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<document id="D7E9CABA6157E7BB657E24E16DCCF2DD" ID-DOI="10.11646/zootaxa.5403.4.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10562280" ID-ZooBank="CDF2D3A6-334D-4208-B0F7-ACFC268316AA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1706105492649" checkinUser="plazi" docAuthor="Peng, Yun-Fei, Bian, Dong-Ju &amp; Wang, Ju-Ping" docDate="2024" docId="038287B7FD20FFE5FF54FD2871BBFDC7" docLanguage="en" docName="zootaxa.5403.4.7.pdf" docOrigin="Zootaxa 5403 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5403.4.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zaitzevia triangularis Peng &amp; Bian &amp; Wang 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="493" masterDocId="FFBBFFCFFD23FFE0FFC3FFF67143FFDE" masterDocTitle="Heterlimnius luyashanensis sp. n. and Zaitzevia triangularis sp. n. from Shanxi Province, China (Coleoptera: Elmidae)" masterLastPageNumber="494" masterPageNumber="488" pageNumber="491" updateTime="1706106029902" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="29D18B002B84E6DD78A1761B454E67A0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="94AABF2E1FFBA1D960FF54CF8F2295F5">
<mods:title id="7B7A5CB0578C292FBA80C84A55133D6C">Heterlimnius luyashanensis sp. n. and Zaitzevia triangularis sp. n. from Shanxi Province, China (Coleoptera: Elmidae)</mods:title>
</mods:titleInfo>
<mods:name id="E162362570BC546CD7FC20144E98E6B8" type="personal">
<mods:role id="0BFBADDD93C607FDE0C05A8D1A6D3243">
<mods:roleTerm id="3E7DF3CC3D48F533B6004E0E5D57B738">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1CF380D0CC2EF164929A3B03AACC25AD">Peng, Yun-Fei</mods:namePart>
<mods:nameIdentifier id="1F1A147F01C3E66C73E5F4DD329757BC" type="ORCID">0000-0002-8091-2282</mods:nameIdentifier>
<mods:affiliation id="1ABB35DDDD0BAC09BE77FEDCCCA6DEB0">Shanxi Insect Herbarium, College of Plant Protection, Shanxi Agricultural University, Taiyuan, 030031, China.</mods:affiliation>
</mods:name>
<mods:name id="558C1AA47CC6CEC83E18D0F52282628C" type="personal">
<mods:role id="BEF1E415FDBE667C9453F86E6450DBA4">
<mods:roleTerm id="EA2753689D43A37D6556EEEA2D324704">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="21BB735EC1F8A72C171A51F8F972F7D5">Bian, Dong-Ju</mods:namePart>
<mods:affiliation id="7548377EEF4D9166C2F39C6ED6DD110A">Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Shenyang, 110016, Chinese Academy of Sciences.</mods:affiliation>
</mods:name>
<mods:name id="F2DFBD0D125369F90E15A219C62003FE" type="personal">
<mods:role id="55AAACC908C0B239F9A3D5869A2129CD">
<mods:roleTerm id="538E37E9BD4FF84E5F54CEFD0099AE93">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5BF54DFEB6D4BAD3BBCAA2EAC5730EDB">Wang, Ju-Ping</mods:namePart>
<mods:nameIdentifier id="09FCF4A881B348DB1C1E554050AA58E5" type="ORCID">0000-0002-8091-2282</mods:nameIdentifier>
<mods:affiliation id="C49E1D3CED8D0F90804A1EA7D4B33C90">Shanxi Insect Herbarium, College of Plant Protection, Shanxi Agricultural University, Taiyuan, 030031, China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="07BDEFDDB352F3D6C863477CA1B0B614">text</mods:typeOfResource>
<mods:relatedItem id="98FE4691BA12EBC44EDB13E797E6C8D1" type="host">
<mods:titleInfo id="7BE4FE5EDB3408DAA6359B99C24850DC">
<mods:title id="D0B2D77BB8EC893C8CE36EF8B80E6B20">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E9E0D918F423535B0E419AC38299552B">
<mods:date id="CC449055D12074DAF3D6F27D6B2FEF03">2024</mods:date>
<mods:detail id="FE2B0D5D249EDAA01E55891E42473A9A" type="pubDate">
<mods:number id="021134C335347D431A14E6DF686F4DBF">2024-01-23</mods:number>
</mods:detail>
<mods:detail id="19ACD6A11729AA6446DCDA281F092CA9" type="volume">
<mods:number id="F6750004EB18E89D3752B8E66D61EC66">5403</mods:number>
</mods:detail>
<mods:detail id="CCEC26B0520D30FF806986D534E44DE0" type="issue">
<mods:number id="D83D22455BEFDE6A0D82BC8DE570BB3A">4</mods:number>
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<mods:start id="5A596BDEACBE326F1F408909846E3366">488</mods:start>
<mods:end id="A458906A56D3FD3112F54A9C38543157">494</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="A3DA8B8AE0FFFDB39CC4A304E5487005">
<mods:url id="7D0E1583D9054E268BDF319122766332">http://dx.doi.org/10.11646/zootaxa.5403.4.7</mods:url>
</mods:location>
<mods:classification id="6F2784910819B28FB0596A067DE0DF83">journal article</mods:classification>
<mods:identifier id="EB4D1DD24F5B901068CF7BF5CABE7569" type="DOI">10.11646/zootaxa.5403.4.7</mods:identifier>
<mods:identifier id="B18541E2F6BC98D613547E21790A4A50" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="83425DB29F68A41FDC9DA306D785A509" type="Zenodo-Dep">10562280</mods:identifier>
<mods:identifier id="63D56B284D1DDF0D149312BC47A72BE7" type="ZooBank">CDF2D3A6-334D-4208-B0F7-ACFC268316AA</mods:identifier>
</mods:mods>
