFD00840CF889FA79" box="[641,666,1390,1414]" italics="true" pageId="10" pageNumber="288">ad</emphasis>
<subScript id="63676C78D61FFF8EFD1B8415F8B0FA77" attach="left" box="[666,675,1399,1416]" fontSize="7" pageId="10" pageNumber="288">3</subScript>
lateral to anal opening. Lyrifissures 
<emphasis id="CD97B22FD61FFF8EFD8984F3F83BFA56" box="[520,552,1425,1449]" italics="true" pageId="10" pageNumber="288">iad</emphasis>
apoanal, oblique or almost horizontal, positioned lateral to anterolateral angles of ventral opening, closer to anal opening than insertions of 
<emphasis id="CD97B22FD61FFF8EFE1E8498FBABF9ED" box="[415,440,1530,1554]" italics="true" pageId="10" pageNumber="288">ad</emphasis>
<subScript id="63676C78D61FFF8EFE398766FBD2F9EA" attach="left" box="[440,449,1540,1557]" fontSize="7" pageId="10" pageNumber="288">3</subScript>
(
<figureCitation id="67D872B8D61FFF8EFE508498F80EF9ED" box="[465,541,1530,1554]" captionStart="FIGURE" captionStartId="3.[159,171,1296,1315]" captionTargetBox="[243,1435,324,1242]" captionTargetId="figure-6@3.[159,1520,302,1264]" captionTargetPageId="3" captionText="FIGURE 1: Damaeus lupus n. sp. (adult, paratype): A – dorsal view, B – ventral view. Acronyms: alp – anterolateral protuberance, abr – anterobothridial ridge, dis – discidium, pbr – postbothridial ridge, rop – rostral protuberance, s. a. – spina adnatum, tl – translamellar line, tpf – tectum of podocephalic fossa (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667402" httpUri="https://zenodo.org/record/4667402/files/figure.png" pageId="10" pageNumber="288">Fig 1B</figureCitation>
).
</paragraph>
<paragraph id="FF5C6E3DD61FFF8EFF448744F99DFAEB" blockId="10.[159,762,715,1949]" lastBlockId="10.[826,1429,715,1949]" pageId="10" pageNumber="288">
Legs — 
<figureCitation id="67D872B8D61FFF8EFEAA8744FB8AF9C1" box="[299,409,1574,1598]" captionStart="FIGURE" captionStartId="5.[159,171,1847,1866]" captionTargetBox="[286,1314,355,1768]" captionTargetId="figure-6@5.[265,1323,320,1816]" captionTargetPageId="5" captionText="FIGURE 3: Legs of Damaeus lupus n. sp. (adult, paratype): A – femur and genu of leg I, B – genu, tarsus and tibia of leg I, C – femur and genu of leg II, D – tibia and tarsus of leg II, E – trochanter, femur and genu of leg III, F – genu, tibia and tarsus of leg III, G – trochanter, femur and genu of leg IV, H – genu, tibia and tarsus of leg IV (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667408" httpUri="https://zenodo.org/record/4667408/files/figure.png" pageId="10" pageNumber="288">Figures 3</figureCitation>
A-H, 4A, 5A-D. Legs moderately long, both leg I (about 1,2 times) and leg IV (1,4-1,5 times) longer than ventral body length. Segments of legs I and II only slightly expanded, of leg IV-elongated and more slender. Leg setae generally short, dorsal and lateral setae usually stronger than ventral, thorn-like, slightly bent or curved with rather blunt tips, ventral setae more setiform, bent, with elongated and acute tips. Only few setae of tarsi and tibiae longer (around 
<quantity id="381BC3D8D61FFF8EFD8B8600F8B2F886" box="[522,673,1889,1914]" metricMagnitude="-1" metricUnit="m" metricValue="1.225" metricValueMax="1.35" metricValueMin="1.1" pageId="10" pageNumber="288" unit="mm" value="122.5" valueMax="135.0" valueMin="110.0">110 – 135 µm</quantity>
): tarsal 
<emphasis id="CD97B22FD61FFF8EFF1E86E7FAAFF862" box="[159,188,1925,1949]" italics="true" pageId="10" pageNumber="288">ft&quot;</emphasis>
and tibial 
<emphasis id="CD97B22FD61FFF8EFEB786E6FB56F863" box="[310,325,1924,1948]" italics="true" pageId="10" pageNumber="288">l’</emphasis>
and 
<emphasis id="CD97B22FD61FFF8EFEFC86E7FB81F863" box="[381,402,1925,1948]" italics="true" pageId="10" pageNumber="288">v’</emphasis>
of leg IV and tarsal 
<emphasis id="CD97B22FD61FFF8EFDF686E7F8ABF863" box="[631,696,1924,1948]" italics="true" pageId="10" pageNumber="288">tc and</emphasis>
tibial 
<emphasis id="CD97B22FD61FFF8EFCBB83A9F95AFD1C" box="[826,841,715,739]" italics="true" pageId="10" pageNumber="288">l’</emphasis>
and 
