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<document id="CE7FCA186AE743119DE69D8724BABE2E" ID-DOI="10.11646/zootaxa.5428.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10847364" ID-ZooBank="E438D8DA-B142-4A0C-BA25-F6B6EC45C197" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1711009717928" checkinUser="plazi" docAuthor="Almeida-Silva, Lina M. &amp; Brescovit, Antonio D." docDate="2024" docId="B86D8799FFA1FFC28CDDFA47FDC877E1" docLanguage="en" docName="zootaxa.5428.2.1.pdf" docOrigin="Zootaxa 5428 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5428.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goeldia utcuyacu Almeida-Silva &amp; Brescovit 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="187" masterDocId="4454FFE1FF83FFE68C4AFFDCFFC37030" masterDocTitle="Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae)" masterLastPageNumber="193" masterPageNumber="151" pageNumber="185" updateTime="1711010311489" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="39F4B958C2FEFBBB691A2BF551652E9D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F8321EAAF562EBBD993084144EDB3C93">
<mods:title id="2606FDBC68B9B55AD74BC6A965318DBA">Unraveling the mysteries of Goeldia Keyserling, 1891: revision, description of seven new species and first record from USA (Araneae: Titanoecidae)</mods:title>
</mods:titleInfo>
<mods:name id="94FBC00A3F58EEA765D5182AA67A7CB6" type="personal">
<mods:role id="9BF4691D4DD0CC302B474F90DF37E8E8">
<mods:roleTerm id="54B9022DC72A88DFBFE43A2B264D4AE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="407D1531970B8B52E576832506D2B562">Almeida-Silva, Lina M.</mods:namePart>
<mods:affiliation id="D98946AC683A71E4E05CB89F6867E763">Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil.</mods:affiliation>
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<mods:name id="6C78FCC3E45D401E01887B898CF07A38" type="personal">
<mods:role id="AFC0895BCFE2E151512CA2AEB4DE8269">
<mods:roleTerm id="F4727C554352157DB239FC913D53B08A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="77689A29699EA6EA06F9670A55E31BD7">Brescovit, Antonio D.</mods:namePart>
<mods:affiliation id="4E4E726FD91D77F1D75626CFBDB265D3">Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil. &amp; Laboratório de Coleções Zoológicas-LCZ, Instituto Butantan, Avenida Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil &amp; Laboratório de Genômica Evolutiva, Universidade Estadual de Campinas, Instituto de Biologia, Departamento de Genética, Evolução, Microbiologia e Imunologia, Cidade Universitária “ Zeferino Vaz ” Rua Monteiro Lobato, 255, 13083 - 970 - Campinas, SP, Brazil.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="A840B923D34591F3B9A9C4505232D36B">text</mods:typeOfResource>
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<mods:titleInfo id="994689B62A79A4E1B4B3CFD8EEEDF686">
<mods:title id="71B7CE3E7E411C49438233AE2B1C2D6B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B4B99223C430D25C9831980099D95892">
<mods:date id="3888BDC8E7EC835DB595B7A4F415D654">2024</mods:date>
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<mods:number id="B91AA60C26B597D4060B2E32B46A0E93">2024-03-21</mods:number>
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<mods:number id="14E0E8004E9B0E25ADE0364032D822BE">5428</mods:number>
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<mods:number id="1EB35A3FD028462AD6EE0794669A053F">2</mods:number>
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<mods:start id="0E8B134F83FC98FDA5CC0286B3C52B2D">151</mods:start>
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<mods:url id="B6BE8CD5374ABAB45433F81D9960924A">http://dx.doi.org/10.11646/zootaxa.5428.2.1</mods:url>
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<mods:classification id="C366FBD3214C97A11150AD498F6BA3B3">journal article</mods:classification>
<mods:identifier id="EF4A157C8EFB0AE479EEE61BA540C84A" type="DOI">10.11646/zootaxa.5428.2.1</mods:identifier>
<mods:identifier id="CDD7DDCE0D7A2A3C6BA4E5311E28CC9C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F8BE5E0E0CC62999A520F53459EE7925" type="Zenodo-Dep">10847364</mods:identifier>
<mods:identifier id="4BC0330DFEEC5BB92BCC6BF361CD1AD3" type="ZooBank">E438D8DA-B142-4A0C-BA25-F6B6EC45C197</mods:identifier>
</mods:mods>
<treatment id="B86D8799FFA1FFC28CDDFA47FDC877E1" ID-DOI="http://doi.org/10.5281/zenodo.10847439" ID-Zenodo-Dep="10847439" LSID="urn:lsid:plazi:treatment:B86D8799FFA1FFC28CDDFA47FDC877E1" httpUri="http://treatment.plazi.org/id/B86D8799FFA1FFC28CDDFA47FDC877E1" lastPageId="36" lastPageNumber="187" pageId="34" pageNumber="185">
<subSubSection id="78DE6504FFA1FFC48CDDFA47FE127586" box="[151,465,1435,1462]" pageId="34" pageNumber="185" type="nomenclature">
<paragraph id="307B368FFFA1FFC48CDDFA47FE127586" blockId="34.[151,465,1435,1498]" box="[151,465,1435,1462]" pageId="34" pageNumber="185">
<heading id="6B3381E3FFA1FFC48CDDFA47FE127586" bold="true" box="[151,465,1435,1462]" fontSize="11" level="1" pageId="34" pageNumber="185" reason="1">
<emphasis id="02B0EA9DFFA1FFC48CDDFA47FE127586" bold="true" box="[151,465,1435,1462]" pageId="34" pageNumber="185">
<taxonomicName id="F7C44D0CFFA1FFC48CDDFA47FEAF7586" authority="Almeida-Silva &amp; Brescovit, 2024" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[151,364,1435,1462]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="utcuyacu" status="sp. nov.">
<emphasis id="02B0EA9DFFA1FFC48CDDFA47FEAF7586" bold="true" box="[151,364,1435,1462]" italics="true" pageId="34" pageNumber="185">Goeldia utcuyacu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="198357E6FFA1FFC48D39FA40FE127586" box="[371,465,1436,1462]" pageId="34" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FFA1FFC48CDDFA63FE9175EA" box="[151,338,1471,1498]" pageId="34" pageNumber="185" type="description">
<paragraph id="307B368FFFA1FFC48CDDFA63FE9175EA" blockId="34.[151,465,1435,1498]" box="[151,338,1471,1498]" pageId="34" pageNumber="185">
(
<figureCitation id="A8FF2A0AFFA1FFC48CD5FA63FF3475EA" box="[159,247,1471,1498]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="34" pageNumber="185">Figs 21</figureCitation>
;
<figureCitation id="A8FF2A0AFFA1FFC48D49FA1CFEE175EA" box="[259,290,1472,1498]" captionStart="FIGURE 22" captionStartId="36.[151,250,1291,1315]" captionTargetBox="[151,1437,181,1267]" captionTargetId="figure-21@36.[151,1437,181,1267]" captionTargetPageId="36" captionText="FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view." figureDoi="http://doi.org/10.5281/zenodo.10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="34" pageNumber="185">22</figureCitation>
;
<figureCitation id="A8FF2A0AFFA1FFC48D67FA63FE8975EA" box="[301,330,1471,1498]" captionStart="FIGURE 25" captionStartId="39.[151,250,1041,1065]" captionTargetBox="[184,1404,188,1011]" captionTargetId="figure-21@39.[177,1411,181,1018]" captionTargetPageId="39" captionText="FIGURE 25. Distribution of Goedia spp. in northern South America: Goeldia diva (pentagon—pink); G. obscura (square— yellow); G. guayaquilensis (circle—purple); G. mirim (star—red); G.utcuyacu (rectangle—blue). Type species are marked as black versions of each symbol when there are records to multiple localities." figureDoi="http://doi.org/10.5281/zenodo.10847445" httpUri="https://zenodo.org/record/10847445/files/figure.png" pageId="34" pageNumber="185">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FFA1FFC48CDDF9D4FA927772" pageId="34" pageNumber="185" type="materials_examined">
<paragraph id="307B368FFFA1FFC48CDDF9D4FB957676" blockId="34.[151,1437,1543,2038]" pageId="34" pageNumber="185">
<emphasis id="02B0EA9DFFA1FFC48CDDF9D4FEF87611" bold="true" box="[151,315,1543,1570]" pageId="34" pageNumber="185">Type material</emphasis>
.