<treatment id="038287B7FD20FFE5FF54FD2871BBFDC7" ID-DOI="http://doi.org/10.5281/zenodo.10562266" ID-Zenodo-Dep="10562266" LSID="urn:lsid:plazi:treatment:038287B7FD20FFE5FF54FD2871BBFDC7" httpUri="http://treatment.plazi.org/id/038287B7FD20FFE5FF54FD2871BBFDC7" lastPageId="5" lastPageNumber="493" pageId="3" pageNumber="491">
<subSubSection id="C331652AFD20FFE3FF54FD2870A6FD26" box="[151,485,734,761]" pageId="3" pageNumber="491" type="nomenclature">
<paragraph id="8B9436A1FD20FFE3FF54FD2870A6FD26" blockId="3.[151,485,734,796]" box="[151,485,734,761]" pageId="3" pageNumber="491">
<heading id="D0DC81CDFD20FFE3FF54FD2870A6FD26" bold="true" box="[151,485,734,761]" fontSize="11" level="1" pageId="3" pageNumber="491" reason="1">
<emphasis id="B95FEAB3FD20FFE3FF54FD2870A6FD26" bold="true" box="[151,485,734,761]" pageId="3" pageNumber="491">
<taxonomicName id="4C2B4D22FD20FFE3FF54FD2870D8FD27" authority="Peng &amp; Bian &amp; Wang, 2024" authorityName="Peng &amp; Bian &amp; Wang" authorityYear="2024" box="[151,411,734,761]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
<emphasis id="B95FEAB3FD20FFE3FF54FD2870D8FD27" bold="true" box="[151,411,734,761]" italics="true" pageId="3" pageNumber="491">Zaitzevia triangularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57C8FD20FFE3FE61FD2870A6FD26" box="[418,485,734,760]" pageId="3" pageNumber="491" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331652AFD20FFE3FF54FCF470CAFCC2" box="[151,393,770,796]" pageId="3" pageNumber="491" type="description">
<paragraph id="8B9436A1FD20FFE3FF54FCF470CAFCC2" blockId="3.[151,485,734,796]" box="[151,393,770,796]" pageId="3" pageNumber="491">
(
<figureCitation id="13102A24FD20FFE3FF5CFCF4706FFCC2" box="[159,300,770,796]" captionStart="FIGURE 3" captionStartId="4.[151,250,1781,1805]" captionTargetBox="[151,1435,277,1758]" captionTargetId="figure-55@4.[151,1436,273,1758]" captionTargetPageId="4" captionText="FIGURE 3. Zaitzevia triangularis sp. n. A. Habitus, holotype, male, in dorsal view. B. Habitus, in ventral view C. Pronotum D. Prosternal process and metaventrite." figureDoi="http://doi.org/10.5281/zenodo.10562288" httpUri="https://zenodo.org/record/10562288/files/figure.png" pageId="3" pageNumber="491">Figs. 3 A–D</figureCitation>
;
<figureCitation id="13102A24FD20FFE3FEF4FCF470C2FCC2" box="[311,385,770,796]" captionStart="FIGURE 4" captionStartId="5.[151,250,1528,1552]" captionTargetBox="[313,1274,561,1505]" captionTargetId="figure-170@5.[313,1274,561,1505]" captionTargetPageId="5" captionText="FIGURE 4. Aedeagus, holotype, male, Zaitzevia triangularis sp. n. A. in ventral view B. in dorsal view C in lateral view." figureDoi="http://doi.org/10.5281/zenodo.10562290" httpUri="https://zenodo.org/record/10562290/files/figure.png" pageId="3" pageNumber="491">4 A–C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C331652AFD20FFE3FF54FCBC70CEFBC6" pageId="3" pageNumber="491" type="materials_examined">
<paragraph id="8B9436A1FD20FFE3FF54FCBC70CEFBC6" blockId="3.[151,1437,842,1949]" pageId="3" pageNumber="491">
<emphasis id="B95FEAB3FD20FFE3FF54FCBC7009FCBA" bold="true" box="[151,330,842,868]" pageId="3" pageNumber="491">Type material:</emphasis>
<materialsCitation id="3B433CFCFD20FFE3FE97FCBC73BAFC56" collectingDate="2021-08-14" collectionCode="IAECAS" country="China" elevation="1550" latitude="38.735558" location="Luya Mountains" longLatPrecision="1" longitude="112.17861" municipality="Xinzhou City" pageId="3" pageNumber="491" specimenCount="1" stateProvince="Shanxi" typeStatus="holotype">
<typeStatus id="54908803FD20FFE3FE97FCBC70FAFCBA" box="[340,441,842,868]" pageId="3" pageNumber="491" type="holotype">holotype</typeStatus>
, male (
<collectionCode id="ED3AAE64FD20FFE3FDCCFCBC7338FCBA" box="[527,635,842,868]" pageId="3" pageNumber="491">IAECAS</collectionCode>
): “
<collectingCountry id="F33C7631FD20FFE3FD58FCBC73B9FCBA" box="[667,762,842,868]" name="China" pageId="3" pageNumber="491">CHINA</collectingCountry>
:
<collectingRegion id="49EFF843FD20FFE3FCCBFCBC7215FCBA" box="[776,854,842,868]" country="China" name="Shanxi" pageId="3" pageNumber="491">Shanxi</collectingRegion>
/
<collectingMunicipality id="6BF0ACDBFD20FFE3FCB1FCBC754FFCBA" box="[882,1036,842,868]" pageId="3" pageNumber="491">Xinzhou City</collectingMunicipality>
/
<location id="8EF4607AFD20FFE3FBEBFCBD75A7FCBA" LSID="urn:lsid:plazi:treatment:038287B7FD20FFE5FF54FD2871BBFDC7:8EF4607AFD20FFE3FBEBFCBD75A7FCBA" box="[1064,1252,842,869]" country="China" latitude="38.735558" longLatPrecision="1" longitude="112.17861" municipality="Xinzhou City" name="Luya Mountains" pageId="3" pageNumber="491" stateProvince="Shanxi">Luya Mountains</location>
|
<geoCoordinate id="EE1F5066FD20FFE3FB3EFCBC74DFFCBA" box="[1277,1436,842,868]" degrees="112.178611" direction="east" orientation="longitude" pageId="3" pageNumber="491" precision="1" value="112.17861">112.178611°E</geoCoordinate>
<geoCoordinate id="EE1F5066FD20FFE3FF54FC98706EFC57" box="[151,301,878,905]" degrees="38.735556" direction="north" orientation="latitude" pageId="3" pageNumber="491" precision="1" value="38.735558">38.735556°N</geoCoordinate>
/ ca.