<emphasis id="CD97B22FD61FFF8EFC0583A9F98AFD1D" box="[900,921,715,738]" italics="true" pageId="10" pageNumber="288">v’</emphasis>
of leg I being the longest (
<figureCitation id="67D872B8D61FFF8EFB5383A9FF9BFD1C" box="[1234,1416,715,739]" captionStart="FIGURE" captionStartId="7.[159,171,1810,1829]" captionTargetBox="[227,1361,385,1778]" captionTargetId="figure-6@7.[227,1361,385,1778]" captionTargetPageId="7" captionText="FIGURE 5: Damaeus lupus n. sp. (adult, paratype): A – genu and tibia of leg I, B – tarsus of leg I, C – tibia and tarsus of leg II, D – tibia of leg IV." figureDoi="http://doi.org/10.5281/zenodo.4667414" httpUri="https://zenodo.org/record/4667414/files/figure.png" pageId="10" pageNumber="288">Figures 5A,B,D</figureCitation>
). Famulus short, straight, simple setiform. All solenidia also rather short, always shorter than longest seta of respective leg segment, genual solenidia fine, gently curved and slightly longer or subequal to accompanied seta 
<emphasis id="CD97B22FD61FFF8EFC778218FE10FC6D" box="[1014,1027,890,914]" italics="true" pageId="10" pageNumber="288">d</emphasis>
, tibial solenidia short, more or less straight, erect, short. Tarsal solenidia of leg I and II setiform, slightly bent and subequal in length to the other setae of the segment. Companion seta 
<emphasis id="CD97B22FD61FFF8EFCBB8564F954FBE1" box="[826,839,1030,1054]" italics="true" pageId="10" pageNumber="288">d</emphasis>
absent on all tibia, all tibial solenidia free and short. All solenidia of genua I-III with companion seta 
<emphasis id="CD97B22FD61FFF8EFCF0852FF96DFB9A" box="[881,894,1101,1125]" italics="true" pageId="10" pageNumber="288">d</emphasis>
. Both 
<emphasis id="CD97B22FD61FFF8EFC55852FF9F2FB9B" box="[980,993,1101,1124]" italics="true" pageId="10" pageNumber="288">v</emphasis>
<subScript id="63676C78D61FFF8EFC638534F9F8FB98" attach="left" box="[994,1003,1110,1127]" fontSize="7" pageId="10" pageNumber="288">2</subScript>
setae present on tarsus I. Setal formula of legs as follows (famulus included, solenidia in parentheses): leg I: 1-7-4[1]-4(2)-22(2), leg II: 1-5-4[1]-4(1)-18(2), leg III: 2-4-3[1]-3(1)-18, leg IV: 1- 4-3-3(1)-15. Homology of the setae is presented in table 2.
</paragraph>
<paragraph id="FF5C6E3DD61FFF8EFCE18447FEB4FA7C" blockId="10.[826,1429,715,1949]" pageId="10" pageNumber="288">
Ontogeny — Single individual of larva found together with adults (
<figureCitation id="67D872B8D61FFF8EFB9D842AFE9AFA9F" box="[1052,1161,1352,1376]" captionStart="FIGURE" captionStartId="11.[207,219,1296,1315]" captionTargetBox="[159,1520,302,1264]" captionTargetId="figure-6@11.[159,1520,302,1264]" captionTargetPageId="11" captionText="FIGURE 6: Damaeus lupus n. sp. (larva): A – dorsal view, B – ventral view, C – leg I, D – leg II, E – leg III (scale bars = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667416" httpUri="https://zenodo.org/record/4667416/files/figure.png" pageId="10" pageNumber="288">Figures 6</figureCitation>
, 
<figureCitation id="67D872B8D61FFF8EFB15842AFEB0FA9F" box="[1172,1187,1352,1376]" captionStart="FIGURE" captionStartId="12.[177,189,1805,1824]" captionTargetBox="[227,1361,418,1773]" captionTargetId="figure-2@12.[227,1361,418,1773]" captionTargetPageId="12" captionText="FIGURE 7: Damaeus lupus n. sp. (larva): A – dorsal view of the body, B – leg I and II, C – leg III, D – details of setae of opisthonotum." figureDoi="http://doi.org/10.5281/zenodo.4667418" httpUri="https://zenodo.org/record/4667418/files/figure.png" pageId="10" pageNumber="288">7</figureCitation>
), nymphs unknown. Description of larva as follows:
</paragraph>
<paragraph id="FF5C6E3DD61FFF8EFCE184F6F95DF9EA" blockId="10.[826,1429,715,1949]" pageId="10" pageNumber="288">
Dimensions — Body length 426, maximum width 240. Legs I and II longer than body (470 and 430, respectively), other measures are given in 
<tableCitation id="B2615B86D61FFF8EFAD784B8F95BF9EA" captionStart="TABLE" captionStartId="8.[159,172,308,327]" captionText="TABLE 1: Selected measures of Damaeus lupus n. sp." pageId="10" pageNumber="288">Table 1</tableCitation>
.
