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<document id="9A6E8A1AD18D51545FAA0CB130E030FA" ID-DOI="10.1080/00222930802277616" ID-ISSN="1464-5262" ID-Zenodo-Dep="5223730" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629391701861" checkinUser="felipe" docAuthor="Csuzdi, Csaba" docDate="2008" docId="038287A1F315FFA96ABCFB42FBE1CA42" docLanguage="en" docName="JNatHist.42.33-34.2227-2241.pdf" docOrigin="Journal of Natural History 42 (33 - 34)" docSource="http://dx.doi.org/10.1080/00222930802277616" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Dichogaster (Diplothecodrilus) gasanai Csuzdi 2008, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2233" masterDocId="FFBBFFD9F310FFAE6B16FF8BFFA6CE6E" masterDocTitle="Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae)" masterLastPageNumber="2241" masterPageNumber="2227" pageNumber="2231" updateTime="1699071437077" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="FE5E89AF3414E988F75D884E9A74DBBE">
<mods:title id="FBCA3712EFA31CD3078912DDE83005BA">Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae)</mods:title>
</mods:titleInfo>
<mods:name id="5897AEC9501EB458A1AE208454FFE9A1" type="personal">
<mods:role id="0B4574F1CED6E7B8D068F1B10FA8CCD8">
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</mods:role>
<mods:namePart id="AEEA0B08598F0AEA11A317EAF0F8DDE3">Csuzdi, Csaba</mods:namePart>
</mods:name>
<mods:typeOfResource id="8B2FBCA52062EF04F0E78519BF6082AD">text</mods:typeOfResource>
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<mods:titleInfo id="3412CDB1B0149C915CAB3748FCBBC9A0">
<mods:title id="EF5DD07336D38F56F756ED7637DB3658">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="0D362309D12994B85C1E5B0500839CDD">
<mods:date id="21D91F1E8B3A8604FA740A9478B7DB24">2008</mods:date>
<mods:detail id="1234E0448C709223DEC3BC8A56C34E71" type="pubDate">
<mods:number id="40A76F8EED0F97189C2E0F6ED961CD82">2008-09-30</mods:number>
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<mods:number id="6039B84E85D6704CC3BF98413BB0CD95">42</mods:number>
</mods:detail>
<mods:detail id="11C15B92F396EE47A515BB5A77A18267" type="issue">
<mods:number id="175A1336F093655A4FED55EAE2B544E0">33 - 34</mods:number>
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<mods:start id="C481F59147C7477EB3E3A0EBF1B18780">2227</mods:start>
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<mods:url id="74FB4305111054B2356C109275AFB9CB">http://dx.doi.org/10.1080/00222930802277616</mods:url>
</mods:location>
<mods:classification id="844806B4E1AED97BA8482B557AF4CE55">journal article</mods:classification>
<mods:identifier id="D6F4ACD3728C557491FDF5C8AC4C1F97" type="DOI">10.1080/00222930802277616</mods:identifier>
<mods:identifier id="446073FBDE4658B6E7A435589E972434" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="D277B4FBAF35E91298EB0DA2EB35216F" type="Zenodo-Dep">5223730</mods:identifier>
</mods:mods>
<treatment id="038287A1F315FFA96ABCFB42FBE1CA42" LSID="urn:lsid:plazi:treatment:038287A1F315FFA96ABCFB42FBE1CA42" httpUri="http://treatment.plazi.org/id/038287A1F315FFA96ABCFB42FBE1CA42" lastPageId="7" lastPageNumber="2233" pageId="5" pageNumber="2231">
<subSubSection id="C331653CF315FFAB6ABCFB42FC1ACA8F" box="[426,956,1225,1249]" pageId="5" pageNumber="2231" type="nomenclature">
<paragraph id="8B9436B7F315FFAB6ABCFB42FC1ACA8F" blockId="5.[426,956,1225,1281]" box="[426,956,1225,1249]" pageId="5" pageNumber="2231">