<quantity id="4CD39B44FD20FFE3FE9CFC9870FBFC57" box="[351,440,878,905]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="491" unit="m" value="1550.0">
<elevation id="0006D192FD20FFE3FE9CFC9870FBFC57" box="[351,440,878,905]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="491" unit="m" value="1550.0">1550 m</elevation>
</quantity>
,
<date id="FF951061FD20FFE3FE01FC997322FC56" box="[450,609,878,905]" pageId="3" pageNumber="491" value="2021-08-14">
<collectingDate id="EFD1E989FD20FFE3FE01FC997322FC56" box="[450,609,878,905]" pageId="3" pageNumber="491" value="2021-08-14">VIII. 14. 2021</collectingDate>
</date>
/ Leg. Peng.”
</materialsCitation>
<materialsCitation id="3B433CFCFD20FFE3FD3CFC9970C9FBC6" collectingDate="2020-06-15" collectingDateMax="2021-08-13" collectingDateMin="2020-06-15" collectionCode="IAECAS, SIH" country="China" county="Xinzhou City" elevation="1920" latitude="38.69361" location="Xinzhou City" longLatPrecision="1" longitude="111.875275" municipality="Luya Mountains" pageId="3" pageNumber="491" specimenCount="16" specimenCount-female="7" specimenCount-male="9" stateProvince="Shanxi" typeStatus="paratype">
<typeStatus id="54908803FD20FFE3FD3CFC997233FC57" box="[767,880,879,905]" pageId="3" pageNumber="491" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D2DFD28FD20FFE3FCB9FC987299FC56" box="[890,986,878,904]" count="1" pageId="3" pageNumber="491" type="female">1 female</specimenCount>
(
<collectionCode id="ED3AAE64FD20FFE3FC2BFC987510FC56" box="[1000,1107,878,904]" pageId="3" pageNumber="491">IAECAS</collectionCode>
), the same data as holotype.
<specimenCount id="9D2DFD28FD20FFE3FA4DFC987193FC72" count="1" pageId="3" pageNumber="491" type="male">1 male</specimenCount>
,
<specimenCount id="9D2DFD28FD20FFE3FF1FFC64707EFC72" box="[220,317,914,940]" count="1" pageId="3" pageNumber="491" type="female">1 female</specimenCount>
(
<collectionCode id="ED3AAE64FD20FFE3FE8FFC6470FBFC72" box="[332,440,914,940]" pageId="3" pageNumber="491">IAECAS</collectionCode>
): “
<collectingCountry id="F33C7631FD20FFE3FE17FC647370FC72" box="[468,563,914,940]" name="China" pageId="3" pageNumber="491">CHINA</collectingCountry>
:
<collectingRegion id="49EFF843FD20FFE3FDFDFC6473CFFC72" box="[574,652,914,940]" country="China" name="Shanxi" pageId="3" pageNumber="491">Shanxi</collectingRegion>
/
<collectingCounty id="62F54E2DFD20FFE3FD61FC64727AFC72" box="[674,825,914,940]" pageId="3" pageNumber="491">Xinzhou City</collectingCounty>
/
<collectingMunicipality id="6BF0ACDBFD20FFE3FC8CFC657544FC72" box="[847,1031,914,941]" pageId="3" pageNumber="491">Luya Mountains</collectingMunicipality>
|
<geoCoordinate id="EE1F5066FD20FFE3FBD8FC6475F9FC73" box="[1051,1210,914,941]" degrees="111.875278" direction="east" orientation="longitude" pageId="3" pageNumber="491" precision="1" value="111.875275">111.875278°E</geoCoordinate>
<geoCoordinate id="EE1F5066FD20FFE3FB02FC64741AFC72" box="[1217,1369,914,940]" degrees="38.693611" direction="north" orientation="latitude" pageId="3" pageNumber="491" precision="1" value="38.69361">38.693611°N</geoCoordinate>
,
<quantity id="4CD39B44FD20FFE3FAA7FC6471EEFC0F" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="3" pageNumber="491" unit="m" value="1920.0">
<elevation id="0006D192FD20FFE3FAA7FC6471EEFC0F" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="3" pageNumber="491" unit="m" value="1920.0">1920 m</elevation>
</quantity>
/
<date id="FF951061FD20FFE3FF07FC417025FC0E" box="[196,358,950,977]" pageId="3" pageNumber="491" value="2021-08-13">
<collectingDate id="EFD1E989FD20FFE3FF07FC417025FC0E" box="[196,358,950,977]" pageId="3" pageNumber="491" value="2021-08-13">VIII. 13. 2021</collectingDate>
</date>
/ Leg. Peng”;
<specimenCount id="9D2DFD28FD20FFE3FDCEFC407324FC0E" box="[525,615,950,977]" count="5" pageId="3" pageNumber="491" type="male">5 males</specimenCount>
,
<specimenCount id="9D2DFD28FD20FFE3FDB0FC4073A3FC0E" box="[627,736,950,976]" count="3" pageId="3" pageNumber="491" type="female">3 females</specimenCount>
(