</paragraph>
<paragraph id="FF5C6E3DD61FFF8EFCE18744F974F862" blockId="10.[826,1429,715,1949]" pageId="10" pageNumber="288">
Integument — Body colour pale or white, with some parts (legs, mouthparts, apophyses) more sclerotised and slightly brownish, darker. Surface of body irregular, with transverse elevated parts around setal insertions, whole body covered with layer of transparent, mostly granular cerotegument, forming lines and small spots (
<figureCitation id="67D872B8D61FFF8EFB2B879AFF5EF8EF" box="[1194,1357,1784,1808]" captionStart="FIGURE" captionStartId="5.[159,171,1847,1866]" captionTargetBox="[286,1314,355,1768]" captionTargetId="figure-6@5.[265,1323,320,1816]" captionTargetPageId="5" captionText="FIGURE 3: Legs of Damaeus lupus n. sp. (adult, paratype): A – femur and genu of leg I, B – genu, tarsus and tibia of leg I, C – femur and genu of leg II, D – tibia and tarsus of leg II, E – trochanter, femur and genu of leg III, F – genu, tibia and tarsus of leg III, G – trochanter, femur and genu of leg IV, H – genu, tibia and tarsus of leg IV (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667408" httpUri="https://zenodo.org/record/4667408/files/figure.png" pageId="10" pageNumber="288">Figures 3A, B</figureCitation>
; Plate 3A – 
<emphasis id="CD97B22FD61FFF8EFCFB867EF988F8CC" box="[890,923,1820,1843]" italics="true" pageId="10" pageNumber="288">ctg</emphasis>
). Legs covered with more amorphous, almost transparent cerotegument with folds, which appear as irregular lines and nets (
<figureCitation id="67D872B8D61FFF8EFB718603FF72F885" box="[1264,1377,1889,1914]" captionStart="FIGURE" captionStartId="5.[159,171,1847,1866]" captionTargetBox="[286,1314,355,1768]" captionTargetId="figure-6@5.[265,1323,320,1816]" captionTargetPageId="5" captionText="FIGURE 3: Legs of Damaeus lupus n. sp. (adult, paratype): A – femur and genu of leg I, B – genu, tarsus and tibia of leg I, C – femur and genu of leg II, D – tibia and tarsus of leg II, E – trochanter, femur and genu of leg III, F – genu, tibia and tarsus of leg III, G – trochanter, femur and genu of leg IV, H – genu, tibia and tarsus of leg IV (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667408" httpUri="https://zenodo.org/record/4667408/files/figure.png" pageId="10" pageNumber="288">Figures 3</figureCitation>
C-E, 7B).
</paragraph>
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<paragraph id="FF5C6E3DD61EFF8FFF4E8472FF76FADC" blockId="11.[207,1381,1296,1315]" box="[207,1381,1296,1315]" pageId="11" pageNumber="289">
FIGURE 6: 
<taxonomicName id="38E315BED61EFF8FFEB58473FBA9FADC" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[308,442,1296,1315]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="289" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD61EFF8FFEB58473FBA9FADC" box="[308,442,1296,1315]" italics="true" pageId="11" pageNumber="289">Damaeus lupus</emphasis>
</taxonomicName>
<emphasis id="CD97B22FD61EFF8FFE418472FBE7FADC" bold="true" box="[448,500,1296,1315]" pageId="11" pageNumber="289">
<taxonomicNameLabel id="D6A40F54D61EFF8FFE418472FBE7FADC" box="[448,500,1296,1315]" pageId="11" pageNumber="289" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(larva): A – dorsal view, 
<emphasis id="CD97B22FD61EFF8FFD638472F8E3FADC" bold="true" box="[738,752,1296,1315]" pageId="11" pageNumber="289">B</emphasis>
– ventral view, C – leg I, D – leg II, E – leg III (scale bars = 200 µm).
</paragraph>
</caption>
<paragraph id="FF5C6E3DD61EFF89FF448406F8DAFD52" blockId="11.[159,762,1380,1930]" lastBlockId="13.[159,762,311,1948]" lastPageId="13" lastPageNumber="291" pageId="11" pageNumber="289">
Dorsal characters — Pronotum generally triangular, with deep incisions above acetabula I. Prodorsum laterally with broad, more sclerotised, S shaped ridges resembling lamellar ridges of adults of higher oribatids (
<emphasis id="CD97B22FD61EFF8FFE098493FBA3F9F6" box="[392,432,1521,1545]" italics="true" pageId="11" pageNumber="289">lam</emphasis>
– 
<figureCitation id="67D872B8D61EFF8FFE4B8493F851F9F6" box="[458,578,1521,1545]" captionStart="FIGURE" captionStartId="12.[177,189,1805,1824]" captionTargetBox="[227,1361,418,1773]" captionTargetId="figure-2@12.[227,1361,418,1773]" captionTargetPageId="12" captionText="FIGURE 7: Damaeus lupus n. sp. (larva): A – dorsal view of the body, B – leg I and II, C – leg III, D – details of setae of opisthonotum." figureDoi="http://doi.org/10.5281/zenodo.4667418" httpUri="https://zenodo.org/record/4667418/files/figure.png" pageId="11" pageNumber="289">Figure 7A</figureCitation>
). Anterolaterad to bothridia, between acetabula I and II, with irregularly triangular, pointed pedotectum-like projections. Lamellar setae very strong, rugose, regularly curved, inserted on distinct apophyses (
<figureCitation id="67D872B8D61EFF8FFD49871FFAD0F947" captionStart="FIGURE" captionStartId="11.[207,219,1296,1315]" captionTargetBox="[159,1520,302,1264]" captionTargetId="figure-6@11.[159,1520,302,1264]" captionTargetPageId="11" captionText="FIGURE 6: Damaeus lupus n. sp. (larva): A – dorsal view, B – ventral view, C – leg I, D – leg II, E – leg III (scale bars = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667416" httpUri="https://zenodo.org/record/4667416/files/figure.png" pageId="11" pageNumber="289">Figs 6A</figureCitation>