<heading id="D0DC81DBF315FFAB6ABCFB42FC1ACA8F" box="[426,956,1225,1249]" centered="true" fontSize="36" level="2" pageId="5" pageNumber="2231" reason="7">
<taxonomicName id="4C2B4D34F315FFAB6ABCFB42FCFDCA8F" authority="Csuzdi, 2008" authorityName="Csuzdi" authorityYear="2008" box="[426,859,1225,1249]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="2231" phylum="Annelida" rank="species" species="gasanai" status="sp. nov." subGenus="Diplothecodrilus">
<emphasis id="B95FEAA5F315FFAB6ABCFB42FCFDCA8F" bold="true" box="[426,859,1225,1249]" italics="true" pageId="5" pageNumber="2231">Dichogaster (Diplothecodrilus) gasanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57DEF315FFAB6875FB42FC1ACA8F" box="[867,956,1225,1249]" pageId="5" pageNumber="2231" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331653CF315FFA96977FB62FBE1CA42" lastPageId="7" lastPageNumber="2233" pageId="5" pageNumber="2231" type="description">
<paragraph id="8B9436B7F315FFAB6977FB62FCA2CB6F" blockId="5.[426,956,1225,1281]" box="[609,772,1257,1281]" pageId="5" pageNumber="2231">
(
<figureCitation id="13102A32F315FFAB697FFB62FD5BCB6F" box="[617,765,1257,1281]" captionStart="Figure 3" captionStartId="6.[146,211,861,883]" captionTargetBox="[146,1101,165,824]" captionTargetId="figure-189@6.[146,1104,165,824]" captionTargetPageId="6" captionText="Figure 3. Dichogaster (Dt.) gasanai sp.nov. (A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores." figureDoi="http://doi.org/10.5281/zenodo.5223736" httpUri="https://zenodo.org/record/5223736/files/figure.png" pageId="5" pageNumber="2231">Figure 3AD</figureCitation>
)
</paragraph>
<paragraph id="8B9436B7F315FFAB6BC3FAB1FE0FCB3C" blockId="5.[213,425,1338,1362]" box="[213,425,1338,1362]" pageId="5" pageNumber="2231">
<emphasis id="B95FEAA5F315FFAB6BC3FAB1FE0FCB3C" box="[213,425,1338,1362]" italics="true" pageId="5" pageNumber="2231">Material examined</emphasis>
</paragraph>
<paragraph id="8B9436B7F315FFAB6BC3FAE8FD58CBF4" blockId="5.[213,1169,1378,1434]" pageId="5" pageNumber="2231">
<materialsCitation id="3B433CEAF315FFAB6BC3FAE8FD58CBF4" collectingDate="1973-07-15" collectionCode="R" collectorName="Gasana N'Doli" country="Democratic Republic of the Congo" location="Kivu Region" pageId="5" pageNumber="2231" specimenCode="BMNH 1997.1561" specimenCount="1" typeStatus="holotype">
<emphasis id="B95FEAA5F315FFAB6BC3FAE8FE9ACB15" box="[213,316,1379,1403]" italics="true" pageId="5" pageNumber="2231">
<typeStatus id="54908815F315FFAB6BC3FAE8FE9ACB15" box="[213,316,1379,1403]" pageId="5" pageNumber="2231" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="DB8D9ECCF315FFAB6A46FAE9FD8DCB14" box="[336,555,1378,1402]" collectionCode="BMNH" pageId="5" pageNumber="2231">BMNH 1997.1561</specimenCode>
. D.
<collectionCode id="ED3AAE72F315FFAB694EFAE9FDD6CB14" box="[600,624,1378,1402]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="2231">R</collectionCode>
.
<collectingCountry id="F33C7627F315FFAB6997FAE9FD76CB14" box="[641,720,1378,1402]" name="Democratic Republic of the Congo" pageId="5" pageNumber="2231">Congo</collectingCountry>
,
<location id="8EF4606CF315FFAB69F7FAE9FCDACB14" LSID="urn:lsid:plazi:treatment:038287A1F315FFA96ABCFB42FBE1CA42:8EF4606CF315FFAB69F7FAE9FCDACB14" box="[737,892,1378,1402]" country="Democratic Republic of the Congo" name="Kivu Region" pageId="5" pageNumber="2231">Kivu Region</location>
, Butembo, in stems of banana trees, leg.