<collectionCode id="ED3AAE64FD20FFE3FD33FC407260FC0E" box="[752,803,950,976]" pageId="3" pageNumber="491">SIH</collectionCode>
): same label data as last one, but
<date id="FF951061FD20FFE3FB5CFC417471FC0E" box="[1183,1330,950,977]" pageId="3" pageNumber="491" value="2020-06-15">
<collectingDate id="EFD1E989FD20FFE3FB5CFC417471FC0E" box="[1183,1330,950,977]" pageId="3" pageNumber="491" value="2020-06-15">VI. 15. 2020</collectingDate>
</date>
;
<specimenCount id="9D2DFD28FD20FFE3FAFDFC4074DBFC0E" box="[1342,1432,950,976]" count="3" pageId="3" pageNumber="491" type="male">3 males</specimenCount>
,
<specimenCount id="9D2DFD28FD20FFE3FF54FC2C7045FC2A" box="[151,262,986,1012]" count="2" pageId="3" pageNumber="491" type="female">2 females</specimenCount>
(
<collectionCode id="ED3AAE64FD20FFE3FEDAFC2C70C6FC2A" box="[281,389,986,1012]" pageId="3" pageNumber="491">IAECAS</collectionCode>
): “
<collectingCountry id="F33C7631FD20FFE3FE66FC2C7347FC2A" box="[421,516,986,1012]" name="China" pageId="3" pageNumber="491">CHINA</collectingCountry>
:
<collectingRegion id="49EFF843FD20FFE3FDD1FC2C7323FC2A" box="[530,608,986,1012]" country="China" name="Shanxi" pageId="3" pageNumber="491">Shanxi</collectingRegion>
/
<location id="8EF4607AFD20FFE3FDBFFC2C7254FC2A" LSID="urn:lsid:plazi:treatment:038287B7FD20FFE5FF54FD2871BBFDC7:8EF4607AFD20FFE3FDBFFC2C7254FC2A" box="[636,791,986,1012]" country="China" county="Xinzhou City" latitude="38.69361" longLatPrecision="1" longitude="111.875275" municipality="Luya Mountains" name="Xinzhou City" pageId="3" pageNumber="491" stateProvince="Shanxi">Xinzhou City</location>
/
<location id="8EF4607AFD20FFE3FCF0FC2D72ACFC2A" LSID="urn:lsid:plazi:treatment:038287B7FD20FFE5FF54FD2871BBFDC7:8EF4607AFD20FFE3FCF0FC2D72ACFC2A" box="[819,1007,986,1013]" country="China" county="Xinzhou City" latitude="38.69361" longLatPrecision="1" longitude="111.875275" municipality="Luya Mountains" name="Luya Mountains" pageId="3" pageNumber="491" stateProvince="Shanxi">Luya Mountains</location>
|
<geoCoordinate id="EE1F5066FD20FFE3FBCAFC2C75EAFC2B" box="[1033,1193,986,1013]" degrees="111.885272" direction="east" orientation="longitude" pageId="3" pageNumber="491" precision="1" value="111.88527">111.885272°E</geoCoordinate>
<geoCoordinate id="EE1F5066FD20FFE3FB70FC2C7409FC2A" box="[1203,1354,986,1012]" degrees="38.729431" direction="north" orientation="latitude" pageId="3" pageNumber="491" precision="1" value="38.72943">38.729431°N</geoCoordinate>
/ VIII. 13.2021 / Leg. Peng”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331652AFD20FFE3FF04FBD47230F9EA" pageId="3" pageNumber="491" type="diagnosis">
<paragraph id="8B9436A1FD20FFE3FF04FBD47230F9EA" blockId="3.[151,1437,842,1949]" pageId="3" pageNumber="491">
<emphasis id="B95FEAB3FD20FFE3FF04FBD47079FBE2" bold="true" box="[199,314,1058,1084]" pageId="3" pageNumber="491">Diagnosis</emphasis>
. In the genus
<taxonomicName id="4C2B4D22FD20FFE3FE18FBD57301FBE2" authorityName="Champion" authorityYear="1923" box="[475,578,1059,1084]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAB3FD20FFE3FE18FBD57301FBE2" box="[475,578,1059,1084]" italics="true" pageId="3" pageNumber="491">Zaitzevia</emphasis>
</taxonomicName>
, the new species has the similar size with the five known species:
<taxonomicName id="4C2B4D22FD20FFE3FAF5FBD5734DFBBE" authority="Jiang &amp; Wang, 2020" authorityName="Jiang &amp; Wang" authorityYear="2020" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="chenzhitengi">
<emphasis id="B95FEAB3FD20FFE3FAF5FBD57066FBBE" italics="true" pageId="3" pageNumber="491">Zaitzevia chenzhitengi</emphasis>
<bibRefCitation id="EFBA4B50FD20FFE3FEE9FBB0734DFBBE" author="Jiang, R. - X. &amp; Wang, S." box="[298,526,1094,1121]" pageId="3" pageNumber="491" pagination="231 - 238" refId="ref3028" refString="Jiang, R. - X. &amp; Wang, S. (2020) Two new species of the genus Zaitzevia Champion, 1923 from China (Coleoptera: Elmidae: Macronychini). Zootaxa, 4852 (2), 231 - 238. https: // doi. org / 10.11646 / zootaxa. 4852.2.8" type="journal article" year="2020">Jiang &amp; Wang, 2020</bibRefCitation>
</taxonomicName>
, Z.