, 
<figureCitation id="67D872B8D61EFF8FFF5087C2FAE5F947" box="[209,246,1696,1720]" captionStart="FIGURE" captionStartId="12.[177,189,1805,1824]" captionTargetBox="[227,1361,418,1773]" captionTargetId="figure-2@12.[227,1361,418,1773]" captionTargetPageId="12" captionText="FIGURE 7: Damaeus lupus n. sp. (larva): A – dorsal view of the body, B – leg I and II, C – leg III, D – details of setae of opisthonotum." figureDoi="http://doi.org/10.5281/zenodo.4667418" httpUri="https://zenodo.org/record/4667418/files/figure.png" pageId="11" pageNumber="289">7A</figureCitation>
). Rostral setae shorter and much narrower than 
<emphasis id="CD97B22FD61EFF8FFF5A87A1FAFEF924" box="[219,237,1731,1755]" italics="true" pageId="11" pageNumber="289">le</emphasis>
, finely rugose as well and inserted on small apophyses. Interlamellar setae straight, setiform, rugose. Exobothridial setae shortest on pronotum, inserted on small apophyses, curved, rugose, with few longer hairs distally. Sensilli rather long, setiform, with sparse spinuli on the surface except of parts in close vicinity to bothridium opening. Bothridia rather small, almost circular, with expanded transparent funnel-like rims. Opisthonotum soft, irregularly hexagonal, with lateral margins more or less parallel in first two thirds of length. Standard set of 9 setae present dorsally (
<emphasis id="CD97B22FD61EFF8FFB318776F9B9F9B0" italics="true" pageId="11" pageNumber="289">c1, c2, c3, la, lm, lp, da, dm, dp</emphasis>
). All lateral and most of centronotal setae inserted on tubercles, however c 
<subScript id="63676C78D61EFF8FFB3C8701FED5F98B" attach="left" box="[1213,1222,1635,1652]" fontSize="7" pageId="11" pageNumber="289">2</subScript>
, 
<emphasis id="CD97B22FD61EFF8FFB568738FEE1F98D" box="[1239,1266,1626,1650]" italics="true" pageId="11" pageNumber="289">dp</emphasis>
, 
<emphasis id="CD97B22FD61EFF8FFA808738FF05F98D" box="[1281,1302,1626,1650]" italics="true" pageId="11" pageNumber="289">lp</emphasis>
and 
<emphasis id="CD97B22FD61EFF8FFAD58738FF7CF98D" box="[1364,1391,1626,1650]" italics="true" pageId="11" pageNumber="289">h1</emphasis>
on large, distinct apophyses. Opisthonotal setae very variable in size, colour and shape, but all well developed and visible. Three pairs of setae particularly long (
<emphasis id="CD97B22FD61EFF8FFC3B8784FE0FF901" box="[954,1052,1766,1790]" italics="true" pageId="11" pageNumber="289">c2, lp, h1</emphasis>
), of which 
<emphasis id="CD97B22FD61EFF8FFB1F8784FEA0F901" box="[1182,1203,1766,1790]" italics="true" pageId="11" pageNumber="289">lp</emphasis>
clearly the longest, longer than the whole body. These setae are darkly coloured, appearing as almost black, covered by dense spinuli on all they length except a very short base close to insertions (
<figureCitation id="67D872B8D61EFF8FFBD28610FEDFF874" box="[1107,1228,1906,1931]" captionStart="FIGURE" captionStartId="12.[177,189,1805,1824]" captionTargetBox="[227,1361,418,1773]" captionTargetId="figure-2@12.[227,1361,418,1773]" captionTargetPageId="12" captionText="FIGURE 7: Damaeus lupus n. sp. (larva): A – dorsal view of the body, B – leg I and II, C – leg III, D – details of setae of opisthonotum." figureDoi="http://doi.org/10.5281/zenodo.4667418" httpUri="https://zenodo.org/record/4667418/files/figure.png" pageId="11" pageNumber="289">Figure 7A</figureCitation>
). Setae 
<emphasis id="CD97B22FD61EFF8FFAA38610FF2EF875" box="[1314,1341,1906,1930]" italics="true" pageId="11" pageNumber="289">dp</emphasis>
strong, dark, setiform, of medium length, blunt distally, covered by dense spinuli. Three pairs of setae closest to plane of symmetry (
<emphasis id="CD97B22FD618FF89FE56801FF804FE6A" box="[471,535,381,405]" italics="true" pageId="13" pageNumber="291">c1, da</emphasis>
and 
<emphasis id="CD97B22FD618FF89FDD5801FF865FE6A" box="[596,630,381,405]" italics="true" pageId="13" pageNumber="291">dm</emphasis>
) with very specific shape, with proximal part dark and covered by spinuli, resembling setae 
<emphasis id="CD97B22FD618FF89FD8280A1F80DFE24" box="[515,542,451,475]" italics="true" pageId="13" pageNumber="291">dp</emphasis>
, but distally prolonged by transparent, hyaline extension, extended along end of dark part of seta and then narrowing and elongated, these elongated hyaline parts are not easily visible (
<figureCitation id="67D872B8D618FF89FEC7832DFBACFD98" box="[326,447,591,615]" captionStart="FIGURE" captionStartId="12.[177,189,1805,1824]" captionTargetBox="[227,1361,418,1773]" captionTargetId="figure-2@12.[227,1361,418,1773]" captionTargetPageId="12" captionText="FIGURE 7: Damaeus lupus n. sp. (larva): A – dorsal view of the body, B – leg I and II, C – leg III, D – details of setae of opisthonotum." figureDoi="http://doi.org/10.5281/zenodo.4667418" httpUri="https://zenodo.org/record/4667418/files/figure.png" pageId="13" pageNumber="291">Figure 7D</figureCitation>
, black arrows). Lateral setae (
<emphasis id="CD97B22FD618FF89FF508311FB21FD75" box="[209,306,626,651]" italics="true" pageId="13" pageNumber="291">c3, la, lm</emphasis>
) shorter, strongly curved or S- shaped, distally sometimes also with hyaline extension.
