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<document id="C1E998F96B3453108C9F191C20DA17D1" ID-DOI="10.1016/j.sajb.2018.11.021" ID-ISSN="1727-9321" ID-Zenodo-Dep="10496596" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1705062987214" checkinUser="felipe" docAuthor="Elliott, T. L., Barrett, R. L. &amp; Muasya, A. M." docDate="2019" docId="68465A3B6B751D420008D77BF184FAAA" docLanguage="en" docName="SAfrJBot.121.519-535.pdf" docOrigin="South African Journal of Botany 121" docSource="http://dx.doi.org/10.1016/j.sajb.2018.11.021" docStyle="DocumentStyle:AD72C92759D651943977BAEA6E2218B4.1:SAfrJBot.2014-.journal_article" docStyleId="AD72C92759D651943977BAEA6E2218B4" docStyleName="SAfrJBot.2014-.journal_article" docStyleVersion="1" docTitle="Schoenus cuspidatus Rottb." docType="key" docVersion="1" lastPageNumber="521" masterDocId="947F22436B741D40032DD230F45EFFF9" masterDocTitle="A taxonomic revision of Schoenus cuspidatus and allies (Cyperaceae, tribe Schoeneae) - Part 1 *" masterLastPageNumber="535" masterPageNumber="519" pageNumber="520" updateTime="1705673609269" updateUser="guilherme">
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<mods:title id="5A4F3C25B48CCC3746D6F3615B24DF63">A taxonomic revision of Schoenus cuspidatus and allies (Cyperaceae, tribe Schoeneae) - Part 1 *</mods:title>
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<mods:namePart id="5B69AC7C1C2ED75B1409053B630DFEF2">Elliott, T. L.</mods:namePart>
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<heading id="BB185C416B751D410008D77BF1D6FAA7" bold="true" box="[805,1416,1354,1374]" fontSize="36" level="1" pageId="1" pageNumber="520" reason="1">
<emphasis id="D29B373F6B751D410008D77BF1D6FAA7" bold="true" box="[805,1416,1354,1374]" pageId="1" pageNumber="520">
4. Key to the species in the
<taxonomicName id="27EF90AE6B751D41071FD77AF0A4FAA7" ID-CoL="4VF4H" authority="Rottb." authorityName="Rottb." box="[1074,1274,1354,1374]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="1" pageNumber="520" phylum="Tracheophyta" rank="species" species="cuspidatus">
<emphasis id="D29B373F6B751D41071FD77AF0A4FAA7" bold="true" box="[1074,1274,1354,1374]" italics="true" pageId="1" pageNumber="520">Schoenus cuspidatus</emphasis>
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group (part I)
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1a. Proximal primary inflorescence bract conspicuously expanded at base; longitudinal veins at base of bract often prominent: 2a. Culms usually thick (
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wide) and appearing semi-succulent; plants widespread throughout
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(usually found within
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of the S. and E. coasts): ……………………………………………………………
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.
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<emphasis id="D29B373F6B751D410631D43DF19CF9D8" bold="true" box="[1308,1474,1549,1569]" italics="true" pageId="1" pageNumber="520">Schoenus auritus</emphasis>
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2b. Culms thin (
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) and not appearing semi-succulent; plants restricted to granitic sites on the
<collectingRegion id="222B25CF6B751D410635D476F119F9A0" box="[1304,1351,1606,1625]" country="South Africa" name="Western Cape" pageId="1" pageNumber="520">Cape</collectingRegion>
Peninsula: ……………………………………………………
<emphasis id="D29B373F6B751D4107E4D452F08BF98C" bold="true" box="[1225,1237,1634,1653]" pageId="1" pageNumber="520">2</emphasis>
.