<collectorName id="26DE5361F315FFAB6AB0FA09FDFFCBF4" box="[422,601,1410,1434]" pageId="5" pageNumber="2231">Gasana NDoli</collectorName>
,
<date id="FF951077F315FFAB6970FA09FD5CCBF4" box="[614,762,1410,1434]" pageId="5" pageNumber="2231" value="1973-07-15">
<collectingDate id="EFD1E99FF315FFAB6970FA09FD5CCBF4" box="[614,762,1410,1434]" pageId="5" pageNumber="2231" value="1973-07-15">15 July 1973</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9436B7F315FFA86BC3FA58FB93CD82" blockId="5.[213,1170,1491,1706]" lastBlockId="6.[146,1103,948,1004]" lastPageId="6" lastPageNumber="2232" pageId="5" pageNumber="2231">
<materialsCitation id="3B433CEAF315FFAB6BC3FA58FD43CB85" box="[213,741,1491,1515]" collectionCode="HNHM" pageId="5" pageNumber="2231" specimenCount="1" typeStatus="paratype">
<emphasis id="B95FEAA5F315FFAB6BC3FA58FEE3CB85" box="[213,325,1491,1515]" italics="true" pageId="5" pageNumber="2231">
<typeStatus id="54908815F315FFAB6BC3FA58FEE3CB85" box="[213,325,1491,1515]" pageId="5" pageNumber="2231" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectionCode id="ED3AAE72F315FFAB6A4DFA58FE11CB85" box="[347,439,1491,1515]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="2231">HNHM</collectionCode>
AF/3564, one exemplar
</materialsCitation>
,
<materialsCitation id="3B433CEAF315FFAB69EFFA58FD84C865" collectionCode="BMNH" pageId="5" pageNumber="2231" specimenCode="BMNH 1997.1562" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8D9ECCF315FFAB69EFFA58FC73CB85" box="[761,981,1491,1515]" collectionCode="BMNH" pageId="5" pageNumber="2231">BMNH 1997.1562</specimenCode>
, one exemplar. Locality same as that of the
</materialsCitation>
<materialsCitation id="3B433CEAF315FFA8693BFA78FB94CD82" collectingDate="1973-07-15" collectingDateMax="1973-07-25" collectingDateMin="1973-07-15" collectionCode="HNHM, R" collectorName="Bugabo &amp; Gasana N'Doli &amp; Mewa" country="Democratic Republic of the Congo" county="Kivu Region" lastPageId="6" lastPageNumber="2232" location="Kivu Region" municipality="River Mululu" pageId="5" pageNumber="2231" specimenCode="BMNH 1997.1618, 1629, 12, BMNH 1997.1607, 1616, BMNH 1997.1574, 1582, BMNH 1997.1479, 1484" specimenCount="1" typeStatus="holotype">
<typeStatus id="54908815F315FFAB693BFA78FD3AC865" box="[557,668,1523,1547]" pageId="5" pageNumber="2231" type="holotype">Holotype</typeStatus>
.
<collectionCode id="ED3AAE72F315FFAB69BCFA78FCA0C865" box="[682,774,1523,1547]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="2231">HNHM</collectionCode>
AF/3561, ten exemplars,
<specimenCode id="DB8D9ECCF315FFAB6F2FFA78FEEFC844" collectionCode="BMNH" pageId="5" pageNumber="2231">BMNH 1997.1618</specimenCode>
<specimenCode id="DB8D9ECCF315FFAB6A41F999FE2AC844" box="[343,396,1554,1578]" collectionCode="BMNH" pageId="5" pageNumber="2231">1629</specimenCode>
,
<specimenCode id="DB8D9ECCF315FFAB6A8BF999FE1EC844" box="[413,440,1554,1578]" collectionCode="BMNH" pageId="5" pageNumber="2231">12</specimenCode>
exemplars D.
<collectionCode id="ED3AAE72F315FFAB6974F999FDDCC844" box="[610,634,1554,1578]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="2231">R</collectionCode>
.
<collectingCountry id="F33C7627F315FFAB699DF999FD7CC844" box="[651,730,1554,1578]" name="Democratic Republic of the Congo" pageId="5" pageNumber="2231">Congo</collectingCountry>
,
<location id="8EF4606CF315FFAB69FAF999FC21C844" LSID="urn:lsid:plazi:treatment:038287A1F315FFA96ABCFB42FBE1CA42:8EF4606CF315FFAB69FAF999FC21C844" box="[748,903,1554,1578]" country="Democratic Republic of the Congo" county="Kivu Region" municipality="River Mululu" name="Kivu Region" pageId="5" pageNumber="2231">Kivu Region</location>
,
<collectorName id="26DE5361F315FFAB688FF999FC52C844" box="[921,1012,1554,1578]" pageId="5" pageNumber="2231">Bugabo</collectorName>
, leg.