<materialsCitation id="80AC3CD2FFA1FFC48D0DF9DBFB957676" collectingDate="1984-03" collectionCode="AMNH" collectorName="F. Woytkowski" country="Peru" elevation="1900" latitude="-11.186639" location="Utcuyacu" longLatPrecision="1" longitude="-75.46427" pageId="34" pageNumber="185" specimenCount="1" specimenCount-male="1" stateProvince="Junin" typeStatus="holotype">
<typeStatus id="EF7F882DFFA1FFC48D0DF9DBFE6C7611" box="[327,431,1543,1569]" pageId="34" pageNumber="185" type="holotype">Holotype</typeStatus>
<specimenCount id="26C2FD06FFA1FFC48DFEF9D4FE0A7612" box="[436,457,1544,1570]" count="1" pageId="34" pageNumber="185" type="male">♂</specimenCount>
from
<location id="351B6054FFA1FFC48E43F9D4FDB77612" LSID="urn:lsid:plazi:treatment:B86D8799FFA1FFC28CDDFA47FDC877E1:351B6054FFA1FFC48E43F9D4FDB77612" box="[521,628,1544,1570]" country="Peru" latitude="-11.186639" longLatPrecision="1" longitude="-75.46427" name="Utcuyacu" pageId="34" pageNumber="185" stateProvince="Junin">Utcuyacu</location>
(
<geoCoordinate id="55F05048FFA1FFC48EC8F9D4FCDB7612" box="[642,792,1544,1570]" degrees="11" direction="south" minutes="11" orientation="latitude" pageId="34" pageNumber="185" precision="1" seconds="11.9" value="-11.186639">11°11’11.9”S</geoCoordinate>
<geoCoordinate id="55F05048FFA1FFC48F57F9DBFC067612" box="[797,965,1543,1570]" degrees="75" direction="west" minutes="27" orientation="longitude" pageId="34" pageNumber="185" precision="1" seconds="51.4" value="-75.46427">75°27’51.4”W</geoCoordinate>
),
<collectingRegion id="F200F86DFFA1FFC48F9CF9DBFBD67611" box="[982,1045,1543,1569]" country="Peru" name="Junin" pageId="34" pageNumber="185">Junin</collectingRegion>
,
<collectingCountry id="48D3761FFFA1FFC48855F9D4FB967612" box="[1055,1109,1544,1570]" name="Peru" pageId="34" pageNumber="185">Peru</collectingCountry>
, elev.
<quantity id="F73C9B6AFFA1FFC488DCF9DBFAF27612" box="[1174,1329,1543,1570]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="34" pageNumber="185" unit="m" value="1900.0" valueMax="2200.0" valueMin="1600.0">
<elevation id="BBE9D1BCFFA1FFC488DCF9DBFAF27612" box="[1174,1329,1543,1570]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="34" pageNumber="185" unit="m" value="1900.0" valueMax="2200.0" valueMin="1600.0">1600–2200 m</elevation>
</quantity>
,
<date id="447A104FFFA1FFC48970F9D4FA5B7612" box="[1338,1432,1544,1570]" pageId="34" pageNumber="185" value="1984-03">
<collectingDate id="543EE9A7FFA1FFC48970F9D4FA5B7612" box="[1338,1432,1544,1570]" pageId="34" pageNumber="185" value="1984-03">III.1984</collectingDate>
</date>
,
<collectorName id="9D315359FFA1FFC48CDDF9F0FEFC7675" box="[151,319,1579,1606]" pageId="34" pageNumber="185">F. Woytkowski</collectorName>
coll. (
<collectionCode id="56D5AE4AFFA1FFC48DCCF9F0FE217676" box="[390,482,1580,1606]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="34" pageNumber="185">AMNH</collectionCode>
);
<typeStatus id="EF7F882DFFA1FFC48DBFF9F0FD967676" box="[501,597,1580,1606]" pageId="34" pageNumber="185" type="paratype">paratype</typeStatus>
female with same data as holotype (
<collectionCode id="56D5AE4AFFA1FFC48FA4F9F0FB897676" box="[1006,1098,1580,1606]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="34" pageNumber="185">AMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="307B368FFFA1FFC48C8DF993FA927772" blockId="34.[151,1437,1543,2038]" pageId="34" pageNumber="185">
<emphasis id="02B0EA9DFFA1FFC48C8DF993FDFF765A" bold="true" box="[199,572,1615,1642]" pageId="34" pageNumber="185">Additional Material Examined.</emphasis>
<materialsCitation id="80AC3CD2FFA1FFC48E09F98CFCDC76BE" collectingDate="1947-04-06" collectingDateMax="1947-04-11" collectingDateMin="1947-04-06" collectionCode="AMNH" collectorName="J. C. Pallister" country="Peru" elevation="3670" latitude="-13.629167" location="Hacienda Capana" longLatPrecision="21" longitude="-71.38611" municipality="Ocongate" pageId="34" pageNumber="185" specimenCount="2" specimenCount-female="2" stateProvince="Cusco">
<collectingCountry id="48D3761FFFA1FFC48E09F98CFD4D765A" box="[579,654,1616,1642]" name="Peru" pageId="34" pageNumber="185">PERU</collectingCountry>
.