<taxonomicName id="4C2B4D22FD20FFE3FDF6FBB072D3FBBE" authority="Bian &amp; Zhang, 2022" authorityName="Bian &amp; Zhang" authorityYear="2022" box="[565,912,1094,1120]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="muchenae">
<emphasis id="B95FEAB3FD20FFE3FDF6FBB073E4FBBE" box="[565,679,1094,1120]" italics="true" pageId="3" pageNumber="491">muchenae</emphasis>
<bibRefCitation id="EFBA4B50FD20FFE3FD6FFBB072D3FBBE" author="Bian, D. - J. &amp; Zhang, Y." box="[684,912,1094,1120]" pageId="3" pageNumber="491" pagination="257 - 266" refId="ref2679" refString="Bian, D. - J. &amp; Zhang, Y. (2022) Three new species of the genus Zaitzevia Champion, 1923 from China (Coleoptera: Elmidae: Macronychini). Zootaxa, 5190 (2), 257 - 266. https: // doi. org / 10.11646 / zootaxa. 5190.2.5" type="journal article" year="2022">Bian &amp; Zhang, 2022</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2B4D22FD20FFE3FC59FBB175A4FBBE" authority="Nomura &amp; Baba, 1961" authorityName="Nomura &amp; Baba" authorityYear="1961" box="[922,1255,1094,1121]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B95FEAB3FD20FFE3FC59FBB172A6FBBE" box="[922,997,1094,1120]" italics="true" pageId="3" pageNumber="491">Z. rufa</emphasis>
Nomura &amp; Baba, 1961
</taxonomicName>
,
<taxonomicName id="4C2B4D22FD20FFE3FB32FBB17036FB5A" authority="Jiang &amp; Wang, 2020" authorityName="Jiang &amp; Wang" authorityYear="2020" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="xiongzichuni">
<emphasis id="B95FEAB3FD20FFE3FB32FBB174DFFBBE" box="[1265,1436,1094,1120]" italics="true" pageId="3" pageNumber="491">Z. xiongzichuni</emphasis>
<bibRefCitation id="EFBA4B50FD20FFE3FF54FB9C7036FB5A" author="Jiang, R. - X. &amp; Wang, S." box="[151,373,1130,1157]" pageId="3" pageNumber="491" pagination="231 - 238" refId="ref3028" refString="Jiang, R. - X. &amp; Wang, S. (2020) Two new species of the genus Zaitzevia Champion, 1923 from China (Coleoptera: Elmidae: Macronychini). Zootaxa, 4852 (2), 231 - 238. https: // doi. org / 10.11646 / zootaxa. 4852.2.8" type="journal article" year="2020">Jiang &amp; Wang, 2020</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2B4D22FD20FFE3FE65FB9D7272FB5A" authority="Jiang &amp; Chen, 2023" authorityName="Jiang &amp; Chen" authorityYear="2023" box="[422,817,1130,1156]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="B95FEAB3FD20FFE3FE65FB9D7312FB5A" box="[422,593,1130,1156]" italics="true" pageId="3" pageNumber="491">Z. sichuanensis</emphasis>
<bibRefCitation id="EFBA4B50FD20FFE3FD96FB9C7272FB5A" author="Jiang, R. - X. &amp; Chen, X. - S." box="[597,817,1130,1156]" pageId="3" pageNumber="491" pagination="191 - 206" refId="ref2972" refString="Jiang, R. - X. &amp; Chen, X. - S. (2023) Three new species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from China. ZooKeys, 1174, 191 - 206. https: // doi. org / 10.3897 / zookeys. 1174.101046" type="journal article" year="2023">Jiang &amp; Chen, 2023</bibRefCitation>
</taxonomicName>
; it is distinctly smaller than
<taxonomicName id="4C2B4D22FD20FFE3FB9DFB9D7054FB76" authority="Jiang &amp; Chen, 2023" authorityName="Jiang &amp; Chen" authorityYear="2023" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="fengtongzhaiensis">
<emphasis id="B95FEAB3FD20FFE3FB9DFB9D7400FB5A" box="[1118,1347,1130,1156]" italics="true" pageId="3" pageNumber="491">Z. fengtongzhaiensis</emphasis>
<bibRefCitation id="EFBA4B50FD20FFE3FA84FB9C7054FB76" author="Jiang, R. - X. &amp; Chen, X. - S." pageId="3" pageNumber="491" pagination="191 - 206" refId="ref2972" refString="Jiang, R. - X. &amp; Chen, X. - S. (2023) Three new species of the genus Zaitzevia Champion, 1923 (Coleoptera, Elmidae) from China. ZooKeys, 1174, 191 - 206. https: // doi. org / 10.3897 / zookeys. 1174.101046" type="journal article" year="2023">Jiang &amp; Chen, 2023</bibRefCitation>
</taxonomicName>
and larger than other species in
<emphasis id="B95FEAB3FD20FFE3FDBBFB7973A4FB76" box="[632,743,1167,1192]" italics="true" pageId="3" pageNumber="491">
<taxonomicName id="4C2B4D22FD20FFE3FDBBFB7973A1FB76" authorityName="Champion" authorityYear="1923" box="[632,738,1167,1192]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="genus">Zaitzevia</taxonomicName>
.
