treatments-xml/data/76/5D/6E/765D6E15FFAEB569FF7AFAACD7A8FBE7.xml

<document id="3756AF5D719D7A0DEA7044C15C9B3D3A" ID-DOI="10.11865/zs.2024206" ID-ISSN="2095-6827" ID-Zenodo-Dep="12172203" ID-ZooBank="5FB03411-A88D-4DFD-9253-41B59CD2D1E1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718829292756" checkinUser="felipe" docAuthor="Yao, Chonghui, Jiang, Chao &amp; Li, Weichun" docDate="2024" docId="765D6E15FFAEB569FF7AFAACD7A8FBE7" docLanguage="en" docName="ZoolSyst.49.2.181-183.pdf" docOrigin="Zoological Systematics 49 (2)" docSource="http://zoobank.org/5fb03411-a88d-4dfd-9253-41b59cd2d1e1" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Rhyscotus subrisus Li &amp; Jiang 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="183" masterDocId="8A64166DFFAEB56BFFF2FFBAD403FFA0" masterDocTitle="Rhyscotus subrisus Li &amp; Jiang, sp. nov., representing a new record family Rhyscotidae from China (Isopoda: Oniscidea)" masterLastPageNumber="183" masterPageNumber="181" pageNumber="181" updateTime="1718829649839" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4CDE3A0B7AE0033A2CE33EC73362D605">Rhyscotus subrisus Li &amp; Jiang, sp. nov., representing a new record family Rhyscotidae from China (Isopoda: Oniscidea)</mods:title>
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<mods:namePart id="459398F5978130A38D4CC681FAE8B1D4">Yao, Chonghui</mods:namePart>
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<taxonomicName id="39F4A480FFAEB56BFF7AFAACD5DCFA8E" authority="Li &amp; Jiang" authorityName="Li &amp; Jiang" authorityYear="2024" box="[136,479,1302,1326]" class="Malacostraca" family="Rhyscotidae" genus="Rhyscotus" kingdom="Animalia" order="Isopoda" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="subrisus" status="sp. nov.">
<emphasis id="CC800311FFAEB56BFF7AFAACD558FA8E" bold="true" box="[136,347,1302,1326]" italics="true" pageId="0" pageNumber="181">Rhyscotus subrisus</emphasis>
Li &amp; Jiang
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, 
<taxonomicNameLabel id="D7B3BE6AFFAEB56BFE18FAADD643FA8F" box="[490,576,1303,1327]" pageId="0" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="66CFC386FFAEB56BFD90FAACD6C2FA8F" box="[610,705,1302,1327]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="1.[155,218,1123,1145]" captionStartId-1="1.[155,218,2019,2041]" captionTargetBox-0="[203,1413,261,1092]" captionTargetBox-1="[206,1410,1207,1981]" captionTargetId-0="figure-93@1.[203,1413,261,1092]" captionTargetId-1="figure-95@1.[203,1413,1204,1983]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionText-0="Figure 1. Habitus of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype.A. Dorsal view. B. Lateral view. C. Head in frontal view. D. Pleon in dorsal view. E. Pleon in ventral view. Scale bars: A–B=1.0mm; C–E=0.5mm." captionText-1="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi-0="http://doi.org/10.5281/zenodo.12172205" figureDoi-1="http://doi.org/10.5281/zenodo.12172207" httpUri-0="https://zenodo.org/record/12172205/files/figure.png" httpUri-1="https://zenodo.org/record/12172207/files/figure.png" pageId="0" pageNumber="181">Figs 1–2</figureCitation>
)
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</paragraph>
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<paragraph id="FE4BDF03FFAEB56BFF4FFAF4D5D9FA0B" blockId="0.[136,1452,1358,1867]" pageId="0" pageNumber="181">
Type material. 