</paragraph>
<caption id="AB9C3EB5D619FF88FF30866FFF97F8E0" ID-DOI="http://doi.org/10.5281/zenodo.4667418" ID-Zenodo-Dep="4667418" box="[177,1412,1804,1824]" httpUri="https://zenodo.org/record/4667418/files/figure.png" pageId="12" pageNumber="290" startId="12.[177,189,1805,1824]" targetBox="[227,1361,418,1773]" targetPageId="12">
<paragraph id="FF5C6E3DD619FF88FF30866FFF97F8E0" blockId="12.[177,1412,1804,1824]" box="[177,1412,1804,1824]" pageId="12" pageNumber="290">
FIGURE 7: 
<taxonomicName id="38E315BED619FF88FE97866FFB8FF8E0" authority="Miko, 2016" authorityName="Miko" authorityYear="2016" box="[278,412,1804,1823]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="290" phylum="Arthropoda" rank="species" species="lupus" status="sp. nov.">
<emphasis id="CD97B22FD619FF88FE97866FFB8FF8E0" box="[278,412,1804,1823]" italics="true" pageId="12" pageNumber="290">Damaeus lupus</emphasis>
</taxonomicName>
<emphasis id="CD97B22FD619FF88FE20866EFBC5F8E0" bold="true" box="[417,470,1804,1823]" pageId="12" pageNumber="290">
<taxonomicNameLabel id="D6A40F54D619FF88FE20866EFBC5F8E0" box="[417,470,1804,1823]" pageId="12" pageNumber="290" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(larva): A – dorsal view of the body, B – leg I and II, C – leg III, D – details of setae of opisthonotum.
</paragraph>
</caption>
<paragraph id="FF5C6E3DD618FF89FF4483A6F862FBE7" blockId="13.[159,762,311,1948]" pageId="13" pageNumber="291">
Ventral characters — 
<figureCitation id="67D872B8D618FF89FE6F83A6F85AFD22" box="[494,585,708,733]" captionStart="FIGURE" captionStartId="5.[159,171,1847,1866]" captionTargetBox="[286,1314,355,1768]" captionTargetId="figure-6@5.[265,1323,320,1816]" captionTargetPageId="5" captionText="FIGURE 3: Legs of Damaeus lupus n. sp. (adult, paratype): A – femur and genu of leg I, B – genu, tarsus and tibia of leg I, C – femur and genu of leg II, D – tibia and tarsus of leg II, E – trochanter, femur and genu of leg III, F – genu, tibia and tarsus of leg III, G – trochanter, femur and genu of leg IV, H – genu, tibia and tarsus of leg IV (scale bar = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667408" httpUri="https://zenodo.org/record/4667408/files/figure.png" pageId="13" pageNumber="291">Fig 3B</figureCitation>
. Gnathosoma and epimeral region without particular characters, epimeral setal formula 2-1-2, ClaparØde’s organ (
<emphasis id="CD97B22FD618FF89FD598269F8E2FCDC" box="[728,753,779,803]" italics="true" pageId="13" pageNumber="291">Cl</emphasis>
) well developed, distinctly rounded distally. Paraproct without setae or setal insertions. Setae 
<emphasis id="CD97B22FD618FF89FD258233F8ACFC96" box="[676,703,849,873]" italics="true" pageId="13" pageNumber="291">h1</emphasis>
very long, inserted on long and distinct apophyses, setae 
<emphasis id="CD97B22FD618FF89FF1E82F5FAAAFC50" box="[159,185,919,943]" italics="true" pageId="13" pageNumber="291">h2</emphasis>
short, curved, inserted on distinct tubercles, setae 
<emphasis id="CD97B22FD618FF89FF4B82D8FAF7FC2D" box="[202,228,954,978]" italics="true" pageId="13" pageNumber="291">h3</emphasis>
absent, present only as insertion points. All lyrifissures (
<emphasis id="CD97B22FD618FF89FEAE82BFFB9DFC0A" box="[303,398,989,1013]" italics="true" pageId="13" pageNumber="291">ih, im, ip</emphasis>
) as well as opisthosomal gland opening (
<emphasis id="CD97B22FD618FF89FE8E8562FB3CFBE7" box="[271,303,1024,1048]" italics="true" pageId="13" pageNumber="291">gla</emphasis>
) present in usual locations.