<collectorName id="26DE5361F315FFAB6F2CF999FE8EC824" pageId="5" pageNumber="2231">Gasana NDoli</collectorName>
,
<date id="FF951077F315FFAB6A20F9B9FE6BC824" box="[310,461,1586,1610]" pageId="5" pageNumber="2231" value="1973-07-25">
<collectingDate id="EFD1E99FF315FFAB6A20F9B9FE6BC824" box="[310,461,1586,1610]" pageId="5" pageNumber="2231" value="1973-07-25">25 July 1973</collectingDate>
</date>
.
<collectionCode id="ED3AAE72F315FFAB6ACDF9B9FD91C824" box="[475,567,1586,1610]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="2231">HNHM</collectionCode>
AF/3550, four exemplars,
<specimenCode id="DB8D9ECCF315FFAB6865F9B9FBEFC824" box="[883,1097,1586,1610]" collectionCode="BMNH" pageId="5" pageNumber="2231">BMNH 1997.1607</specimenCode>
<specimenCode id="DB8D9ECCF315FFAB6F4EF9B9FB2BC824" box="[1112,1165,1586,1610]" collectionCode="BMNH" pageId="5" pageNumber="2231">1616</specimenCode>
, ten exemplars D.
<collectionCode id="ED3AAE72F315FFAB6AB3F9D9FE1BC804" box="[421,445,1618,1642]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="2231">R</collectionCode>
.
<collectingCountry id="F33C7627F315FFAB6AC6F9D9FDB9C804" box="[464,543,1618,1642]" name="Democratic Republic of the Congo" pageId="5" pageNumber="2231">Congo</collectingCountry>
,
<location id="8EF4606CF315FFAB6925F9D9FD76C804" LSID="urn:lsid:plazi:treatment:038287A1F315FFA96ABCFB42FBE1CA42:8EF4606CF315FFAB6925F9D9FD76C804" box="[563,720,1618,1642]" country="Democratic Republic of the Congo" county="Kivu Region" municipality="River Mululu" name="Kivu Region" pageId="5" pageNumber="2231">Kivu Region</location>
,
<locationDeviation id="9A262F95F315FFAB69F5F9D9FBF1C804" box="[739,1111,1618,1642]" firstLocation="Kagundu" pageId="5" pageNumber="2231" secondLocation="Gitovu">between Kagundu and Gitovu</locationDeviation>
, leg.
<collectorName id="26DE5361F315FFAB6BC3F9F9FE23C8E4" box="[213,389,1650,1674]" pageId="5" pageNumber="2231">Gasana NDoli</collectorName>
,
<date id="FF951077F315FFAB6A86F9F9FDB9C8E4" box="[400,543,1650,1674]" pageId="5" pageNumber="2231" value="1973-07-17">
<collectingDate id="EFD1E99FF315FFAB6A86F9F9FDB9C8E4" box="[400,543,1650,1674]" pageId="5" pageNumber="2231" value="1973-07-17">17 July 1973</collectingDate>
</date>
.
<collectionCode id="ED3AAE72F315FFAB693FF9F9FD23C8E4" box="[553,645,1650,1674]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="2231">HNHM</collectionCode>
AF/3551, four exemplars,
<specimenCode id="DB8D9ECCF315FFAB68A2F9F9FB25C8E4" box="[948,1155,1650,1674]" collectionCode="BMNH" pageId="5" pageNumber="2231">BMNH 1997.1574</specimenCode>
<specimenCode id="DB8D9ECCF315FFAB6BC3F919FEABC8C4" box="[213,269,1682,1706]" collectionCode="BMNH" pageId="5" pageNumber="2231">1582</specimenCode>
, nine exemplars. D.
<collectionCode id="ED3AAE72F315FFAB6AFDF919FDA5C8C4" box="[491,515,1682,1706]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="2231">R</collectionCode>
.