<materialsCitation id="4E9CD55EFFAEB56BFE96FAF4D53AFA29" collectingDate="2023-02-09" collectionCode="JXAUM" collectorName="Haocong Yang" country="China" elevation="10" latitude="18.283333" location="Jiyang" longLatPrecision="1275" longitude="109.566666" municipality="Sanya" pageId="0" pageNumber="181" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<typeStatus id="214F61A1FFAEB56BFE96FAF4D5C4FAC6" box="[356,455,1358,1382]" pageId="0" pageNumber="181" type="holotype">Holotype</typeStatus>
<specimenCount id="E8F2148AFFAEB56BFE22FAEBD5E7FAC5" box="[464,484,1361,1381]" count="1" pageId="0" pageNumber="181" type="male">♂</specimenCount>
, 
<collectingCountry id="86E39F93FFAEB56BFE01FAF4D634FAC6" box="[499,567,1358,1382]" name="China" pageId="0" pageNumber="181">China</collectingCountry>
: 
<collectingRegion id="3C3011E1FFAEB56BFDB1FAF4D690FAC6" box="[579,659,1358,1382]" country="China" name="Hainan" pageId="0" pageNumber="181">Hainan</collectingRegion>
, 
<collectingMunicipality id="1E2F4579FFAEB56BFD6DFAF4D6E0FAC6" box="[671,739,1358,1382]" pageId="0" pageNumber="181">Sanya</collectingMunicipality>
, 
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(
<geoCoordinate id="9BC0B9C4FFAEB56BFCB4FAF4D7AEFAC5" box="[838,941,1358,1382]" degrees="109" direction="east" minutes="34" orientation="longitude" pageId="0" pageNumber="181" precision="925" value="109.566666">109°34′E</geoCoordinate>
, 
<geoCoordinate id="9BC0B9C4FFAEB56BFC48FAF4D015FAC6" box="[954,1046,1358,1382]" degrees="18" direction="north" minutes="17" orientation="latitude" pageId="0" pageNumber="181" precision="925" value="18.283333">18°17′N</geoCoordinate>
), elev. 
<quantity id="390C72E6FFAEB56BFB95FAF4D09EFAC5" box="[1127,1181,1358,1382]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="181" unit="m" value="10.0">
<elevation id="75D93830FFAEB56BFB95FAF4D09EFAC5" box="[1127,1181,1358,1382]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="181" unit="m" value="10.0">10 m</elevation>
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, 
<date id="8A4AF9C3FFAEB56BFB58FAF4D10DFAC6" box="[1194,1294,1358,1382]" pageId="0" pageNumber="181" value="2023-02-09">
<collectingDate id="9A0E002BFFAEB56BFB58FAF4D10DFAC6" box="[1194,1294,1358,1382]" pageId="0" pageNumber="181" value="2023-02-09">9.II.2023</collectingDate>
</date>
, leg. 
<collectorName id="5301BAD5FFAEB56BFAB9FAF4D4C1FA29" pageId="0" pageNumber="181">Haocong Yang</collectorName>
(
<collectionCode id="98E547C6FFAEB56BFF3DFACBD532FA29" box="[207,305,1393,1417]" pageId="0" pageNumber="181">JXAUM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="4E9CD55EFFAEB56BFEB0FACBD6FBFA29" box="[322,760,1393,1417]" collectingDate="2023-02-09" collectionCode="JXAUM" collectorName="Haocong Yang" country="China" elevation="10" latitude="18.283333" location="Jiyang" longLatPrecision="1275" longitude="109.566666" municipality="Sanya" pageId="0" pageNumber="181" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<typeStatus id="214F61A1FFAEB56BFEB0FACBD5AEFA29" box="[322,429,1393,1417]" pageId="0" pageNumber="181" type="paratype">Paratypes</typeStatus>
. 
<specimenCount id="E8F2148AFFAEB56BFE45FACBD5DBFA28" box="[439,472,1393,1417]" count="1" pageId="0" pageNumber="181" type="female">1♀</specimenCount>
, same data as the holotype
</materialsCitation>
; 
<materialsCitation id="4E9CD55EFFAEB56BFCF3FACBD5D5FA0B" collectingDate="2023-06-09" collectionCode="CMMI" collectorName="Chao Jiang" country="China" elevation="10" latitude="18.283333" location="Jiyang" longLatPrecision="1275" longitude="109.566666" municipality="Sanya" pageId="0" pageNumber="181" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="E8F2148AFFAEB56BFCF3FACBD720FA28" box="[769,803,1393,1417]" count="2" pageId="0" pageNumber="181" type="male">2♂</specimenCount>
<specimenCount id="E8F2148AFFAEB56BFCD0FACBD747FA28" box="[802,836,1393,1417]" count="2" pageId="0" pageNumber="181" type="female">2♀</specimenCount>
, same data as the holotype except for collected date 
<date id="8A4AF9C3FFAEB56BFA9DFACBD4C3FA0B" pageId="0" pageNumber="181" value="2023-06-09">
<collectingDate id="9A0E002BFFAEB56BFA9DFACBD4C3FA0B" pageId="0" pageNumber="181" value="2023-06-09">9. VI. 2023</collectingDate>
</date>
, leg. 