</emphasis>
This species can be easily distinguished from
<taxonomicName id="4C2B4D22FD20FFE3FB27FB7974DEFB76" authorityName="Jiang &amp; Wang" authorityYear="2020" box="[1252,1437,1166,1192]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="chengzhitengi">
<emphasis id="B95FEAB3FD20FFE3FB27FB7974DEFB76" box="[1252,1437,1166,1192]" italics="true" pageId="3" pageNumber="491">Z. chengzhitengi</emphasis>
</taxonomicName>
by its median longitudinal sulcus wide, apex of penis sub-triangular, not like an arrow as in
<taxonomicName id="4C2B4D22FD20FFE3FB69FB457414FB12" authorityName="Jiang &amp; Wang" authorityYear="2020" box="[1194,1367,1202,1228]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="chenzhitengi">
<emphasis id="B95FEAB3FD20FFE3FB69FB457414FB12" box="[1194,1367,1202,1228]" italics="true" pageId="3" pageNumber="491">Z. chenzhitengi</emphasis>
</taxonomicName>
; from
<taxonomicName id="4C2B4D22FD20FFE3FF54FB217068FB2E" authorityName="Bian &amp; Zhang" authorityYear="2022" box="[151,299,1238,1265]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="muchenae">
Z.
<emphasis id="B95FEAB3FD20FFE3FF7AFB207068FB2E" box="[185,299,1238,1264]" italics="true" pageId="3" pageNumber="491">muchenae</emphasis>
</taxonomicName>
by distal third of elytra distinctly attenuated, not gradually attenuated as in
<taxonomicName id="4C2B4D22FD20FFE3FB62FB217470FB2E" authorityName="Bian &amp; Zhang" authorityYear="2022" box="[1185,1331,1238,1264]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="muchenae">
<emphasis id="B95FEAB3FD20FFE3FB62FB217470FB2E" box="[1185,1331,1238,1264]" italics="true" pageId="3" pageNumber="491">Z. muchenae</emphasis>
</taxonomicName>
and also by distal 0.4 of penis distinctly narrowed, not distal 0.1 of penis distinctly narrowed as in
<taxonomicName id="4C2B4D22FD20FFE3FB67FB0D7475FACA" authorityName="Bian &amp; Zhang" authorityYear="2022" box="[1188,1334,1274,1300]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="muchenae">
<emphasis id="B95FEAB3FD20FFE3FB67FB0D7475FACA" box="[1188,1334,1274,1300]" italics="true" pageId="3" pageNumber="491">Z. muchenae</emphasis>
</taxonomicName>
; from
<taxonomicName id="4C2B4D22FD20FFE3FA45FB0D7185FAE6" authorityName="Nomura &amp; Baba" authorityYear="1961" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B95FEAB3FD20FFE3FA45FB0D7185FAE6" italics="true" pageId="3" pageNumber="491">
<emphasis id="B95FEAB3FD20FFE3FA45FB0D74DFFACB" bold="true" box="[1414,1436,1275,1301]" italics="true" pageId="3" pageNumber="491">Z.</emphasis>
rufa
</emphasis>
</taxonomicName>
by its body colour dark brown, median longitudinal sulcus of pronotum wide and distinctly impressed, distal 0.4 of penis distinctly narrowed, but in
<taxonomicName id="4C2B4D22FD20FFE3FDA2FAB573F3FA82" authorityName="Nomura &amp; Baba" authorityYear="1961" box="[609,688,1346,1372]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B95FEAB3FD20FFE3FDA2FAB573F3FA82" box="[609,688,1346,1372]" italics="true" pageId="3" pageNumber="491">Z. rufa</emphasis>
</taxonomicName>
, body colour reddish brown, middle sulcus shallowly impressed, and distal 0.4 of penis gradually narrowed; from
<taxonomicName id="4C2B4D22FD20FFE3FD13FA91723CFA5E" authorityName="Jiang &amp; Wang" authorityYear="2020" box="[720,895,1382,1408]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="xiongzichuni">
<emphasis id="B95FEAB3FD20FFE3FD13FA91723CFA5E" box="[720,895,1382,1408]" italics="true" pageId="3" pageNumber="491">Z. xiongzichuni</emphasis>
</taxonomicName>
by its median longitudinal sulcus of pronotum broad in middle, apices of elytra distinctly separated rounded, apex of penis sub-triangular, but in
<taxonomicName id="4C2B4D22FD20FFE3FB18FA7D74D6FA7A" authorityName="Jiang &amp; Wang" authorityYear="2020" box="[1243,1429,1418,1444]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="xiongzhichuni">
<emphasis id="B95FEAB3FD20FFE3FB18FA7D74D6FA7A" box="[1243,1429,1418,1444]" italics="true" pageId="3" pageNumber="491">Z. xiongzhichuni</emphasis>
</taxonomicName>
, median longitudinal sulcus narrow, not broad in middle, apices of elytra almost connected, and apex of penis is cuspidal; from
<taxonomicName id="4C2B4D22FD20FFE3FEA7FA257350FA32" authorityName="Jiang &amp; Chen" authorityYear="2023" box="[356,531,1490,1516]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="B95FEAB3FD20FFE3FEA7FA257350FA32" box="[356,531,1490,1516]" italics="true" pageId="3" pageNumber="491">Z. sichuanensis</emphasis>
</taxonomicName>
by basal 1/3 disc of pronotum and elytral intervals coarse, distal 0.4 of penis distinctly narrowed, while in
<taxonomicName id="4C2B4D22FD20FFE3FE1DFA0173C9F9CE" authorityName="Jiang &amp; Chen" authorityYear="2023" box="[478,650,1526,1552]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="491" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="B95FEAB3FD20FFE3FE1DFA0173C9F9CE" box="[478,650,1526,1552]" italics="true" pageId="3" pageNumber="491">Z. sichuanensis</emphasis>
</taxonomicName>
, surface of pronotum and elytral intervals smooth and shiny, subapical part of penis distinctly inflated and apex of penis arrowhead-like.