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<emphasis id="D29B373F6B751D4107F3D450F1E2F98C" bold="true" box="[1246,1468,1632,1653]" italics="true" pageId="1" pageNumber="520">Schoenus graminifolius</emphasis>
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1b. Proximal primary inflorescence bract not conspicuously expanded at base; longitudinal veins at base of bract not prominent: 3a. Spikelets linear-lanceolate (
<figureCitation id="78D4F7A86B751D410754D485F0F2F930" box="[1145,1196,1717,1737]" captionStart="Fig" captionStartId="2.[113,138,384,398]" captionTargetBox="[351,1261,142,346]" captionTargetId="figure-573@2.[350,1261,142,349]" captionTargetPageId="2" captionText="Fig. 1. Spikelet variation in the Schoenus cuspidatus and allies group found in southern Africa: A—S. cuspidatus (lanceolate spikelet), B—S. crassus (lanceolate spikelet), C—S. auritus (lanceolate spikelet), D—S. graminifolius (lanceolate spikelet), E—S. calceolus (ovate spikelet) and F—S. ligulatus (linear-lanceolate spikelet).The black scale bar next to the spikelets represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496598" httpUri="https://zenodo.org/record/10496598/files/figure.png" pageId="1" pageNumber="520">Fig. 1</figureCitation>
); nutlets narrow-ellipsoidal (
<figureCitation id="78D4F7A86B751D410003D4E1F7D4F91D" box="[814,906,1745,1764]" captionStart="Fig" captionStartId="3.[90,115,602,616]" captionTargetBox="[328,1238,143,565]" captionTargetId="figure-246@3.[327,1238,142,566]" captionTargetPageId="3" captionText="Fig.2. Examples of nutlets from 11 species of the Schoenus cuspidatus and allies group found in southern Africa:A—S.auritus; B—S.graminifolius; C—S.ligulatus; D—S.exilis; E—S.bolusii; F— S.submarginalis; G—S.crassiculmis; H—S.quartziticus; I—S.limosus; J—S.purpurascens; and K—S.calceolus (immature).The black scale bar represents 1 mm.The mature fruits of S.calceolus and S.prophyllus are unknown." figureDoi="http://doi.org/10.5281/zenodo.10496600" httpUri="https://zenodo.org/record/10496600/files/figure.png" pageId="1" pageNumber="520">Fig. 2C, D</figureCitation>
): 4a. Plants usually&gt;
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in height; spikelets
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in length, often reduced sexually with only male or female organs present:………………………………………………
<emphasis id="D29B373F6B751D4107DCD515F0A3F8C1" bold="true" box="[1265,1277,1829,1848]" pageId="1" pageNumber="520">3</emphasis>
.
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<emphasis id="D29B373F6B751D41062AD514F1E5F8C1" bold="true" box="[1287,1467,1828,1848]" italics="true" pageId="1" pageNumber="520">Schoenus ligulatus</emphasis>
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4b. Plants &lt;
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in height; spikelets mostly &lt;
<quantity id="271746C86B751D41060BD571F13EF8AD" box="[1318,1376,1857,1876]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="520" unit="mm" value="4.0">4 mm</quantity>
in length, bisexual spikelets always present:………………………
<emphasis id="D29B373F6B751D410634D56DF17BF889" bold="true" box="[1305,1317,1885,1904]" pageId="1" pageNumber="520">4</emphasis>
.
<taxonomicName id="27EF90AE6B751D410603D56CF1E2F889" ID-CoL="8X5LH" box="[1326,1468,1884,1904]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="1" pageNumber="520" phylum="Tracheophyta" rank="species" species="exilis">
<emphasis id="D29B373F6B751D410603D56CF1E2F889" bold="true" box="[1326,1468,1884,1904]" italics="true" pageId="1" pageNumber="520">Schoenus exilis</emphasis>
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3b. Spikelets lanceolate or ovate (
<figureCitation id="78D4F7A86B751D4107A1D549F0E1F875" box="[1164,1215,1913,1932]" captionStart="Fig" captionStartId="2.[113,138,384,398]" captionTargetBox="[351,1261,142,346]" captionTargetId="figure-573@2.[350,1261,142,349]" captionTargetPageId="2" captionText="Fig. 1. Spikelet variation in the Schoenus cuspidatus and allies group found in southern Africa: A—S. cuspidatus (lanceolate spikelet), B—S. crassus (lanceolate spikelet), C—S. auritus (lanceolate spikelet), D—S. graminifolius (lanceolate spikelet), E—S. calceolus (ovate spikelet) and F—S. ligulatus (linear-lanceolate spikelet).The black scale bar next to the spikelets represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496598" httpUri="https://zenodo.org/record/10496598/files/figure.png" pageId="1" pageNumber="520">Fig. 1</figureCitation>
); nutlets broad-ellipsoidal (