<docAuthor id="559CDE8EFFAEB56BFF0BFA29D573FA0B" box="[249,368,1427,1451]" pageId="0" pageNumber="181">
<collectorName id="5301BAD5FFAEB56BFF0BFA29D573FA0B" box="[249,368,1427,1451]" pageId="0" pageNumber="181">Chao Jiang</collectorName>
</docAuthor>
(
<collectionCode id="98E547C6FFAEB56BFE8CFA29D5CDFA0B" box="[382,462,1427,1451]" country="China" lsid="urn:lsid:biocol.org:col:12805" name="Chinese Academy of Traditional Medicine" pageId="0" pageNumber="181" type="Herbarium">CMMI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="FE4BDF03FFAEB56BFF4FFA0CD7A3FA51" blockId="0.[136,1452,1358,1867]" pageId="0" pageNumber="181">
Etymology. The specific name is derived from the Latin 
<emphasis id="CC800311FFAEB56BFCC5FA0CD7A5FA6E" box="[823,934,1462,1486]" italics="true" pageId="0" pageNumber="181">
<taxonomicName id="39F4A480FFAEB56BFCC5FA0CD793FA6E" authorityName="Li &amp; Jiang" authorityYear="2024" box="[823,912,1462,1486]" class="Malacostraca" family="Rhyscotidae" genus="Rhyscotus" kingdom="Animalia" order="Isopoda" pageId="0" pageNumber="181" phylum="Arthropoda" rank="species" species="subrisus">subrisus</taxonomicName>
=
</emphasis>
smile, in reference to the dorsal surface of the species with a pattern like a smiling face on the first and second pleonites.
</paragraph>
<paragraph id="FE4BDF03FFAEB56BFF4FFA41D640F996" blockId="0.[136,1452,1358,1867]" pageId="0" pageNumber="181">Diagnosis. Pleopod 1 endopodite almost three times as long as the longest part of exopodite, and male pleopod 2 exopodite with a finger-shaped apical tip.</paragraph>
<paragraph id="FE4BDF03FFAEB56BFF4FF9FBD618F961" blockId="0.[136,1452,1358,1867]" pageId="0" pageNumber="181">
Description. Body length 5.0–7.0 mm. Colour of dorsal surface white with brown blotches. Each pereonite with two triangular blotches and two incurved stipes near lateral side (
<figureCitation id="66CFC386FFAEB56BFCC7F9D9D7B1F9DB" box="[821,946,1635,1659]" captionStart="Figure 1" captionStartId="1.[155,218,1123,1145]" captionTargetBox="[203,1413,261,1092]" captionTargetId="figure-93@1.[203,1413,261,1092]" captionTargetPageId="1" captionText="Figure 1. Habitus of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype.A. Dorsal view. B. Lateral view. C. Head in frontal view. D. Pleon in dorsal view. E. Pleon in ventral view. Scale bars: A–B=1.0mm; C–E=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12172205" httpUri="https://zenodo.org/record/12172205/files/figure.png" pageId="0" pageNumber="181">Figs 1A–B</figureCitation>
). Pleon narrower than pereon, epimera short; pleonites 1 and 2 with smiling face pattern near middle (
<figureCitation id="66CFC386FFAEB56BFD14F93CD739F93E" box="[742,826,1670,1694]" captionStart="Figure 1" captionStartId="1.[155,218,1123,1145]" captionTargetBox="[203,1413,261,1092]" captionTargetId="figure-93@1.[203,1413,261,1092]" captionTargetPageId="1" captionText="Figure 1. Habitus of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype.A. Dorsal view. B. Lateral view. C. Head in frontal view. D. Pleon in dorsal view. E. Pleon in ventral view. Scale bars: A–B=1.0mm; C–E=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12172205" httpUri="https://zenodo.org/record/12172205/files/figure.png" pageId="0" pageNumber="181">Fig. 1D</figureCitation>
). Telson nearly triangular, distinctively concave on lateral margin, distal apex pointed (
<figureCitation id="66CFC386FFAEB56BFE4BF913D60EF961" box="[441,525,1705,1729]" captionStart="Figure 1" captionStartId="1.[155,218,1123,1145]" captionTargetBox="[203,1413,261,1092]" captionTargetId="figure-93@1.[203,1413,261,1092]" captionTargetPageId="1" captionText="Figure 1. Habitus of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype.A. Dorsal view. B. Lateral view. C. Head in frontal view. D. Pleon in dorsal view. E. Pleon in ventral view. Scale bars: A–B=1.0mm; C–E=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12172205" httpUri="https://zenodo.org/record/12172205/files/figure.png" pageId="0" pageNumber="181">Fig. 1D</figureCitation>
).