</paragraph>
<paragraph id="FF5C6E3DD618FF89FF44854DFB91F8B5" blockId="13.[159,762,311,1948]" pageId="13" pageNumber="291">
Legs — 
<figureCitation id="67D872B8D618FF89FEA5854DFB9CFBB8" box="[292,399,1071,1095]" captionStart="FIGURE" captionStartId="11.[207,219,1296,1315]" captionTargetBox="[159,1520,302,1264]" captionTargetId="figure-6@11.[159,1520,302,1264]" captionTargetPageId="11" captionText="FIGURE 6: Damaeus lupus n. sp. (larva): A – dorsal view, B – ventral view, C – leg I, D – leg II, E – leg III (scale bars = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667416" httpUri="https://zenodo.org/record/4667416/files/figure.png" pageId="13" pageNumber="291">Figures 6</figureCitation>
C-E, 7B-C. Legs with usual larval setation, leg formula as follows (famulus not included, solenidia in parentheses). Leg I: 0-2-3(1)- 4(1)-15(1); leg II: 0-2-3(1)-3(1)-14(1), leg III: 0-2-2(1)- 3(1)-13. Homology of setae indicated on 
<figureCitation id="67D872B8D618FF89FD2385D9FABFFB09" captionStart="FIGURE" captionStartId="11.[207,219,1296,1315]" captionTargetBox="[159,1520,302,1264]" captionTargetId="figure-6@11.[159,1520,302,1264]" captionTargetPageId="11" captionText="FIGURE 6: Damaeus lupus n. sp. (larva): A – dorsal view, B – ventral view, C – leg I, D – leg II, E – leg III (scale bars = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667416" httpUri="https://zenodo.org/record/4667416/files/figure.png" pageId="13" pageNumber="291">Figures 6</figureCitation>
C-E, 7B-C, one additional seta observed on tarsus of leg II behind seta 
<emphasis id="CD97B22FD618FF89FE1E8460FBA9FAE6" box="[415,442,1282,1305]" italics="true" pageId="13" pageNumber="291">tc’</emphasis>
(indicated by? in 
<figureCitation id="67D872B8D618FF89FD2C8463FAD0FAC3" captionStart="FIGURE" captionStartId="11.[207,219,1296,1315]" captionTargetBox="[159,1520,302,1264]" captionTargetId="figure-6@11.[159,1520,302,1264]" captionTargetPageId="11" captionText="FIGURE 6: Damaeus lupus n. sp. (larva): A – dorsal view, B – ventral view, C – leg I, D – leg II, E – leg III (scale bars = 200 µm)." figureDoi="http://doi.org/10.5281/zenodo.4667416" httpUri="https://zenodo.org/record/4667416/files/figure.png" pageId="13" pageNumber="291">Figure 6D</figureCitation>
). Distal setae of tarsi simple setiform, rest of setae differently shaped. Setae 
<emphasis id="CD97B22FD618FF89FE7A8425F81BFAA0" box="[507,520,1351,1375]" italics="true" pageId="13" pageNumber="291">d</emphasis>
on tibia I, 
<emphasis id="CD97B22FD618FF89FD0A842AF8B0FA9F" box="[651,675,1352,1376]" italics="true" pageId="13" pageNumber="291">ft’</emphasis>
on tarsus I and 
<emphasis id="CD97B22FD618FF89FE908409FB3DFA7C" box="[273,302,1387,1411]" italics="true" pageId="13" pageNumber="291">ft&quot;</emphasis>
on tarsi II and III long, strong, straight and pointed, at least partly rugose. Dorsal setae of femora curved, very strong, thick, with dense spinuli. Other setae of proximal part of leg (femora, genua, partly also tibiae) also thickened and covered by spinuli, but usually short and less curved, ventral setae narrower and with hairs or smooth. Famulus (
<emphasis id="CD97B22FD618FF89FE958703FB0DF987" box="[276,286,1633,1656]" italics="true" pageId="13" pageNumber="291">e</emphasis>
) sunken in small sclerotised cup. Solenidia of different size and shape, solenidia 
<emphasis id="CD97B22FD618FF89FD0F87E1F88CF964" box="[654,671,1667,1691]" italics="true" pageId="13" pageNumber="291">φ</emphasis>
and 
<emphasis id="CD97B22FD618FF89FD5C87E1F8EBF961" box="[733,760,1667,1694]" italics="true" pageId="13" pageNumber="291">
ω 
<subScript id="63676C78D618FF89FD6E87EFF8EBF961" attach="left" box="[751,760,1677,1694]" fontSize="7" pageId="13" pageNumber="291">1</subScript>
</emphasis>
of leg I rather long, setiform, solenidion 
<emphasis id="CD97B22FD618FF89FD0287C4F887F941" box="[643,660,1702,1726]" italics="true" pageId="13" pageNumber="291">ω</emphasis>
of leg II shorter, ceratiform, all remaining solenidia (on genu I-III, tibia II and III) thin, almost filiform, about as long as their respective companion seta, and often difficult to observe.
</paragraph>
</subSubSection>
<subSubSection id="B7F93DB6D618FF8AFF448603FBF4FB15" lastPageId="14" lastPageNumber="292" pageId="13" pageNumber="291" type="discussion">
<paragraph id="FF5C6E3DD618FF89FF448603F9D0F97C" blockId="13.[159,762,311,1948]" lastBlockId="13.[826,1429,311,1667]" pageId="13" pageNumber="291">
Remarks — The new species clearly belongs to the nominate subgenus of 
<taxonomicName id="38E315BED618FF89FE6986E7F85FF863" authorityName="C. L. Koch" authorityYear="1835" box="[488,588,1925,1948]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FE6986E7F85FF863" box="[488,588,1925,1948]" italics="true" pageId="13" pageNumber="291">Damaeus</emphasis>
</taxonomicName>
. It shares all important characters of the taxon such as free tibial solenidia, presence of apophysis 
<emphasis id="CD97B22FD618FF89FB6C8039FEEEFE8D" box="[1261,1277,347,370]" italics="true" pageId="13" pageNumber="291">P</emphasis>
, presence of additional ventral setae (
<emphasis id="CD97B22FD618FF89FBE7801FFE9BFE6A" box="[1126,1160,381,405]" italics="true" pageId="13" pageNumber="291">v2’</emphasis>
) on tarsi I and IV, development of prodorsum with set of tubercles (
<emphasis id="CD97B22FD618FF89FAF880C3F9A6FE24" italics="true" pageId="13" pageNumber="291">Ba Bp and Da</emphasis>
) and with rugose surface with various ridges. However, it also bears some very specific characters, which to my knowledge are in present combination unique among 
<emphasis id="CD97B22FD618FF89FB10834FFEEEFDBB" box="[1169,1277,557,580]" italics="true" pageId="13" pageNumber="291">
<taxonomicName id="38E315BED618FF89FB10834FFEEAFDBB" authorityName="C. L. Koch" authorityYear="1835" box="[1169,1273,557,580]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">Damaeus</taxonomicName>
.