<emphasis id="02B0EA9DFFA1FFC48EDDF98CFD26765A" box="[663,741,1616,1642]" italics="true" pageId="34" pageNumber="185">
<collectingRegion id="F200F86DFFA1FFC48EDDF98CFD23765A" box="[663,736,1616,1642]" country="Peru" name="Cusco" pageId="34" pageNumber="185">Cuzco</collectingRegion>
.
</emphasis>
<collectingMunicipality id="D01FACF5FFA1FFC48EA1F98CFC95765A" box="[747,854,1616,1642]" pageId="34" pageNumber="185">Ocongate</collectingMunicipality>
(
<geoCoordinate id="55F05048FFA1FFC48F2FF993FC29765A" box="[869,1002,1615,1642]" degrees="13" direction="south" minutes="37" orientation="latitude" pageId="34" pageNumber="185" precision="15" seconds="45" value="-13.629167">13°37’45”S</geoCoordinate>
<geoCoordinate id="55F05048FFA1FFC48FBBF98CFB40765A" box="[1009,1155,1616,1642]" degrees="71" direction="west" minutes="23" orientation="longitude" pageId="34" pageNumber="185" precision="15" seconds="10" value="-71.38611">71°23’10”W</geoCoordinate>
),
<location id="351B6054FFA1FFC488DDF993FA9D765A" LSID="urn:lsid:plazi:treatment:B86D8799FFA1FFC28CDDFA47FDC877E1:351B6054FFA1FFC488DDF993FA9D765A" box="[1175,1374,1615,1642]" country="Peru" latitude="-13.629167" longLatPrecision="21" longitude="-71.38611" municipality="Ocongate" name="Hacienda Capana" pageId="34" pageNumber="185" stateProvince="Cusco">Hacienda Capana</location>
, elev.
<quantity id="F73C9B6AFFA1FFC48CDDF9AFFF3376BE" box="[151,240,1651,1678]" metricMagnitude="3" metricUnit="m" metricValue="3.67" pageId="34" pageNumber="185" unit="m" value="3670.0">
<elevation id="BBE9D1BCFFA1FFC48CDDF9AFFF3376BE" box="[151,240,1651,1678]" metricMagnitude="3" metricUnit="m" metricValue="3.67" pageId="34" pageNumber="185" unit="m" value="3670.0">3670 m</elevation>
</quantity>
,
<specimenCount id="26C2FD06FFA1FFC48CB4F9A8FEE176BE" box="[254,290,1652,1678]" count="2" pageId="34" pageNumber="185" type="female">2♀</specimenCount>
,
<date id="447A104FFFA1FFC48D7BF9AFFE0976BD" box="[305,458,1651,1677]" pageId="34" pageNumber="185" value="1947-04-06" valueMax="1947-04-11" valueMin="1947-04-06">
<collectingDate id="543EE9A7FFA1FFC48D7BF9AFFE0976BD" box="[305,458,1651,1677]" pageId="34" pageNumber="185" value="1947-04-06" valueMax="1947-04-11" valueMin="1947-04-06">6–11.IV.1947</collectingDate>
</date>
,
<collectorName id="9D315359FFA1FFC48D92F9A8FDB276BD" box="[472,625,1651,1678]" pageId="34" pageNumber="185">J. C. Pallister</collectorName>
coll. (
<collectionCode id="56D5AE4AFFA1FFC48EF1F9A8FCD476BE" box="[699,791,1652,1678]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="34" pageNumber="185">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="80AC3CD2FFA1FFC48F66F9AFFC417682" collectingDate="1964-08-01" collectingDateMax="1964-08-05" collectingDateMin="1964-08-01" collectionCode="AMNH" collectorName="B. Malkin" country="Peru" elevation="2700" latitude="-13.163055" location="Machu-Picchu" longLatPrecision="21" longitude="-72.545555" municipality="Urubamba" pageId="34" pageNumber="185" specimenCount="1" specimenCount-female="1" stateProvince="Cuzco">
<collectingMunicipality id="D01FACF5FFA1FFC48F66F9AFFC6676BD" box="[812,933,1651,1677]" pageId="34" pageNumber="185">Urubamba</collectingMunicipality>
,
<location id="351B6054FFA1FFC48FF9F9AFFB9A76BD" LSID="urn:lsid:plazi:treatment:B86D8799FFA1FFC28CDDFA47FDC877E1:351B6054FFA1FFC48FF9F9AFFB9A76BD" box="[947,1113,1651,1677]" country="Peru" latitude="-13.163055" longLatPrecision="21" longitude="-72.545555" municipality="Urubamba" name="Machu-Picchu" pageId="34" pageNumber="185" stateProvince="Cuzco">Machu-Picchu</location>
, (
<geoCoordinate id="55F05048FFA1FFC48824F9A8FB3076BE" box="[1134,1267,1652,1678]" degrees="13" direction="south" minutes="09" orientation="latitude" pageId="34" pageNumber="185" precision="15" seconds="47" value="-13.163055">13°09’47”S</geoCoordinate>
<geoCoordinate id="55F05048FFA1FFC488B7F9A8FA4C76BE" box="[1277,1423,1652,1678]" degrees="72" direction="west" minutes="32" orientation="longitude" pageId="34" pageNumber="185" precision="15" seconds="44" value="-72.545555">72°32’44”W</geoCoordinate>
), elev.