</paragraph>
<paragraph id="FE4BDF03FFAEB56BFF4FF971D5DBF8EB" blockId="0.[136,1452,1358,1867]" pageId="0" pageNumber="181">
Head white with irregular brown patterns; frons brown, bearing fourteen white stripes and three white dots (
<figureCitation id="66CFC386FFAEB56BFAD4F971D19CF943" box="[1318,1439,1739,1763]" captionStart="Figure 1" captionStartId="1.[155,218,1123,1145]" captionTargetBox="[203,1413,261,1092]" captionTargetId="figure-93@1.[203,1413,261,1092]" captionTargetPageId="1" captionText="Figure 1. Habitus of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype.A. Dorsal view. B. Lateral view. C. Head in frontal view. D. Pleon in dorsal view. E. Pleon in ventral view. Scale bars: A–B=1.0mm; C–E=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12172205" httpUri="https://zenodo.org/record/12172205/files/figure.png" pageId="0" pageNumber="181">Figs 1A− C</figureCitation>
). Antenna with second flagellar article almost three times as long as first one, apical organ thin and short (
<figureCitation id="66CFC386FFAEB56BFB27F954D129F8A6" box="[1237,1322,1774,1798]" captionStart="Figure 2" captionStartId="1.[155,218,2019,2041]" captionTargetBox="[206,1410,1207,1981]" captionTargetId="figure-95@1.[203,1413,1204,1983]" captionTargetPageId="1" captionText="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="0" pageNumber="181">Fig. 2A</figureCitation>
). Pereopods without apparent sexual specializations, pereopod 1 without particular modifications, ischium of pereopod 7 deeply concave on ventral margin (
<figureCitation id="66CFC386FFAEB56BFEA6F889D5CFF8EB" box="[340,460,1843,1867]" captionStart="Figure 2" captionStartId="1.[155,218,2019,2041]" captionTargetBox="[206,1410,1207,1981]" captionTargetId="figure-95@1.[203,1413,1204,1983]" captionTargetPageId="1" captionText="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="0" pageNumber="181">Figs 2B–C</figureCitation>
).
</paragraph>
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<paragraph id="FE4BDF03FFAFB56AFF69FBD9D7ADFB39" blockId="1.[155,1461,1123,1177]" pageId="1" pageNumber="182">
Figure 1. Habitus of 
<taxonomicName id="39F4A480FFAFB56AFE92FBD9D680FBD9" authority="Li &amp; Jiang" authorityName="Li &amp; Jiang" authorityYear="2024" box="[352,643,1123,1145]" class="Malacostraca" family="Rhyscotidae" genus="Rhyscotus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="182" phylum="Arthropoda" rank="species" species="subrisus" status="sp. nov.">
<emphasis id="CC800311FFAFB56AFE92FBD9D615FBD8" box="[352,534,1123,1144]" italics="true" pageId="1" pageNumber="182">Rhyscotus subrisus</emphasis>
Li &amp; Jiang
</taxonomicName>
, 
<emphasis id="CC800311FFAFB56AFD7EFBD9D6DBFBD8" bold="true" box="[652,728,1123,1144]" pageId="1" pageNumber="182">
<taxonomicNameLabel id="D7B3BE6AFFAFB56AFD7EFBD9D6DBFBD8" box="[652,728,1123,1144]" pageId="1" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype. A. Dorsal view. B. Lateral view. C. Head in frontal view. D. Pleon in dorsal view. E. Pleon in ventral view. Scale bars: A–B = 1.0 mm; C–E = 0.5 mm.