</emphasis>
First of all, bothridia are shifted laterad and partly anteriad, so they are positioned quite close to prodorsum margins, above acetabulum I. This is unusual, as this character is very typical for most of the species closer to genus 
<taxonomicName id="38E315BED618FF89FC6E83BEFF17FD0B" authority="von Heyden, 1826" authorityName="von Heyden" authorityYear="1826" box="[1007,1284,732,756]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FC6E83BEFE3AFD0B" box="[1007,1065,732,756]" italics="true" pageId="13" pageNumber="291">Belba</emphasis>
von Heyden, 1826
</taxonomicName>
and usually do not appear within 
<taxonomicName id="38E315BED618FF89FBCB839DFEBDFCE9" authorityName="C. L. Koch" authorityYear="1835" box="[1098,1198,767,790]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="CD97B22FD618FF89FBCB839DFEBDFCE9" box="[1098,1198,767,790]" italics="true" pageId="13" pageNumber="291">Damaeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6A40F54D618FF89FB38839DFF26FCE8" box="[1209,1333,767,791]" pageId="13" pageNumber="291" sensu="lato">sensu lato</taxonomicNameLabel>
. I consider this character being an apomorphy of 
<taxonomicName id="38E315BED618FF89FADA8240FF86FCC5" authorityName="von Heyden" authorityYear="1826" box="[1371,1429,802,826]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FADA8240FF86FCC5" box="[1371,1429,802,826]" italics="true" pageId="13" pageNumber="291">Belba</emphasis>
</taxonomicName>
related group of species, with quite high taxonomic importance at generic or even higher level, and it´s appearance within 
<taxonomicName id="38E315BED618FF89FB0E82EEFEE0FC5C" authorityName="C. L. Koch" authorityYear="1835" box="[1167,1267,908,931]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FB0E82EEFEE0FC5C" box="[1167,1267,908,931]" italics="true" pageId="13" pageNumber="291">Damaeus</emphasis>
</taxonomicName>
could be interpreted as very rare, perhaps parallel developed trait. Other special characters of this species are rather of lower, specific level. Presence of transversal curved ridges at anterior end of notogaster is not surprising, as cuticular thickenings here, even if of different kind, are quite common within 
<taxonomicName id="38E315BED618FF89FAAB853CFF9DFB8A" authorityName="C. L. Koch" authorityYear="1835" box="[1322,1422,1118,1141]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FAAB853CFF9DFB8A" box="[1322,1422,1118,1141]" italics="true" pageId="13" pageNumber="291">Damaeus</emphasis>
</taxonomicName>
. Development of rostral protuberance, even if much more pronounced, have been observed in some of broader relatives, namely in 
<taxonomicName id="38E315BED618FF89FB3585A5FEBFFAFE" authority="Behan-Pelletier &amp; Norton 1985" authorityName="Behan-Pelletier &amp; Norton" authorityYear="1985" class="Arachnida" family="Damaeidae" genus="Epidamaeus" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="CD97B22FD618FF89FB3585A5FF86FB21" box="[1204,1429,1223,1247]" italics="true" pageId="13" pageNumber="291">Epidamaeus nasutus</emphasis>
<bibRefCitation id="9B7213CCD618FF89FCBB8588FEBFFAFE" author="Behan-Pelletier V. M. &amp; Norton R. A." box="[826,1196,1257,1282]" pageId="13" pageNumber="291" pagination="277 - 319" refId="ref6271" refString="Behan-Pelletier V. M., Norton R. A. 1985 - Epidamaeus (Acari: Damaeidae) of subarctic western North America and extreme northeastern USSR. - Canadian Entomologist, 117, 277 - 319. doi: 10.4039 / Ent 117277 - 3" type="journal article" year="1985">Behan-Pelletier &amp; Norton 1985</bibRefCitation>
</taxonomicName>
. Other remarkable characters are epimeral neotrichy, slightly ovoid form of notogaster and presence of load of soil particles on the notogaster providing &quot;camouflage&quot; effect. These are all characters more commonly present in genus 
<taxonomicName id="38E315BED618FF89FB9384FBFE5FFA4E" authorityName="von Heyden" authorityYear="1826" box="[1042,1100,1433,1457]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FB9384FBFE5FFA4E" box="[1042,1100,1433,1457]" italics="true" pageId="13" pageNumber="291">Belba</emphasis>
</taxonomicName>
, similarly as shifted bothridia. If all these characters indicate more derived state of 
<taxonomicName id="38E315BED618FF89FC258482FE1BFA08" authorityName="C. L. Koch" authorityYear="1835" box="[932,1032,1504,1527]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FC258482FE1BFA08" box="[932,1032,1504,1527]" italics="true" pageId="13" pageNumber="291">Damaeus</emphasis>
</taxonomicName>
&quot;towards&quot; the line of 
<taxonomicName id="38E315BED618FF89FAAA84BDFF76FA08" authorityName="von Heyden" authorityYear="1826" box="[1323,1381,1503,1527]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FAAA84BDFF76FA08" box="[1323,1381,1503,1527]" italics="true" pageId="13" pageNumber="291">Belba</emphasis>
</taxonomicName>
, or rather combination of parallel developed derived characters and more ancient ones which could have been shared by some common ancestors remains to be clarified.