<quantity id="F73C9B6AFFA1FFC48C99F94BFEB17682" box="[211,370,1687,1714]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="34" pageNumber="185" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">
<elevation id="BBE9D1BCFFA1FFC48C99F94BFEB17682" box="[211,370,1687,1714]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="34" pageNumber="185" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">2600–2800 m</elevation>
</quantity>
,
<specimenCount id="26C2FD06FFA1FFC48D35F944FE617682" box="[383,418,1688,1714]" count="1" pageId="34" pageNumber="185" type="female">1♀</specimenCount>
,
<date id="447A104FFFA1FFC48DFBF94BFD977682" box="[433,596,1687,1714]" pageId="34" pageNumber="185" value="1964-08-01" valueMax="1964-08-05" valueMin="1964-08-01">
<collectingDate id="543EE9A7FFA1FFC48DFBF94BFD977682" box="[433,596,1687,1714]" pageId="34" pageNumber="185" value="1964-08-01" valueMax="1964-08-05" valueMin="1964-08-01">1–5.VIII.1964</collectingDate>
</date>
,
<collectorName id="9D315359FFA1FFC48E2BF944FD177681" box="[609,724,1687,1714]" pageId="34" pageNumber="185">B. Malkin</collectorName>
coll. (
<collectionCode id="56D5AE4AFFA1FFC48F54F944FCB97682" box="[798,890,1688,1714]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="34" pageNumber="185">AMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="80AC3CD2FFA1FFC48FC7F944FD1176E6" collectingDate="1965-07-07" collectingDateMax="1965-07-20" collectingDateMin="1965-07-07" collectionCode="AMNH" country="Peru" latitude="-11.1247225" location="Estancia Naranjal" longLatPrecision="21" longitude="-75.35694" municipality="San Ramon" pageId="34" pageNumber="185" specimenCount="2" specimenCount-female="2" stateProvince="Junin">
<emphasis id="02B0EA9DFFA1FFC48FC7F944FC087681" box="[909,971,1688,1713]" italics="true" pageId="34" pageNumber="185">
<collectingRegion id="F200F86DFFA1FFC48FC7F944FC087681" box="[909,971,1688,1713]" country="Peru" name="Junin" pageId="34" pageNumber="185">Junin</collectingRegion>
</emphasis>
:
<collectingMunicipality id="D01FACF5FFA1FFC48F96F944FB9C7682" box="[988,1119,1688,1714]" pageId="34" pageNumber="185">San Ramon</collectingMunicipality>
(
<geoCoordinate id="55F05048FFA1FFC4883AF944FB377682" box="[1136,1268,1688,1714]" degrees="11" direction="south" minutes="07" orientation="latitude" pageId="34" pageNumber="185" precision="15" seconds="29" value="-11.1247225">11°07’29”S</geoCoordinate>
<geoCoordinate id="55F05048FFA1FFC488B7F94BFA4C7682" box="[1277,1423,1687,1714]" degrees="75" direction="west" minutes="21" orientation="longitude" pageId="34" pageNumber="185" precision="15" seconds="25" value="-75.35694">75°21’25”W</geoCoordinate>
),
<location id="351B6054FFA1FFC48CDDF967FEA676E5" LSID="urn:lsid:plazi:treatment:B86D8799FFA1FFC28CDDFA47FDC877E1:351B6054FFA1FFC48CDDF967FEA676E5" box="[151,357,1723,1749]" country="Peru" latitude="-11.1247225" longLatPrecision="21" longitude="-75.35694" municipality="San Ramon" name="Estancia Naranjal" pageId="34" pageNumber="185" stateProvince="Junin">Estância Naranjal</location>
,
<specimenCount id="26C2FD06FFA1FFC48D3EF960FE5A76E6" box="[372,409,1724,1750]" count="2" pageId="34" pageNumber="185" type="female">2♀</specimenCount>
,
<date id="447A104FFFA1FFC48DE0F960FD9976E6" box="[426,602,1723,1750]" pageId="34" pageNumber="185" value="1965-07-20" valueMax="1965-07-07" valueMin="1965-07-20">
<collectingDate id="543EE9A7FFA1FFC48DE0F960FD9976E6" box="[426,602,1723,1750]" pageId="34" pageNumber="185" value="1965-07-20" valueMax="1965-07-07" valueMin="1965-07-20">20– 7.VII.1965</collectingDate>
</date>
(
<collectionCode id="56D5AE4AFFA1FFC48E24F960FD0976E6" box="[622,714,1724,1750]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="34" pageNumber="185">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="80AC3CD2FFA1FFC48EABF967FDF076CA" collectingDate="1965-07-27" collectingDateMax="1965-07-30" collectingDateMin="1965-07-27" collectionCode="AMNH" country="Peru" elevation="1800" latitude="-11.233055" location="Huacapistana" longLatPrecision="21" longitude="-75.483055" pageId="34" pageNumber="185" specimenCount="4" specimenCount-female="2" specimenCount-male="1" stateProvince="Junin">
<location id="351B6054FFA1FFC48EABF967FCBB76E5" LSID="urn:lsid:plazi:treatment:B86D8799FFA1FFC28CDDFA47FDC877E1:351B6054FFA1FFC48EABF967FCBB76E5" box="[737,888,1723,1749]" country="Peru" latitude="-11.233055" longLatPrecision="21" longitude="-75.483055" name="Huacapistana" pageId="34" pageNumber="185" stateProvince="Junin">Huacapistana</location>
(
<geoCoordinate id="55F05048FFA1FFC48FC7F967FBD276E6" box="[909,1041,1723,1750]" degrees="11" direction="south" minutes="13" orientation="latitude" pageId="34" pageNumber="185" precision="15" seconds="59" value="-11.233055">11°13’59”S</geoCoordinate>
<geoCoordinate id="55F05048FFA1FFC48856F967FB6C76E6" box="[1052,1199,1723,1750]" degrees="75" direction="west" minutes="28" orientation="longitude" pageId="34" pageNumber="185" precision="15" seconds="59" value="-75.483055">75°28’59”W</geoCoordinate>
), elev.