</paragraph>
</caption>
<caption id="AA8B8F8BFFAFB56AFF69F859D0ADF7B9" ID-DOI="http://doi.org/10.5281/zenodo.12172207" ID-Zenodo-Dep="12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="1" pageNumber="182" startId="1.[155,218,2019,2041]" targetBox="[206,1410,1207,1981]" targetPageId="1" targetType="figure">
<paragraph id="FE4BDF03FFAFB56AFF69F859D0ADF7B9" blockId="1.[155,1461,2019,2073]" pageId="1" pageNumber="182">
Figure 2. Appendages of 
<taxonomicName id="39F4A480FFAFB56AFE61F859D6BEF859" authority="Li &amp; Jiang" authorityName="Li &amp; Jiang" authorityYear="2024" box="[403,701,2019,2041]" class="Malacostraca" family="Rhyscotidae" genus="Rhyscotus" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="182" phylum="Arthropoda" rank="species" species="subrisus" status="sp. nov.">
<emphasis id="CC800311FFAFB56AFE61F859D648F858" box="[403,587,2019,2040]" italics="true" pageId="1" pageNumber="182">Rhyscotus subrisus</emphasis>
Li &amp; Jiang
</taxonomicName>
, 
<emphasis id="CC800311FFAFB56AFD3AF859D716F858" bold="true" box="[712,789,2019,2040]" pageId="1" pageNumber="182">
<taxonomicNameLabel id="D7B3BE6AFFAFB56AFD3AF859D716F858" box="[712,789,2019,2040]" pageId="1" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="FE4BDF03FFACB569FF4FFF56D698FE2F" blockId="2.[136,1452,236,642]" pageId="2" pageNumber="183">
Pleopods, sexual differentiation. Male pleopod 1 exopodite small and elliptic; endopodite approximate three times as long as longest part of exopodite, basal part broad, narrowed towards apical tip (
<figureCitation id="66CFC386FFACB569FC29FEB5D032FE87" box="[987,1073,271,295]" captionStart="Figure 2" captionStartId="1.[155,218,2019,2041]" captionTargetBox="[206,1410,1207,1981]" captionTargetId="figure-95@1.[203,1413,1204,1983]" captionTargetPageId="1" captionText="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="2" pageNumber="183">Fig. 2D</figureCitation>
). Male pleopod 2 exopodite convex on outer margin, apical tip finger-shaped; endopodite thin and long, distinctly longer than exopodite (
<figureCitation id="66CFC386FFACB569FB3FFE88D122FEEA" box="[1229,1313,306,330]" captionStart="Figure 2" captionStartId="1.[155,218,2019,2041]" captionTargetBox="[206,1410,1207,1981]" captionTargetId="figure-95@1.[203,1413,1204,1983]" captionTargetPageId="1" captionText="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="2" pageNumber="183">Fig. 2E</figureCitation>
). Pleopods 3 and 4 exopodites convex on outer margin, ending with triangular apical tip (
<figureCitation id="66CFC386FFACB569FC45FEEED02DFECD" box="[951,1070,340,365]" captionStart="Figure 2" captionStartId="1.[155,218,2019,2041]" captionTargetBox="[206,1410,1207,1981]" captionTargetId="figure-95@1.[203,1413,1204,1983]" captionTargetPageId="1" captionText="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="2" pageNumber="183">Figs 2F–G</figureCitation>
). Pleopod 5 nearly straight on outer margin, concave at distal one-fifth area (
<figureCitation id="66CFC386FFACB569FDCAFECDD68CFE2F" box="[568,655,375,399]" captionStart="Figure 2" captionStartId="1.[155,218,2019,2041]" captionTargetBox="[206,1410,1207,1981]" captionTargetId="figure-95@1.[203,1413,1204,1983]" captionTargetPageId="1" captionText="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="2" pageNumber="183">Fig. 2H</figureCitation>
).
</paragraph>
<paragraph id="FE4BDF03FFACB569FF4FFE20D5FBFE12" blockId="2.[136,1452,236,642]" box="[189,504,410,434]" pageId="2" pageNumber="183">
Distribution. 
<collectingCountry id="86E39F93FFACB569FEBEFE20D58FFE12" box="[332,396,410,434]" name="China" pageId="2" pageNumber="183">China</collectingCountry>
(
<collectingRegion id="3C3011E1FFACB569FE68FE20D5EEFE12" box="[410,493,410,434]" country="China" name="Hainan" pageId="2" pageNumber="183">Hainan</collectingRegion>
).