</paragraph>
</subSubSection>
<subSubSection id="C331652AFD20FFE5FF04F9C875EBFE73" lastPageId="5" lastPageNumber="493" pageId="3" pageNumber="491" type="description">
<paragraph id="8B9436A1FD20FFE3FF04F9C870EDF91A" blockId="3.[151,1437,842,1949]" pageId="3" pageNumber="491">
<emphasis id="B95FEAB3FD20FFE3FF04F9C87012F986" bold="true" box="[199,337,1598,1624]" pageId="3" pageNumber="491">Description</emphasis>
. BL 3.0 mm, EW
<quantity id="4CD39B44FD20FFE3FDEAF9C873C2F987" box="[553,641,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="491" unit="mm" value="1.2">1.2 mm</quantity>
(
<typeStatus id="54908803FD20FFE3FD52F9C873B9F986" box="[657,762,1598,1624]" pageId="3" pageNumber="491" type="holotype">holotype</typeStatus>
). Habitus see fig. 3A–B. Head and pronotum black, elytra, femora, tibiae and ventral surface brown. Tarsi light brown. Antennae yellowish brown. Labrum wider than long; surface smooth and shiny, densely pubescent; lateral margin with long yellowish fringe. Clypeus and frons densely granulate and pubescent.
</paragraph>
<paragraph id="8B9436A1FD20FFE3FF04F93975A7F843" blockId="3.[151,1437,842,1949]" pageId="3" pageNumber="491">
Pronotum (fig. 3C) broadest at basal 0.4, distinctly attenuated anteriorly and slightly attenuated posteriorly. PL
<quantity id="4CD39B44FD20FFE3FF02F904705EF8D3" box="[193,285,1778,1805]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="491" unit="mm" value="0.8">0.8 mm</quantity>
, PW
<quantity id="4CD39B44FD20FFE3FEA3F90470FFF8D3" box="[352,444,1778,1805]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="491" unit="mm" value="0.9">0.9 mm</quantity>
. Disc of basal third coarse, and base with some granules medially. Disc of distal 2/3 smooth and shiny, punctures moderately dense. Lateral areas coarse, densely granulate. Median longitudinal sulcus is present from basal 0.2 to 0.7, or even shorter, broadest at middle, then gradually narrowed anteriorly and posteriorly. Sublateral carinae present in basal 0.4, subparallel in 0.3, then bend outwards. Anterior angles sharp, slightly produced, and posterior angles almost right-angled. Anterior half of lateral margin serrate.
</paragraph>
<paragraph id="8B9436A1FD27FFE4FF04FF6170EEFF27" blockId="4.[151,1437,151,249]" pageId="4" pageNumber="492">Elytra broadest at basal 2/3, slightly attenuated anteriorly and distinctly attenuated posteriorly. Strial punctures small and deeply impressed. Intervals slightly convex, rugose. Intervals 1, 3, 5 with a row of longitudinal pubescence. Intervals 5, 7, 8 carinate.</paragraph>
<caption id="DF546629FD27FFE4FF54F9037365F8EF" ID-DOI="http://doi.org/10.5281/zenodo.10562288" ID-Zenodo-Dep="10562288" httpUri="https://zenodo.org/record/10562288/files/figure.png" pageId="4" pageNumber="492" startId="4.[151,250,1781,1805]" targetBox="[151,1435,277,1758]" targetPageId="4" targetType="figure">
<paragraph id="8B9436A1FD27FFE4FF54F9037365F8EF" blockId="4.[151,1436,1781,1841]" pageId="4" pageNumber="492">
<emphasis id="B95FEAB3FD27FFE4FF54F9037057F8D3" bold="true" box="[151,276,1781,1805]" pageId="4" pageNumber="492">FIGURE 3.</emphasis>
<taxonomicName id="4C2B4D22FD27FFE4FED8F90070BBF8D3" authority="Peng &amp; Bian &amp; Wang, 2024" authorityName="Peng &amp; Bian &amp; Wang" authorityYear="2024" box="[283,504,1781,1805]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="492" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
<emphasis id="B95FEAB3FD27FFE4FED8F90070BBF8D3" box="[283,504,1781,1805]" italics="true" pageId="4" pageNumber="492">Zaitzevia triangularis</emphasis>
</taxonomicName>
<emphasis id="B95FEAB3FD27FFE4FE3CF903731BF8D3" bold="true" box="[511,600,1781,1805]" pageId="4" pageNumber="492">
<taxonomicNameLabel id="A26C57C8FD27FFE4FE3CF903737AF8D3" box="[511,569,1781,1805]" pageId="4" pageNumber="492" rank="species">sp. n.</taxonomicNameLabel>
A.