<quantity id="F73C9B6AFFA1FFC4894CF960FAA176E6" box="[1286,1378,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="185" unit="m" value="1800.0">
<elevation id="BBE9D1BCFFA1FFC4894CF960FAA176E6" box="[1286,1378,1724,1750]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="185" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<specimenCount id="26C2FD06FFA1FFC48938F960FA5476E6" box="[1394,1431,1724,1750]" count="1" pageId="34" pageNumber="185" type="male">1♂</specimenCount>
,
<specimenCount id="26C2FD06FFA1FFC48CDDF93CFF7976CA" box="[151,186,1760,1786]" count="2" pageId="34" pageNumber="185" type="female">2♀</specimenCount>
<specimenCount id="26C2FD06FFA1FFC48C8FF903FF3D76C9" box="[197,254,1759,1785]" count="1" pageId="34" pageNumber="185" type="generic">1juv.</specimenCount>
,
<date id="447A104FFFA1FFC48D47F903FE0376CA" box="[269,448,1759,1786]" pageId="34" pageNumber="185" value="1965-07-27" valueMax="1965-07-30" valueMin="1965-07-27">
<collectingDate id="543EE9A7FFA1FFC48D47F903FE0376CA" box="[269,448,1759,1786]" pageId="34" pageNumber="185" value="1965-07-27" valueMax="1965-07-30" valueMin="1965-07-27">27–30.VII.1965</collectingDate>
</date>
(
<collectionCode id="56D5AE4AFFA1FFC48D98F93CFDE876CA" box="[466,555,1760,1786]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="34" pageNumber="185">AMNH</collectionCode>
)
</materialsCitation>
<materialsCitation id="80AC3CD2FFA1FFC48E77F903FC6C772E" collectingDate="1948-03" collectionCode="AMNH" collectorName="P. &amp; B. Wygodzinsky &amp; F. Woytkowski" country="Peru" latitude="-11.186111" location="San Jose de Utcuyacu" longLatPrecision="21" longitude="-75.46305" pageId="34" pageNumber="185" specimenCount="8" specimenCount-female="3" specimenCount-male="1" stateProvince="Junin">
all collected by
<collectorName id="9D315359FFA1FFC48EB3F93CFC3A76C9" box="[761,1017,1759,1786]" pageId="34" pageNumber="185">P. &amp; B. Wygodzinsky</collectorName>
;
<location id="351B6054FFA1FFC4884DF93CFAC576CA" LSID="urn:lsid:plazi:treatment:B86D8799FFA1FFC28CDDFA47FDC877E1:351B6054FFA1FFC4884DF93CFAC576CA" box="[1031,1286,1759,1786]" country="Peru" latitude="-11.186111" longLatPrecision="21" longitude="-75.46305" name="San Jose de Utcuyacu" pageId="34" pageNumber="185" stateProvince="Junin">San José de Utcuyacu</location>
(
<geoCoordinate id="55F05048FFA1FFC48950F93CFA5F76CA" box="[1306,1436,1760,1786]" degrees="11" direction="south" minutes="11" orientation="latitude" pageId="34" pageNumber="185" precision="15" seconds="10" value="-11.186111">11°11’10”S</geoCoordinate>
<geoCoordinate id="55F05048FFA1FFC48CDDF8DFFEE9772E" box="[151,298,1795,1822]" degrees="75" direction="west" minutes="27" orientation="longitude" pageId="34" pageNumber="185" precision="15" seconds="47" value="-75.46305">75°27’47”W</geoCoordinate>
),
<specimenCount id="26C2FD06FFA1FFC48D75F8D8FEA0772E" box="[319,355,1796,1822]" count="3" pageId="34" pageNumber="185" type="female">3♀</specimenCount>
,
<specimenCount id="26C2FD06FFA1FFC48D3BF8D8FE56772E" box="[369,405,1796,1822]" count="1" pageId="34" pageNumber="185" type="male">1♂</specimenCount>
,
<specimenCount id="26C2FD06FFA1FFC48DE8F8D8FE27772D" box="[418,484,1795,1822]" count="4" pageId="34" pageNumber="185" type="generic">4 juv.</specimenCount>
,
<date id="447A104FFFA1FFC48DBAF8D8FD8E772E" box="[496,589,1796,1822]" pageId="34" pageNumber="185" value="1948-03">
<collectingDate id="543EE9A7FFA1FFC48DBAF8D8FD8E772E" box="[496,589,1796,1822]" pageId="34" pageNumber="185" value="1948-03">III.1948</collectingDate>
</date>
,
<collectorName id="9D315359FFA1FFC48E10F8D8FCC1772D" box="[602,770,1795,1822]" pageId="34" pageNumber="185">F. Woytkowski</collectorName>
coll. (
<collectionCode id="56D5AE4AFFA1FFC48F01F8D8FC64772E" box="[843,935,1796,1822]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="34" pageNumber="185">AMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="80AC3CD2FFA1FFC48FF1F8DFFA8E7772" collectingDate="1955-01-03" collectionCode="CAS" collectorName="E. Schlinger &amp; E. Ross" country="Peru" location="Rio Perene" municipality="Colonia Perene" pageId="34" pageNumber="185" specimenCode="CAS 9021317" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Junin">
<collectingMunicipality id="D01FACF5FFA1FFC48FF1F8DFFBA9772E" box="[955,1130,1795,1822]" pageId="34" pageNumber="185">Colonia Perene</collectingMunicipality>
,
<location id="351B6054FFA1FFC4883CF8DFFB36772E" LSID="urn:lsid:plazi:treatment:B86D8799FFA1FFC28CDDFA47FDC877E1:351B6054FFA1FFC4883CF8DFFB36772E" box="[1142,1269,1795,1822]" country="Peru" municipality="Colonia Perene" name="Rio Perene" pageId="34" pageNumber="185" stateProvince="Junin">Rio Perene</location>
,
<locationDeviation id="21C92FADFFA1FFC4894BF8D8FF2C7771" location="La Merced" pageId="34" pageNumber="185">
<quantity id="F73C9B6AFFA1FFC4894BF8D8FA89772D" box="[1281,1354,1795,1822]" metricMagnitude="4" metricUnit="m" metricValue="2.8968192" pageId="34" pageNumber="185" unit="mi" value="18.0">18 mi.</quantity>
NE La Merced
</locationDeviation>
,
<specimenCount id="26C2FD06FFA1FFC48CB1F8F4FEDE7772" box="[251,285,1832,1858]" count="2" pageId="34" pageNumber="185" type="male">2♂</specimenCount>
<specimenCount id="26C2FD06FFA1FFC48D6FF8F4FE8A7772" box="[293,329,1832,1858]" count="2" pageId="34" pageNumber="185" type="female">2♀</specimenCount>
,
<date id="447A104FFFA1FFC48D1CF8FBFE767772" box="[342,437,1831,1858]" pageId="34" pageNumber="185" value="1955-01-03">
<collectingDate id="543EE9A7FFA1FFC48D1CF8FBFE767772" box="[342,437,1831,1858]" pageId="34" pageNumber="185" value="1955-01-03">3.I.1955</collectingDate>
</date>
,
<collectorName id="9D315359FFA1FFC48D8BF8F4FD887771" box="[449,587,1831,1858]" pageId="34" pageNumber="185">E. Schlinger</collectorName>
&amp;
<collectorName id="9D315359FFA1FFC48E25F8F4FD077772" box="[623,708,1832,1858]" pageId="34" pageNumber="185">E. Ross</collectorName>
coll. (
<specimenCode id="60629EF4FFA1FFC48F46F8F4FC737772" box="[780,944,1832,1858]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="34" pageNumber="185" type="Museum">CAS 9021317</specimenCode>
, SEM, male palp and female leg IV)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FFA1FFC48C8DF890FAB97756" box="[199,1402,1867,1894]" pageId="34" pageNumber="185" type="etymology">
<paragraph id="307B368FFFA1FFC48C8DF890FAB97756" blockId="34.[151,1437,1543,2038]" box="[199,1402,1867,1894]" pageId="34" pageNumber="185">
<emphasis id="02B0EA9DFFA1FFC48C8DF890FE8B7756" bold="true" box="[199,328,1868,1894]" pageId="34" pageNumber="185">Etymology</emphasis>
. The specific epithet is a noun in apposition from the
<typeStatus id="EF7F882DFFA1FFC48FEFF890FC167756" box="[933,981,1868,1894]" pageId="34" pageNumber="185">type</typeStatus>
locality, San José de Utcuyacu,
<collectingCountry id="48D3761FFFA1FFC48975F890FAB57756" box="[1343,1398,1868,1894]" name="Peru" pageId="34" pageNumber="185">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FFA1FFC48C8DF8B3FA9C77C6" pageId="34" pageNumber="185" type="diagnosis">
<paragraph id="307B368FFFA1FFC48C8DF8B3FA9C77C6" blockId="34.[151,1437,1543,2038]" pageId="34" pageNumber="185">
<emphasis id="02B0EA9DFFA1FFC48C8DF8B3FEF977B9" bold="true" box="[199,314,1903,1929]" pageId="34" pageNumber="185">Diagnosis</emphasis>
.