</paragraph>
<paragraph id="FE4BDF03FFACB569FF4FFE06D6FEFD22" blockId="2.[136,1452,236,642]" pageId="2" pageNumber="183">
Remarks. The new species is similar to 
<taxonomicName id="39F4A480FFACB569FD97FE07D02AFE75" authority="Schmalfuss &amp; Ferrara, 1978" authorityName="Schmalfuss &amp; Ferrara" authorityYear="1978" box="[613,1065,444,469]" class="Malacostraca" family="Rhyscotidae" genus="Rhyscotus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="CC800311FFACB569FD97FE07D6F5FE74" box="[613,758,444,468]" italics="true" pageId="2" pageNumber="183">R. rotundatus</emphasis>
Schmalfuss &amp; Ferrara, 1978
</taxonomicName>
by pointed apex of telson and small and elliptic pleopod 1 exopodite (
<figureCitation id="66CFC386FFACB569FE19FE65D640FE57" box="[491,579,479,503]" captionStart="Figure 1" captionStartId="1.[155,218,1123,1145]" captionTargetBox="[203,1413,261,1092]" captionTargetId="figure-93@1.[203,1413,261,1092]" captionTargetPageId="1" captionText="Figure 1. Habitus of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype.A. Dorsal view. B. Lateral view. C. Head in frontal view. D. Pleon in dorsal view. E. Pleon in ventral view. Scale bars: A–B=1.0mm; C–E=0.5mm." figureDoi="http://doi.org/10.5281/zenodo.12172205" httpUri="https://zenodo.org/record/12172205/files/figure.png" pageId="2" pageNumber="183">Figs 1D</figureCitation>
, 
<figureCitation id="66CFC386FFACB569FDBCFE65D66CFE57" box="[590,623,479,503]" captionStart="Figure 2" captionStartId="1.[155,218,2019,2041]" captionTargetBox="[206,1410,1207,1981]" captionTargetId="figure-95@1.[203,1413,1204,1983]" captionTargetPageId="1" captionText="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="2" pageNumber="183">2D</figureCitation>
<emphasis id="CC800311FFACB569FD87FE5AD691FE57" box="[629,658,480,503]" italics="true" pageId="2" pageNumber="183">vs.</emphasis>
Schmalfuss &amp; Ferrara, 1978: figs 22, 26). But it can be distinguished from the latter species by the features of pleopods 1 and 2: pleopod 1 endopodite of the new species almost three times as long as the longest part of exopodite, and male pleopod 2 exopodite with a finger-shaped apical tip (
<figureCitation id="66CFC386FFACB569FB98FD9ED0E1FD9D" box="[1130,1250,548,573]" captionStart="Figure 2" captionStartId="1.[155,218,2019,2041]" captionTargetBox="[206,1410,1207,1981]" captionTargetId="figure-95@1.[203,1413,1204,1983]" captionTargetPageId="1" captionText="Figure 2. Appendages of Rhyscotus subrisus Li &amp; Jiang, sp. nov., holotype. A. Antenna. B. Pereopod 1. C. Pereopod 7. D. Pleopod 1. E. Pleopod 2. F. Pleopod 3 exopodite. G. Pleopod 4 exopodite. H. Pleopod 5 exopodite. Scale bars=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.12172207" httpUri="https://zenodo.org/record/12172207/files/figure.png" pageId="2" pageNumber="183">Figs 2D, E</figureCitation>
); in 
<taxonomicName id="39F4A480FFACB569FAE1FD9FD1A7FD9C" authorityName="Schmalfuss &amp; Ferrara" authorityYear="1978" box="[1299,1444,548,572]" class="Malacostraca" family="Rhyscotidae" genus="Rhyscotus" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="CC800311FFACB569FAE1FD9FD1A7FD9C" box="[1299,1444,548,572]" italics="true" pageId="2" pageNumber="183">R. rotundatus</emphasis>
</taxonomicName>
, the pleopod 1 endopodite about twice as long as the longest part of exopodite, and the male pleopod 2 exopodite with a round apical tip (Schmalfuss &amp; Ferrara, 1978: figs 26−28).
</paragraph>
<paragraph id="FE4BDF03FFACB569FF7AFD0CD153FD6E" blockId="2.[136,1360,694,718]" box="[136,1360,694,718]" pageId="2" pageNumber="183">
<emphasis id="CC800311FFACB569FF7AFD0CD4EBFD6E" bold="true" box="[136,232,694,718]" inLineHeading="true" pageId="2" pageNumber="183" reason="1">Funding</emphasis>
This study was supported by the National Natural Science Foundation of 
<collectingCountry id="86E39F93FFACB569FBFCFD0CD04DFD6E" box="[1038,1102,694,718]" name="China" pageId="2" pageNumber="183">China</collectingCountry>
(31960100, 82073972).