<collectingCountry id="F33C7627F315FFAB691BF919FDFAC8C4" box="[525,604,1682,1706]" name="Democratic Republic of the Congo" pageId="5" pageNumber="2231">Congo</collectingCountry>
,
<location id="8EF4606CF315FFAB6971F919FD5AC8C4" LSID="urn:lsid:plazi:treatment:038287A1F315FFA96ABCFB42FBE1CA42:8EF4606CF315FFAB6971F919FD5AC8C4" box="[615,764,1682,1706]" country="Democratic Republic of the Congo" county="Kivu Region" municipality="River Mululu" name="Kivu Region" pageId="5" pageNumber="2231">Kivu Region</location>
,
<collectingMunicipality id="6BF0ACCDF315FFAB6811F919FC02C8C4" box="[775,932,1682,1706]" pageId="5" pageNumber="2231">River Mululu</collectingMunicipality>
, leg.
<collectorName id="26DE5361F315FFAB68CAF919FB2BC8C4" box="[988,1165,1682,1706]" pageId="5" pageNumber="2231">Gasana NDoli</collectorName>
,
<date id="FF951077F316FFA86B84FC3FFE9ACDA2" box="[146,316,948,972]" pageId="6" pageNumber="2232" value="1973-07-15">
<collectingDate id="EFD1E99FF316FFA86B84FC3FFE9ACDA2" box="[146,316,948,972]" pageId="6" pageNumber="2232" value="1973-07-15">15. July 1973</collectingDate>
</date>
.
<collectionCode id="ED3AAE72F316FFA86A46FC3FFE0ACDA2" box="[336,428,948,972]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="2232">HNHM</collectionCode>
AF/3557, two exemplars,
<specimenCode id="DB8D9ECCF316FFA869EAFC3FFC7ECDA2" box="[764,984,948,972]" collectionCode="BMNH" pageId="6" pageNumber="2232">BMNH 1997.1479</specimenCode>
<specimenCode id="DB8D9ECCF316FFA868F1FC3FFBBACDA2" box="[999,1052,948,972]" collectionCode="BMNH" pageId="6" pageNumber="2232">1484</specimenCode>
, six exemplars. D.
<collectionCode id="ED3AAE72F316FFA86A27FC5FFEEFCD82" box="[305,329,980,1004]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="2232">R</collectionCode>
.
<collectingCountry id="F33C7627F316FFA86A40FC5FFE03CD82" box="[342,421,980,1004]" name="Democratic Republic of the Congo" pageId="6" pageNumber="2232">Congo</collectingCountry>
,
<collectingCounty id="62F54E3BF316FFA86AA5FC5FFDECCD82" box="[435,586,980,1004]" pageId="6" pageNumber="2232">Kivu Region</collectingCounty>
,
<collectorName id="26DE5361F316FFA86941FC5FFD06CD82" box="[599,672,980,1004]" pageId="6" pageNumber="2232">Mewa</collectorName>
, leg.
<collectorName id="26DE5361F316FFA869C8FC5FFC36CD82" box="[734,912,980,1004]" pageId="6" pageNumber="2232">Gasana NDoli</collectorName>
,
<date id="FF951077F316FFA8688BFC5FFB94CD82" box="[925,1074,980,1004]" pageId="6" pageNumber="2232" value="1973-07-17">
<collectingDate id="EFD1E99FF316FFA8688BFC5FFB94CD82" box="[925,1074,980,1004]" pageId="6" pageNumber="2232" value="1973-07-17">17 July 1973</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="DF54663FF316FFA86B84FCD6FBF6CDFE" ID-DOI="http://doi.org/10.5281/zenodo.5223736" ID-Zenodo-Dep="5223736" httpUri="https://zenodo.org/record/5223736/files/figure.png" pageId="6" pageNumber="2232" startId="6.[146,211,861,883]" targetBox="[146,1101,165,824]" targetPageId="6">
<paragraph id="8B9436B7F316FFA86B84FCD6FBF6CDFE" blockId="6.[146,1104,861,912]" pageId="6" pageNumber="2232">
Figure 3.