<taxonomicName id="F7C44D0CFFA1FFC48D00F8B3FDD177B9" authority="Almeida-Silva &amp; Brescovit, 2024" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[330,530,1903,1929]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="185" phylum="Arthropoda" rank="species" species="utcuyacu" status="sp. nov.">
<emphasis id="02B0EA9DFFA1FFC48D00F8B3FDD177B9" box="[330,530,1903,1929]" italics="true" pageId="34" pageNumber="185">Goeldia utcuyacu</emphasis>
</taxonomicName>
<emphasis id="02B0EA9DFFA1FFC48E51F8ACFDB577BA" bold="true" box="[539,630,1904,1930]" pageId="34" pageNumber="185">
<taxonomicNameLabel id="198357E6FFA1FFC48E51F8ACFDB577BA" box="[539,630,1904,1930]" pageId="34" pageNumber="185" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other species of the genus by the thick patellar apophysis with sinuous tip (
<figureCitation id="A8FF2A0AFFA1FFC48D11F84FFE2A779E" box="[347,489,1939,1966]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="34" pageNumber="185">Figs 21B–D</figureCitation>
;
<figureCitation id="A8FF2A0AFFA1FFC48DBEF848FDE5779E" box="[500,550,1940,1966]" captionStart="FIGURE 22" captionStartId="36.[151,250,1291,1315]" captionTargetBox="[151,1437,181,1267]" captionTargetId="figure-21@36.[151,1437,181,1267]" captionTargetPageId="36" captionText="FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view." figureDoi="http://doi.org/10.5281/zenodo.10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="34" pageNumber="185">22E</figureCitation>
); and by the tegular process subrectangular and with enlarged base in the male palp (
<figureCitation id="A8FF2A0AFFA1FFC48C92F86BFE8477E2" box="[216,327,1975,2002]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="34" pageNumber="185">Figs 21A</figureCitation>
;
<figureCitation id="A8FF2A0AFFA1FFC48D1EF864FE4A77E2" box="[340,393,1976,2002]" captionStart="FIGURE 22" captionStartId="36.[151,250,1291,1315]" captionTargetBox="[151,1437,181,1267]" captionTargetId="figure-21@36.[151,1437,181,1267]" captionTargetPageId="36" captionText="FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view." figureDoi="http://doi.org/10.5281/zenodo.10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="34" pageNumber="185">22A</figureCitation>
). Females can be recognized by the epigynum with median field wide, with posterior edge almost straight (
<figureCitation id="A8FF2A0AFFA1FFC48D06F807FE7377C6" box="[332,432,2011,2038]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="34" pageNumber="185">Fig. 21E</figureCitation>
), and the vertical spermathecae slender and longer than copulatory ducts (
<figureCitation id="A8FF2A0AFFA1FFC488A4F807FA9177C6" box="[1262,1362,2011,2038]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="34" pageNumber="185">Fig. 21F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="64BB6607FFA0FFC58CDDFA19FABA7631" ID-DOI="http://doi.org/10.5281/zenodo.10847431" ID-Zenodo-Dep="10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="35" pageNumber="186" startId="35.[151,250,1477,1501]" targetBox="[159,1429,187,1448]" targetPageId="35" targetType="figure">
<paragraph id="307B368FFFA0FFC58CDDFA19FABA7631" blockId="35.[151,1436,1477,1537]" pageId="35" pageNumber="186">
<emphasis id="02B0EA9DFFA0FFC58CDDFA19FEE375ED" bold="true" box="[151,288,1477,1501]" pageId="35" pageNumber="186">FIGURE 21.</emphasis>
A–F.
<taxonomicName id="F7C44D0CFFA0FFC58D17FA19FDD375ED" authority="Almeida-Silva &amp; Brescovit, 2024" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[349,528,1477,1501]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="utcuyacu" status="sp. nov.">
<emphasis id="02B0EA9DFFA0FFC58D17FA19FDD375ED" box="[349,528,1477,1501]" italics="true" pageId="35" pageNumber="186">Goeldia utcuyacu</emphasis>
</taxonomicName>
<emphasis id="02B0EA9DFFA0FFC58E5DFA19FDA475ED" bold="true" box="[535,615,1477,1501]" pageId="35" pageNumber="186">
<taxonomicNameLabel id="198357E6FFA0FFC58E5DFA19FDA475ED" box="[535,615,1477,1501]" pageId="35" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view.