<figureCitation id="78D4F7A86B751D410003D5A5F73EF851" box="[814,864,1941,1960]" captionStart="Fig" captionStartId="3.[90,115,602,616]" captionTargetBox="[328,1238,143,565]" captionTargetId="figure-246@3.[327,1238,142,566]" captionTargetPageId="3" captionText="Fig.2. Examples of nutlets from 11 species of the Schoenus cuspidatus and allies group found in southern Africa:A—S.auritus; B—S.graminifolius; C—S.ligulatus; D—S.exilis; E—S.bolusii; F— S.submarginalis; G—S.crassiculmis; H—S.quartziticus; I—S.limosus; J—S.purpurascens; and K—S.calceolus (immature).The black scale bar represents 1 mm.The mature fruits of S.calceolus and S.prophyllus are unknown." figureDoi="http://doi.org/10.5281/zenodo.10496600" httpUri="https://zenodo.org/record/10496600/files/figure.png" pageId="1" pageNumber="520">Fig. 2</figureCitation>
): 5a. Spikelets lanceolate:
</paragraph>
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<emphasis id="D29B373F6B761D42035CD3B0F4FEFE77" bold="true" box="[113,160,384,398]" pageId="2" pageNumber="521">Fig. 1.</emphasis>
Spikelet variation in the
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<emphasis id="D29B373F6B761D420249D34FF5A3FE76" box="[356,509,383,399]" italics="true" pageId="2" pageNumber="521">Schoenus cuspidatus</emphasis>
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and allies group found in southern Africa: A—
<taxonomicName id="27EF90AE6B761D420049D34FF79AFE76" box="[868,964,383,399]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="cuspidatus">
<emphasis id="D29B373F6B761D420049D34FF79AFE76" box="[868,964,383,399]" italics="true" pageId="2" pageNumber="521">S. cuspidatus</emphasis>
</taxonomicName>
(lanceolate spikelet), B—
<taxonomicName id="27EF90AE6B761D4207A4D34FF08EFE76" box="[1161,1232,383,399]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="crassus">
<emphasis id="D29B373F6B761D4207A4D34FF08EFE76" box="[1161,1232,383,399]" italics="true" pageId="2" pageNumber="521">S. crassus</emphasis>
</taxonomicName>
(lanceolate spikelet), C—
<taxonomicName id="27EF90AE6B761D4206B9D34FF184FE76" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" baseAuthorityName="T. L. Elliott &amp; Muasya" baseAuthorityYear="2019" box="[1428,1498,383,399]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="auritus">
<emphasis id="D29B373F6B761D4206B9D34FF184FE76" box="[1428,1498,383,399]" italics="true" pageId="2" pageNumber="521">S. auritus</emphasis>
</taxonomicName>
(lanceolate spikelet), D—
<taxonomicName id="27EF90AE6B761D42021FD3A6F5FBFE5C" box="[306,421,406,422]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="graminifolius">
<emphasis id="D29B373F6B761D42021FD3A6F5FBFE5C" box="[306,421,406,422]" italics="true" pageId="2" pageNumber="521">S. graminifolius</emphasis>
</taxonomicName>
(lanceolate spikelet), E—
<taxonomicName id="27EF90AE6B761D42014AD3A6F6E5FE5C" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[615,699,406,422]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="calceolus">
<emphasis id="D29B373F6B761D42014AD3A6F6E5FE5C" box="[615,699,406,422]" italics="true" pageId="2" pageNumber="521">S. calceolus</emphasis>
</taxonomicName>
(ovate spikelet) and F—
<taxonomicName id="27EF90AE6B761D42005BD3A6F799FE5C" box="[886,967,406,422]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="ligulatus">
<emphasis id="D29B373F6B761D42005BD3A6F799FE5C" box="[886,967,406,422]" italics="true" pageId="2" pageNumber="521">S. ligulatus</emphasis>
</taxonomicName>
(linear-lanceolate spikelet). The black scale bar next to the spikelets represents 1 mm.
</paragraph>
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<paragraph id="E050EB2D6B761D4203BCD3CBF757FD4C" blockId="2.[113,784,507,694]" pageId="2" pageNumber="521">
6a. Glume margins with reddish-purple streaks adjacent to the hyaline margins (
<figureCitation id="78D4F7A86B761D4203DBD027F510FDD3" box="[246,334,535,554]" captionStart="Fig" captionStartId="3.[90,115,1926,1940]" captionTargetBox="[252,1313,1076,1890]" captionTargetId="figure-340@3.[251,1313,1075,1890]" captionTargetPageId="3" captionText="Fig.3. Examples of plants from six species of the Schoenus cuspidatus and allies group found in southern Africa,showing the corresponding inflorescences andspikelets beneath each plant: A, G and M—S.auritus; B, H and N—S.graminifolius; C, I and O—S. ligulatus; D,J, and P—S.exilis; E,K and Q—S.bolusii; and F, L and R—S.submarginalis. The black scale bar adjacent to the inflorescences in G—L represents 10 mm, while the scale bars adjacent to the spikelets in M—R represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496602" httpUri="https://zenodo.org/record/10496602/files/figure.png" pageId="2" pageNumber="521">Fig. 3E, F</figureCitation>
): 7a: Older leaves usually curled; membranaceous sheaths with reddish-purple streaks:………………………………
<emphasis id="D29B373F6B761D420164D07FF75EFD98" bold="true" box="[585,768,589,610]" pageId="2" pageNumber="521">
5.