<taxonomicName id="4C2B4D34F316FFA86BE0FCD6FE55CD1C" authority="Csuzdi, 2008" authorityName="Csuzdi" authorityYear="2008" box="[246,499,861,882]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="6" pageNumber="2232" phylum="Annelida" rank="species" species="gasanai" status="sp. nov." subGenus="Diplothecodrilus">
<emphasis id="B95FEAA5F316FFA86BE0FCD6FE55CD1C" box="[246,499,861,882]" italics="true" pageId="6" pageNumber="2232">Dichogaster (Dt.) gasanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57DEF316FFA86AEEFCD6FDE3CD1D" box="[504,581,861,883]" pageId="6" pageNumber="2232" rank="species">sp. nov.</taxonomicNameLabel>
(A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores.
</paragraph>
</caption>
<paragraph id="8B9436B7F316FFA86B84FBAEFEFECA53" blockId="6.[146,344,1061,1085]" box="[146,344,1061,1085]" pageId="6" pageNumber="2232">
<emphasis id="B95FEAA5F316FFA86B84FBAEFEFECA53" box="[146,344,1061,1085]" italics="true" pageId="6" pageNumber="2232">Derivatio nominis</emphasis>
</paragraph>
<paragraph id="8B9436B7F316FFA86B84FBC5FBE9CA08" blockId="6.[146,1103,1102,1126]" box="[146,1103,1102,1126]" pageId="6" pageNumber="2232">The specific epithet is referring to the collector of this interesting material:Gasana NDoli.</paragraph>
<paragraph id="8B9436B7F316FFA86B84FB1CFF59CAC1" blockId="6.[146,255,1175,1199]" box="[146,255,1175,1199]" pageId="6" pageNumber="2232">
<emphasis id="B95FEAA5F316FFA86B84FB1CFF59CAC1" box="[146,255,1175,1199]" italics="true" pageId="6" pageNumber="2232">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B9436B7F316FFA86B84FB34FC96CBF9" blockId="6.[146,1103,1215,1431]" pageId="6" pageNumber="2232">
Length
<quantity id="4CD39B52F316FFA86BFFFB34FEF9CAB9" box="[233,351,1215,1239]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.6" metricValueMin="3.2" pageId="6" pageNumber="2232" unit="mm" value="44.0" valueMax="56.0" valueMin="32.0">3256 mm</quantity>
, D.
<quantity id="4CD39B52F316FFA86A9BFB34FE5BCAB9" box="[397,509,1215,1239]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="6" pageNumber="2232" unit="mm" value="1.9" valueMax="2.0" valueMin="1.8">1.82 mm</quantity>
, no. of segments 75112. Pigmentation lacking. First dorsal pore in 5/6. Clitellum 12
<emphasis id="B95FEAA5F316FFA86917FB54FDBACA96" bold="true" box="[513,540,1247,1272]" pageId="6" pageNumber="2232">K</emphasis>
21.
<specimenCount id="9D2DFD3EF316FFA86953FB55FDF5CA99" box="[581,595,1246,1271]" pageId="6" pageNumber="2232" type="female"></specimenCount>
14 paired between setae aa. Prostatic pores 17, 19. Spermathecal pores 7/8, 8/9, spermathecae with elongated ampoule, and a shorter duct, half as long as the ampoule. Gizzard in 5/6, last pair of hearts in 13. Calciferous glands in 1517, with slightly backwards increasing size. Excretory system meroic with four meronephridia per side. Penial seta L.
<quantity id="4CD39B52F316FFA8688AFAD4FB86CB19" box="[924,1056,1375,1399]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="6" pageNumber="2232" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
, D.
<quantity id="4CD39B52F316FFA86B84FAF4FEA4CBF9" box="[146,258,1407,1431]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="2232" unit="mm" value="0.006">0.006 mm</quantity>
, tip sharply pointed, ornamentation small teeth.
</paragraph>
<paragraph id="8B9436B7F316FFA86B84FA42FEC8CB8F" blockId="6.[146,366,1481,1505]" box="[146,366,1481,1505]" pageId="6" pageNumber="2232">
<emphasis id="B95FEAA5F316FFA86B84FA42FEC8CB8F" box="[146,366,1481,1505]" italics="true" pageId="6" pageNumber="2232">External characters</emphasis>
</paragraph>
<paragraph id="8B9436B7F316FFA86B84FA7AFCE2C867" blockId="6.[146,836,1521,1545]" box="[146,836,1521,1545]" pageId="6" pageNumber="2232">
<emphasis id="B95FEAA5F316FFA86B84FA7AFF5FC867" box="[146,249,1521,1545]" italics="true" pageId="6" pageNumber="2232">
<typeStatus id="54908815F316FFA86B84FA7AFF5FC867" box="[146,249,1521,1545]" pageId="6" pageNumber="2232" type="holotype">Holotype</typeStatus>
</emphasis>
. Length
<quantity id="4CD39B52F316FFA86A75FA7AFE17C867" box="[355,433,1521,1545]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="2232" unit="mm" value="48.0">48 mm</quantity>
, D.