</paragraph>
</caption>
<subSubSection id="78DE6504FFA0FFC28C8DF98FFD92779C" lastPageId="36" lastPageNumber="187" pageId="35" pageNumber="186" type="description">
<paragraph id="307B368FFFA0FFC58C8DF98FFE2177C9" blockId="35.[151,1437,1619,2041]" pageId="35" pageNumber="186">
<emphasis id="02B0EA9DFFA0FFC58C8DF98FFE92765D" bold="true" box="[199,337,1619,1645]" pageId="35" pageNumber="186">Description</emphasis>
. Male (
<typeStatus id="EF7F882DFFA0FFC58DE1F98FFDDB765D" box="[427,536,1619,1645]" pageId="35" pageNumber="186" type="holotype">Holotype</typeStatus>
, AMNH). Carapace brown with cephalic area darker; femur I brown; other leg segments yellowish-brown; chelicerae dark brown, labium and endites brown. Total length 4.8. Carapace: length 2.4; width 1.8; height 1.34. Clypeus: height 0.16. Opisthosoma: length 2.43; width 1.4. Eye diameters and interdistances: AME 0.06; ALE 0.11; PME 0.10; PLE 0.12; AME–AME 0.11; AME–ALE 0.11; ALE–PLE 0.5; PME–PLE 0.20; PME–PME 0.13; AME–PME 0.12. Chelicerae paturon: length 1.25. Leg measurements: I: femur 2.2/ patella 1.3/ tibia 2.3/ metatarsus 2.2/ tarsus 1.0/total 9.0; II: 2.0/ 0.8/ 1.9/ 1.85/ 0.95/ 7.5; III: 1.85/ 0.7/ 1.5/ 1.6/ 0.7/ 6.35; IV: 2.35/ 0.7/ 2.2/ 2.0/ 0.85/ 8.1. Leg formula: 1423. Leg spination: femur I–II p1ap; tibia II v2-0-2 or v0-0-2, p1ap, r1ap; III v2sa or v1-0-2, r 0-0-1, p0-1-1; IV: v1-0-2 or v0-1-2, p1ap, r1ap; metatarsus I v1p-2-
<quantity id="F73C9B6AFFA0FFC588A2F893FACF7759" box="[1256,1292,1871,1897]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="35" pageNumber="186" unit="m" value="1.0">1m</quantity>
, p1ap, r1ap; II v2-1r-2(1p-1r)-2(1p-1r)-2ap, p0-2-2, r2ap. Patellar apophysis dorso-posteriorly projected (
<figureCitation id="A8FF2A0AFFA0FFC5888AF8AFFA8C77BD" box="[1216,1359,1907,1933]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="35" pageNumber="186">Figs 21B–D</figureCitation>
,
<figureCitation id="A8FF2A0AFFA0FFC58915F8AFFA5377BD" box="[1375,1424,1907,1933]" captionStart="FIGURE 22" captionStartId="36.[151,250,1291,1315]" captionTargetBox="[151,1437,181,1267]" captionTargetId="figure-21@36.[151,1437,181,1267]" captionTargetPageId="36" captionText="FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view." figureDoi="http://doi.org/10.5281/zenodo.10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="35" pageNumber="186">22E</figureCitation>
). MLT base enlarged and apex partially sclerotized (
<figureCitation id="A8FF2A0AFFA0FFC58E84F84BFCF47781" box="[718,823,1943,1969]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="35" pageNumber="186">Figs 21C</figureCitation>
,
<figureCitation id="A8FF2A0AFFA0FFC58F09F84BFC547781" box="[835,919,1943,1969]" captionStart="FIGURE 22" captionStartId="36.[151,250,1291,1315]" captionTargetBox="[151,1437,181,1267]" captionTargetId="figure-21@36.[151,1437,181,1267]" captionTargetPageId="36" captionText="FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view." figureDoi="http://doi.org/10.5281/zenodo.10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="35" pageNumber="186">22D–E</figureCitation>
). Cymbium with macrosetae (
<figureCitation id="A8FF2A0AFFA0FFC588ACF84BFA937781" box="[1254,1360,1943,1969]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="35" pageNumber="186">Figs 21B</figureCitation>
,
<figureCitation id="A8FF2A0AFFA0FFC58916F84BFA537781" box="[1372,1424,1943,1969]" captionStart="FIGURE 22" captionStartId="36.[151,250,1291,1315]" captionTargetBox="[151,1437,181,1267]" captionTargetId="figure-21@36.[151,1437,181,1267]" captionTargetPageId="36" captionText="FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view." figureDoi="http://doi.org/10.5281/zenodo.10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="35" pageNumber="186">22D</figureCitation>
). Tegular process with uniform width and base membranous (
<figureCitation id="A8FF2A0AFFA0FFC58F1AF867FC0377E5" box="[848,960,1979,2005]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="35" pageNumber="186">Figs 21A</figureCitation>
;
<figureCitation id="A8FF2A0AFFA0FFC58F84F867FBC077E5" box="[974,1027,1979,2005]" captionStart="FIGURE 22" captionStartId="36.[151,250,1291,1315]" captionTargetBox="[151,1437,181,1267]" captionTargetId="figure-21@36.[151,1437,181,1267]" captionTargetPageId="36" captionText="FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view." figureDoi="http://doi.org/10.5281/zenodo.10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="35" pageNumber="186">22A</figureCitation>
).
<emphasis id="02B0EA9DFFA0FFC58853F860FB7177E5" box="[1049,1202,1979,2005]" italics="true" pageId="35" pageNumber="186">Pars pendula</emphasis>
thickening far from embolus tip (
<figureCitation id="A8FF2A0AFFA0FFC58D63F803FE5677C9" box="[297,405,2015,2041]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="35" pageNumber="186">Figs 21D</figureCitation>
;
<figureCitation id="A8FF2A0AFFA0FFC58DEBF803FE1577C9" box="[417,470,2015,2041]" captionStart="FIGURE 22" captionStartId="36.[151,250,1291,1315]" captionTargetBox="[151,1437,181,1267]" captionTargetId="figure-21@36.[151,1437,181,1267]" captionTargetPageId="36" captionText="FIGURE 22. A–E. Goeldia utcuyacu sp. nov., from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view." figureDoi="http://doi.org/10.5281/zenodo.10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="35" pageNumber="186">22A</figureCitation>
).
</paragraph>
<caption id="64BB6607FFA7FFC28CDDFAD7FEF6755C" ID-DOI="http://doi.org/10.5281/zenodo.10847435" ID-Zenodo-Dep="10847435" httpUri="https://zenodo.org/record/10847435/files/figure.png" pageId="36" pageNumber="187" startId="36.[151,250,1291,1315]" targetBox="[151,1437,181,1267]" targetPageId="36" targetType="figure">
<paragraph id="307B368FFFA7FFC28CDDFAD7FEF6755C" blockId="36.[151,1436,1291,1388]" pageId="36" pageNumber="187">
<emphasis id="02B0EA9DFFA7FFC28CDDFAD7FEE07514" bold="true" box="[151,291,1291,1316]" pageId="36" pageNumber="187">FIGURE 22.</emphasis>
A–E.
<taxonomicName id="F7C44D0CFFA7FFC28D23FAD0FDDC7513" authority="Almeida-Silva &amp; Brescovit, 2024" authorityName="Almeida-Silva &amp; Brescovit" authorityYear="2024" box="[361,543,1292,1316]" class="Arachnida" family="Titanoecidae" genus="Goeldia" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="187" phylum="Arthropoda" rank="species" species="utcuyacu" status="sp. nov.">
<emphasis id="02B0EA9DFFA7FFC28D23FAD0FDDC7513" box="[361,543,1292,1316]" italics="true" pageId="36" pageNumber="187">Goeldia utcuyacu</emphasis>
</taxonomicName>
<emphasis id="02B0EA9DFFA7FFC28E63FAD0FDBF7514" bold="true" box="[553,636,1292,1316]" pageId="36" pageNumber="187">
<taxonomicNameLabel id="198357E6FFA7FFC28E63FAD0FDBF7514" box="[553,636,1292,1316]" pageId="36" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, from Junin, Peru, male genitalia. A. Left palp, bulb, apical view; B. Ditto, retrolateral view; C. Left tibia, ventro-prolateral view; D. Left tibia and cymbium, dorsum apical view; E. Left tibia, dorsum prolateral view.