</paragraph>
</subSubSection>
<subSubSection id="B6EE8C88FFACB569FF7AFCB8D7A8FBE7" pageId="2" pageNumber="183" type="description">
<paragraph id="FE4BDF03FFACB569FF7AFCB8D11AFC9D" blockId="2.[136,1452,770,829]" pageId="2" pageNumber="183">
<emphasis id="CC800311FFACB569FF7AFCB8D566FCBA" bold="true" box="[136,357,770,794]" inLineHeading="true" pageId="2" pageNumber="183" reason="1">Acknowledgements</emphasis>
We are grateful to Mr. Haocong Yang for providing the specimens. Special thanks are given to Dr. Ghasem M. Kashani (University of 
<collectingRegion id="3C3011E1FFACB569FDF5FC9FD657FC9D" box="[519,596,805,829]" country="Iran" name="Zanjan" pageId="2" pageNumber="183">Zanjan</collectingRegion>
, 
<collectingCountry id="86E39F93FFACB569FDACFC9FD688FC9D" box="[606,651,805,829]" name="Iran" pageId="2" pageNumber="183">Iran</collectingCountry>
) and an anonymous reviewer for their insightful suggestions.
</paragraph>
<paragraph id="FE4BDF03FFACB569FC82FC2DD795FC47" blockId="2.[880,1451,918,946]" lastBlockId="2.[136,1451,978,1095]" pageId="2" pageNumber="183">
Chonghui Yao1, Chao Jiang2 *, Weichun Li1 * 
<emphasis id="CC800311FFACB569FF7AFC75D795FC47" box="[136,918,975,1000]" italics="true" pageId="2" pageNumber="183">
1College of Agronomy, 
<collectingRegion id="3C3011E1FFACB569FE95FC69D5ADFC48" box="[359,430,979,1000]" country="China" name="Jiangxi" pageId="2" pageNumber="183">Jiangxi</collectingRegion>
Agricultural University, Nanchang 330045, 
<collectingCountry id="86E39F93FFACB569FCA9FC68D795FC47" box="[859,918,978,999]" name="China" pageId="2" pageNumber="183">China</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="FE4BDF03FFACB569FF7AFC55D777FB87" blockId="2.[136,1451,978,1095]" pageId="2" pageNumber="183">
<emphasis id="CC800311FFACB569FF7AFC55D777FB87" italics="true" pageId="2" pageNumber="183">
<superScript id="0981724BFFACB569FF7AFC55D493FC5D" attach="right" box="[136,144,1007,1021]" fontSize="6" pageId="2" pageNumber="183">2</superScript>
State Key Laboratory for Quality Ensurance and Sustainable Use of Dao-di Herbs, National Resource Center for Chinese Materia Medica, 
<collectingCountry id="86E39F93FFACB569FF2EFBA8D514FB87" box="[220,279,1042,1063]" name="China" pageId="2" pageNumber="183">China</collectingCountry>
Academy of Chinese Medical Sciences, 
<collectingRegion id="3C3011E1FFACB569FD68FBA9D6DCFB88" box="[666,735,1043,1064]" country="China" name="Beijing" pageId="2" pageNumber="183">Beijing</collectingRegion>
100700, 
<collectingCountry id="86E39F93FFACB569FCCBFBA8D777FB87" box="[825,884,1042,1063]" name="China" pageId="2" pageNumber="183">China</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="FE4BDF03FFACB569FF7AFB8AD7A8FBE7" blockId="2.[136,1451,978,1095]" box="[136,939,1072,1095]" pageId="2" pageNumber="183">
<emphasis id="CC800311FFACB569FF7AFB8AD7A8FBE7" box="[136,939,1072,1095]" italics="true" pageId="2" pageNumber="183">
<superScript id="0981724BFFACB569FF7AFB8AD493FB9E" attach="right" box="[136,144,1072,1086]" fontSize="6" pageId="2" pageNumber="183">*</superScript>
Corresponding authors, E-mails: jiangchao0411@126.com, weichunlee@126.com
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>