</emphasis>
Habitus, holotype, male, in dorsal view.
<emphasis id="B95FEAB3FD27FFE4FC3EF9037557F8D3" bold="true" box="[1021,1044,1781,1805]" pageId="4" pageNumber="492">B.</emphasis>
Habitus, in ventral view
<emphasis id="B95FEAB3FD27FFE4FADBF9037469F8D3" bold="true" box="[1304,1322,1781,1805]" pageId="4" pageNumber="492">C</emphasis>
. Pronotum
<emphasis id="B95FEAB3FD27FFE4FF54F8EF71EAF8EF" bold="true" box="[151,169,1817,1841]" pageId="4" pageNumber="492">D</emphasis>
. Prosternal process and metaventrite.
</paragraph>
</caption>
<paragraph id="8B9436A1FD27FFE4FF04F8A970DAF83B" blockId="4.[151,1436,1887,2021]" pageId="4" pageNumber="492">Prosternal process (fig. 3D) elongated and slim, gradually narrowed from base to apex; lateral margin distinctly rimmed; apex broadly rounded; surface coarsely granulated. Disc of metaventrite (fig. 3D) broadly impressed, smooth and shiny, with some sparse punctures; median groove present in posterior 0.8; lateral area densely pubescent and sparsely granulate.</paragraph>
<paragraph id="8B9436A1FD26FFE5FF04FF617411FF27" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="493">Disc of ventrite I coarse, sparsely granulate, and disc of ventrite II densely punctate and sparsely granulate. Disc of ventrites III to IV, and basal 1/3 of ventrite V sparsely punctate and granulate. Lateral areas of ventrites I to IV densely pubescent and sparsely granulate. Distal 2/3 of ventrite V densely granulate and sparsely pubescent.</paragraph>
<paragraph id="8B9436A1FD26FFE5FF04FEF572FAFEBB" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="493">
Aedeagus (figs. 4A–C).
<quantity id="4CD39B44FD26FFE5FE23FEF5737AFEC3" box="[480,569,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="493" unit="mm" value="1.4">1.4 mm</quantity>
long. Penis about 2.4 times as long as phallobase, subparallel in basal 0.6, distinctly narrowed in distal 0.4, with a pair of developed teeth at distal 0.25. Endophallus simple. Parameres completely fused to penis; apices of parameres reach basal 0.6 of penis.
</paragraph>
<paragraph id="8B9436A1FD26FFE5FF04FE9975C4FE57" blockId="5.[151,1437,151,537]" box="[199,1159,367,393]" pageId="5" pageNumber="493">
Males: BL
<quantity id="4CD39B44FD26FFE5FE87FE99708EFE57" box="[324,461,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="5" pageNumber="493" unit="mm" value="3.05" valueMax="3.2" valueMin="2.9">2.9–3.2 mm</quantity>
, EW
<quantity id="4CD39B44FD26FFE5FDC8FE9973D1FE57" box="[523,658,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="5" pageNumber="493" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
(n=13). Apices of elytra separately rounded.
</paragraph>
<paragraph id="8B9436A1FD26FFE5FF04FE6575EBFE73" blockId="5.[151,1437,151,537]" box="[199,1192,403,429]" pageId="5" pageNumber="493">
Females: BL 3.0–
<quantity id="4CD39B44FD26FFE5FE4DFE6570A5FE73" box="[398,486,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="493" unit="mm" value="3.2">3.2 mm</quantity>
, EW
<quantity id="4CD39B44FD26FFE5FDE7FE6573E8FE73" box="[548,683,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="5" pageNumber="493" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2–1.3 mm</quantity>
(n=4). Apices of elytra separately sharpened.
</paragraph>
</subSubSection>
<subSubSection id="C331652AFD26FFE5FF04FE40734DFE0F" box="[199,526,438,465]" pageId="5" pageNumber="493" type="distribution">
<paragraph id="8B9436A1FD26FFE5FF04FE40734DFE0F" blockId="5.[151,1437,151,537]" box="[199,526,438,465]" pageId="5" pageNumber="493">
<emphasis id="B95FEAB3FD26FFE5FF04FE40701AFE0E" bold="true" box="[199,345,438,464]" pageId="5" pageNumber="493">Distribution</emphasis>
.
<collectingCountry id="F33C7631FD26FFE5FEA4FE4170EDFE0F" box="[359,430,439,465]" name="China" pageId="5" pageNumber="493">China</collectingCountry>
:
<collectingRegion id="49EFF843FD26FFE5FE7AFE417349FE0F" box="[441,522,439,465]" country="China" name="Shanxi" pageId="5" pageNumber="493">Shanxi</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C331652AFD26FFE5FF04FE2D71BBFDC7" pageId="5" pageNumber="493" type="etymology">
<paragraph id="8B9436A1FD26FFE5FF04FE2D71BBFDC7" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="493">
<emphasis id="B95FEAB3FD26FFE5FF04FE2D700DFE2B" bold="true" box="[199,334,475,501]" pageId="5" pageNumber="493">Etymology.</emphasis>
The epithet is derived from the Latin adjective
<taxonomicName id="4C2B4D22FD26FFE5FC9AFE2D72A4FE2B" authorityName="Peng &amp; Bian &amp; Wang" authorityYear="2024" box="[857,999,475,501]" class="Insecta" family="Elmidae" genus="Zaitzevia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="493" phylum="Arthropoda" rank="species" species="triangularis">“triangularis</taxonomicName>
” = triangular, refers to the apex’s shape of penis.
</paragraph>
</subSubSection>
</treatment>
</document>
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