</paragraph>
</caption>
<paragraph id="307B368FFFA7FFC28C8DFA62FE537754" blockId="36.[151,1437,1470,2001]" pageId="36" pageNumber="187">
Female (AMNH). Coloration: carapace, chelicerae, and femora dark brown; other leg segments, labium and endites caramel brown. Total length: 6.8. Carapace: length 2.56; width 1.92, height 1.28. Clypeus: height 0.21. Opisthosoma: length 4.4, width 3.2. Eye diameters and interdistances: AME 0.08; ALE 0.13; PME 0.12; PLE 0.13; AME–AME 0.08; AME–ALE 0.13;ALE–PLE 0.04; PME–PLE 0.21; PME–PME 0.14; AME–PME 0.09. Chelicerae paturon: length 1.29. Leg measurements: I: femur 1.85/ patella 0.85/ tibia 1.8/ metatarsus 1.55/ tarsus 0.8/ total 6.85; II: 1.8/ 0.9/ 1.5/ 1.4/ 0.7/ 6.3; III: 1.75/ 0.75/ 1.25/ 1.2/ 0.55/ 5.5; IV: 1.8/ 0.8/ 1.65/ 1.55/ 0.55/ 6.35. Leg formula 1423. Leg spination: femur I–II p1ap; tibia III p1ap; metatarsus I–II v2-2-
<quantity id="F73C9B6AFFA7FFC28F87F94AFC337680" box="[973,1008,1686,1712]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="187" unit="m" value="1.0">1m</quantity>
, p 1ap, r1ap; III v1r-1p-2ap, p1-0-1ap, r0-0-1p; IV v1p-1p-
<quantity id="F73C9B6AFFA7FFC28D39F966FE5876E4" box="[371,411,1722,1748]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="187" unit="m" value="2.0">2m</quantity>
, p1ap, r1ap. Median field of epigynum wide but not closing copulatory openings (
<figureCitation id="A8FF2A0AFFA7FFC28961F966FA5376E4" box="[1323,1424,1722,1748]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="36" pageNumber="187">Fig. 21E</figureCitation>
). Copulatory duct wall visible by transparency on the median field and partially visible at the copulatory opening area (
<figureCitation id="A8FF2A0AFFA7FFC28C9FF8DEFEF8772C" box="[213,315,1794,1820]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="36" pageNumber="187">Fig. 21E</figureCitation>
). Copulatory duct base enlarged (
<figureCitation id="A8FF2A0AFFA7FFC28EFEF8DEFCFB772C" box="[692,824,1794,1820]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="36" pageNumber="187">Fig. 21E–F</figureCitation>
). Vertical spermathecae straight (
<figureCitation id="A8FF2A0AFFA7FFC288E7F8DEFAD2772C" box="[1197,1297,1794,1820]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="36" pageNumber="187">Fig. 21F</figureCitation>
). Horizontal spermathecae straight shorter than vertical spermathecae (
<figureCitation id="A8FF2A0AFFA7FFC28F6DF8FAFC4F7770" box="[807,908,1830,1856]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="36" pageNumber="187">Fig. 21F</figureCitation>
). Fertilization ducts enlarged and separated by their length (
<figureCitation id="A8FF2A0AFFA7FFC28D6DF896FE4B7754" box="[295,392,1866,1892]" captionStart="FIGURE 21" captionStartId="35.[151,250,1477,1501]" captionTargetBox="[159,1429,187,1448]" captionTargetId="figure-21@35.[151,1436,181,1453]" captionTargetPageId="35" captionText="FIGURE 21. A–F. Goeldia utcuyacu sp. nov. A–D. Male left palp; E–F Female epigynum: A. Tegular process frontal view. B. Palp ventral view; C. Tibia and patella, dorsal view; D. Retrolateral view. E. Epigynum ventral view; F. Vulva, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10847431" httpUri="https://zenodo.org/record/10847431/files/figure.png" pageId="36" pageNumber="187">Fig. 21F</figureCitation>
)
</paragraph>
<paragraph id="307B368FFFA7FFC28C8DF8B2FD92779C" blockId="36.[151,1437,1470,2001]" pageId="36" pageNumber="187">
<emphasis id="02B0EA9DFFA7FFC28C8DF8B2FEFB77B8" bold="true" box="[199,312,1902,1928]" pageId="36" pageNumber="187">Variation</emphasis>
. Length (
<specimenCount id="26C2FD06FFA7FFC28DE6F8B2FE1E77B8" box="[428,477,1902,1928]" count="2" pageId="36" pageNumber="187" type="male">2 ♂</specimenCount>
): total 4.20–4.80; carapace 2.00–2.40; femur I 2.10–2.20; (
<specimenCount id="26C2FD06FFA7FFC288DAF8B2FB0F77B8" box="[1168,1228,1902,1928]" count="10" pageId="36" pageNumber="187" type="female">10 ♀</specimenCount>
): total 5.20–7.10; carapace 2.14–2.60; femur I 1.78–1.88.
</paragraph>
</subSubSection>
<subSubSection id="78DE6504FFA7FFC28C8DF86AFDC877E1" box="[199,523,1974,2001]" pageId="36" pageNumber="187" type="distribution">
<paragraph id="307B368FFFA7FFC28C8DF86AFDC877E1" blockId="36.[151,1437,1470,2001]" box="[199,523,1974,2001]" pageId="36" pageNumber="187">
<emphasis id="02B0EA9DFFA7FFC28C8DF86AFE9A77E0" bold="true" box="[199,345,1974,2000]" pageId="36" pageNumber="187">Distribution</emphasis>
.
<collectingCountry id="48D3761FFFA7FFC28D2DF86AFE5977E0" box="[359,410,1974,2000]" name="Peru" pageId="36" pageNumber="187">Peru</collectingCountry>
(
<figureCitation id="A8FF2A0AFFA7FFC28DE0F86AFE3D77E1" box="[426,510,1974,2001]" captionStart="FIGURE 25" captionStartId="39.[151,250,1041,1065]" captionTargetBox="[184,1404,188,1011]" captionTargetId="figure-21@39.[177,1411,181,1018]" captionTargetPageId="39" captionText="FIGURE 25. Distribution of Goedia spp. in northern South America: Goeldia diva (pentagon—pink); G. obscura (square— yellow); G. guayaquilensis (circle—purple); G. mirim (star—red); G.utcuyacu (rectangle—blue). Type species are marked as black versions of each symbol when there are records to multiple localities." figureDoi="http://doi.org/10.5281/zenodo.10847445" httpUri="https://zenodo.org/record/10847445/files/figure.png" pageId="36" pageNumber="187">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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