<quantity id="4CD39B52F316FFA86AF2FA7AFD82C867" box="[484,548,1521,1545]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="2232" unit="mm" value="2.0">2 mm</quantity>
, number of segments 96.
</paragraph>
<paragraph id="8B9436B7F316FFA96B84F9B9FD3BCF4C" blockId="6.[146,1103,1586,1706]" lastBlockId="7.[213,1169,168,290]" lastPageId="7" lastPageNumber="2233" pageId="6" pageNumber="2232">
<emphasis id="B95FEAA5F316FFA86B84F9B9FEA4C824" box="[146,258,1586,1610]" italics="true" pageId="6" pageNumber="2232">
<typeStatus id="54908815F316FFA86B84F9B9FEA4C824" box="[146,258,1586,1610]" pageId="6" pageNumber="2232" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<quantity id="4CD39B52F316FFA86A04F9B9FE23C824" box="[274,389,1586,1610]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="5.6" metricValueMin="3.2" pageId="6" pageNumber="2232" unit="mm" value="44.0" valueMax="56.0" valueMin="32.0">3256 mm</quantity>
in length,
<quantity id="4CD39B52F316FFA86917F9B9FDC8C824" box="[513,622,1586,1610]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="6" pageNumber="2232" unit="mm" value="1.9" valueMax="2.0" valueMin="1.8">1.82 mm</quantity>
in diameter, number of segments 75112. Colour pale, pigmentation lacking. Prostomium schizolobous. First dorsal pore in 5/6. Setae all ventral, setal arrangement after the clitellum aa:ab:bc:cd: dd52.5:1.25:3.75:1:25. Clitellum extends over segments 12
<emphasis id="B95FEAA5F316FFA8688DF91AFC10C8C4" bold="true" box="[923,950,1681,1706]" pageId="6" pageNumber="2232">K</emphasis>
21, with a butterfly-shaped male field. Prostatic pores paired on 17, 19 in the setal line b. Male pores externally not visible, they are on 18, in the seminal grooves. Female pores paired on 14 between setae aa. Spermathecal pores paired in 7/8, 8/9 near to the setae b. Porophores absent (
<figureCitation id="13102A32F317FFA9690DFE81FD37CF4C" box="[539,657,266,290]" captionStart="Figure 3" captionStartId="6.[146,211,861,883]" captionTargetBox="[146,1101,165,824]" captionTargetId="figure-189@6.[146,1104,165,824]" captionTargetPageId="6" captionText="Figure 3. Dichogaster (Dt.) gasanai sp.nov. (A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores." figureDoi="http://doi.org/10.5281/zenodo.5223736" httpUri="https://zenodo.org/record/5223736/files/figure.png" pageId="7" pageNumber="2233">Figure 3A</figureCitation>
).