</paragraph>
<paragraph id="FF5C6E3DD618FF8AFCE187D0FB65FE47" blockId="13.[826,1429,1714,1949]" lastBlockId="14.[159,762,311,1258]" lastPageId="14" lastPageNumber="292" pageId="13" pageNumber="291">
When considering affinities within 
<taxonomicName id="38E315BED618FF89FB7F87D1FF71F935" authorityName="C. L. Koch" authorityYear="1835" box="[1278,1378,1715,1738]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD618FF89FB7F87D1FF71F935" box="[1278,1378,1715,1738]" italics="true" pageId="13" pageNumber="291">Damaeus</emphasis>
</taxonomicName>
, the new species does not seem to belong to any clear species-group. Curved and heterotrichous notogastral setae, as well as mostly very short setae on legs indicate some relations to 
<taxonomicName id="38E315BED618FF89FBE9865DFEF9F8A9" baseAuthorityName="Kulczynski" baseAuthorityYear="1902" box="[1128,1258,1855,1878]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="291" phylum="Arthropoda" rank="species" species="crispatus">
<emphasis id="CD97B22FD618FF89FBE9865DFEF9F8A9" box="[1128,1258,1855,1878]" italics="true" pageId="13" pageNumber="291">D. crispatus</emphasis>
</taxonomicName>
group, but the mentioned specific characters separate this species quite significantly also from species of this group. Identification of the new species is therefore easy, as the combination of characters is unique and species cannot be misidentified with any of known 
<taxonomicName id="38E315BED61BFF8AFD14801CF8EAFE6A" authorityName="C. L. Koch" authorityYear="1835" box="[661,761,382,405]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="292" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="CD97B22FD61BFF8AFD14801CF8EAFE6A" box="[661,761,382,405]" italics="true" pageId="14" pageNumber="292">Damaeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6A40F54D61BFF8AFF1E80C2FB07FE47" box="[159,276,416,440]" pageId="14" pageNumber="292" sensu="lato">sensu lato</taxonomicNameLabel>
species.
</paragraph>
<paragraph id="FF5C6E3DD61BFF8AFF4480ABF886FCC0" blockId="14.[159,762,311,1258]" pageId="14" pageNumber="292">
Unfortunately, only single larva was available in our material, and judgements about ontogeny are therefore very difficult. Nevertheless, the larva is easily recognized by very long notogastral setae, some of them being longer than the whole body. Porose, double-walled sejugal apodeme and saclike porose vestibules from apodemes I and III in immatures, typical for 
<taxonomicName id="38E315BED61BFF8AFE3883DCFBE0FD29" authorityName="von Heyden" authorityYear="1826" box="[441,499,702,726]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD61BFF8AFE3883DCFBE0FD29" box="[441,499,702,726]" italics="true" pageId="14" pageNumber="292">Belba</emphasis>
</taxonomicName>
species were not observed, which supports the placing of the species within 
<taxonomicName id="38E315BED61BFF8AFF728267FB44FCE3" authorityName="C. L. Koch" authorityYear="1835" box="[243,343,773,796]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="292" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="CD97B22FD61BFF8AFF728267FB44FCE3" box="[243,343,773,796]" italics="true" pageId="14" pageNumber="292">Damaeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6A40F54D61BFF8AFEDE8267FBC4FCE3" box="[351,471,772,796]" pageId="14" pageNumber="292" sensu="lato">sensu lato</taxonomicNameLabel>
, despite of some characters being similar to those of 
<taxonomicName id="38E315BED61BFF8AFE738245F83FFCC0" authorityName="von Heyden" authorityYear="1826" box="[498,556,807,831]" class="Arachnida" family="Damaeidae" genus="Belba" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="CD97B22FD61BFF8AFE738245F83FFCC0" box="[498,556,807,831]" italics="true" pageId="14" pageNumber="292">Belba</emphasis>
</taxonomicName>
in adult.
</paragraph>
<paragraph id="FF5C6E3DD61BFF8AFF448232FBF4FB15" blockId="14.[159,762,311,1258]" pageId="14" pageNumber="292">
Derivatio nominis — The species name &quot; 
<taxonomicName id="38E315BED61BFF8AFD328232F8E3FC97" authorityName="Miko" authorityYear="2016" box="[691,752,848,872]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="292" phylum="Arthropoda" rank="species" species="lupus">
<emphasis id="CD97B22FD61BFF8AFD328232F8E3FC97" box="[691,752,848,872]" italics="true" pageId="14" pageNumber="292">lupus</emphasis>
</taxonomicName>
&quot; refers to the Latin name of wolf, an iconic species of remaining European wilderness territories. At the same time, the Slovak word for wolf is &quot;vlk&quot;, which is also the name of well-known Slovak nongovernmental organisation, which is active in protection of natural habitats and large carnivores in Slovakian Carpathians. The new species is dedicated to all friends and colleagues from this organisation as expression of gratitude for their so important work, and also namely to the founder of this initiative, Juraj &quot;Vlk&quot; LukAEˇc.
</paragraph>
</subSubSection>
</treatment>
</document>