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3FED8FE0ECF05" blockId="7.[213,424,339,363]" box="[213,424,339,363]" pageId="7" pageNumber="2233">
<emphasis id="B95FEAA5F317FFA96BC3FED8FE0ECF05" box="[213,424,339,363]" italics="true" pageId="7" pageNumber="2233">Internal characters</emphasis>
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3FEF0FC90CD37" blockId="7.[213,1169,379,857]" pageId="7" pageNumber="2233">
The first visible septum 6/7, thickened septa lacking. Oesophageal gizzards small, in segment 5 and 6. Three pairs of calciferous glands located in segment 1517, their size slightly increasing backwards. Excretory system meroic, with four meronephridia on each side. Paired hearts are present in segments 1013. Typhlosole small, begins in segment 20 and accompanied by a pair of auxiliary typhlosoles between segments 27 and 33. Testes in 10 and 11 enclosed in perioesophageal testis sacs. Seminal vesicles in 11, 12, and a pair of large ovaries in segment 13. Seminal duct convoluted, discharging in segment 18. Two pairs of prostatic glands are present in 17 and 19, slightly intruding on the neighbouring segments and each with a penial setal sac containing a mature and several immature penial setae. The mature penial seta is somewhat undulated, about
<quantity id="4CD39B52F317FFA96995FD34FCA2CCB9" box="[643,772,703,727]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="7" pageNumber="2233" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.41.5 mm</quantity>
long and
<quantity id="4CD39B52F317FFA9689FFD34FC53CCB9" box="[905,1013,703,727]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="2233" unit="mm" value="0.006">0.006 mm</quantity>
wide with a slightly hooked tip. Ornamentation consists of scattered small teeth (
<figureCitation id="13102A32F317FFA968E1FD6BFB25CC96" box="[1015,1155,736,760]" captionStart="Figure 3" captionStartId="6.[146,211,861,883]" captionTargetBox="[146,1101,165,824]" captionTargetId="figure-189@6.[146,1104,165,824]" captionTargetPageId="6" captionText="Figure 3. Dichogaster (Dt.) gasanai sp.nov. (A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores." figureDoi="http://doi.org/10.5281/zenodo.5223736" httpUri="https://zenodo.org/record/5223736/files/figure.png" pageId="7" pageNumber="2233">Figure 3B,C</figureCitation>
). There are two pairs of spermathecae in segment 8 and 9. The ampoule is elongated dumb-bell-shaped and on its basis bears a small rounded diverticulum. The duct is narrow and half as long as the ampoule (
<figureCitation id="13102A32F317FFA969A5FCCAFC8FCD37" box="[691,809,833,857]" captionStart="Figure 3" captionStartId="6.[146,211,861,883]" captionTargetBox="[146,1101,165,824]" captionTargetId="figure-189@6.[146,1104,165,824]" captionTargetPageId="6" captionText="Figure 3. Dichogaster (Dt.) gasanai sp.nov. (A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores." figureDoi="http://doi.org/10.5281/zenodo.5223736" httpUri="https://zenodo.org/record/5223736/files/figure.png" pageId="7" pageNumber="2233">Figure 3D</figureCitation>
).
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3FC01FE9ECDCC" blockId="7.[213,312,906,930]" box="[213,312,906,930]" pageId="7" pageNumber="2233">
<emphasis id="B95FEAA5F317FFA96BC3FC01FE9ECDCC" box="[213,312,906,930]" italics="true" pageId="7" pageNumber="2233">Remarks</emphasis>
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3FC38FBE1CA42" blockId="7.[213,1169,947,1068]" pageId="7" pageNumber="2233">
This species belongs to the
<emphasis id="B95FEAA5F317FFA96912FC38FDEFCDA5" box="[516,585,947,971]" italics="true" pageId="7" pageNumber="2233">affinis</emphasis>
species group and, by the penial setae, resembles
<taxonomicName id="4C2B4D34F317FFA96F63FC38FDE6CD85" authority="Michaelsen, 1912" authorityName="Michaelsen" authorityYear="1912" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="2233" phylum="Annelida" rank="species" species="crateris" subGenus="Diplothecodrilus">
<emphasis id="B95FEAA5F317FFA96F63FC38FEC9CD85" italics="true" pageId="7" pageNumber="2233">D. (Dt.) crateris</emphasis>
<bibRefCitation id="EFBA4B46F317FFA96A61FC58FDE6CD85" author="Michaelsen W" box="[375,576,979,1003]" pageId="7" pageNumber="2233" pagination="1 - 90" refId="ref6109" refString="Michaelsen W. 1912. Die terrestrischen Oligochaten des tropischen Afrikas und ihre geographischen Beziehungen. Wissensch Ergebn Dtsch Zentral-Afrika Exped. 3: 1 - 90." type="journal article" year="1912">Michaelsen, 1912</bibRefCitation>
</taxonomicName>
. Nevertheless it differs from the latter (and from all other member of the
<emphasis id="B95FEAA5F317FFA96AD0FC7FFDADCA62" box="[454,523,1012,1036]" italics="true" pageId="7" pageNumber="2233">affinis</emphasis>
group) in the more backward location of the last pair of hearts, in the position of the clitellum and in the shape of the spermathecae.
</paragraph>
</subSubSection>
</treatment>
</document>