<taxonomicName id="27EF90AE6B761D42014DD07DF75EFD98" ID-CoL="8X5L7" authorityName="T. L. Elliott &amp; Muasya, S." authorityYear="2019" baseAuthorityName="C. B. Clarke" baseAuthorityYear="1898" box="[608,768,589,609]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="bolusii">
<emphasis id="D29B373F6B761D42014DD07DF75EFD98" bold="true" box="[608,768,589,609]" italics="true" pageId="2" pageNumber="521">Schoenus bolusii</emphasis>
</taxonomicName>
</emphasis>
7b. Older leaves usually straight; sheaths firm (i.e. not membranaceous) and solid red to brown in colour: ……………………………………………..……
<emphasis id="D29B373F6B761D420125D092F757FD4C" bold="true" box="[520,777,673,693]" pageId="2" pageNumber="521">
6.
<taxonomicName id="27EF90AE6B761D420132D091F757FD4C" ID-CoL="8XW54" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[543,777,673,693]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="submarginalis">
<emphasis id="D29B373F6B761D420132D091F757FD4C" bold="true" box="[543,777,673,693]" italics="true" pageId="2" pageNumber="521">Schoenus submarginalis</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="E050EB2D6B761D420071D3CBF184FAAA" blockId="2.[828,1499,507,1363]" pageId="2" pageNumber="521">
6b. Glume margins without reddish-purple streaks adjacent to the hyaline margins: 8a. Basal leaves reduced in length (usually a third to a quarter of culm length, never over half of culm length); glumes usually dark-reddish brown in colour; plants of mountainous regions of the Western and
<collectingRegion id="222B25CF6B761D420686D05BF7D1FD60" country="South Africa" name="Eastern Cape" pageId="2" pageNumber="521">Eastern Cape</collectingRegion>
Provinces:……….…….…….......………
<emphasis id="D29B373F6B761D4207C0D0B6F189FD60" bold="true" box="[1261,1495,645,665]" pageId="2" pageNumber="521">
7.
<taxonomicName id="27EF90AE6B761D42062ED0B5F189FD60" ID-CoL="8XW4Z" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[1283,1495,645,665]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="crassiculmis">
<emphasis id="D29B373F6B761D42062ED0B5F189FD60" bold="true" box="[1283,1495,645,665]" italics="true" pageId="2" pageNumber="521">Schoenus crassiculmis</emphasis>
</taxonomicName>
</emphasis>
8b. Basal leaves well-developed in length (often over half of culm length); glumes medium brown to light reddish-brown in colour; plants endemic to the
<collectingRegion id="222B25CF6B761D420732D0EAF0C1FD14" box="[1055,1183,730,749]" country="South Africa" name="Eastern Cape" pageId="2" pageNumber="521">Eastern Cape</collectingRegion>
and
<collectingRegion id="222B25CF6B761D4207F9D0EAF139FD14" box="[1236,1383,730,749]" country="South Africa" name="KwaZulu-Natal" pageId="2" pageNumber="521">KwaZulu-Natal</collectingRegion>
or
<location id="E530BDF66B761D4206A6D0EAF7E8FCF0" LSID="urn:lsid:plazi:treatment:68465A3B6B751D420008D77BF184FAAA:E530BDF66B761D4206A6D0EAF7E8FCF0" country="South Africa" name="Agulhas Plains region" pageId="2" pageNumber="521" stateProvince="Eastern Cape">Agulhas Plains region</location>
of
<collectingCountry id="98F8ABBD6B761D4200FED0C6F010FCF0" box="[979,1102,758,777]" name="South Africa" pageId="2" pageNumber="521">South Africa</collectingCountry>
): 9a. Spikelets &lt;4.0 mm in length; glumes often thin and membranaceous (
<figureCitation id="78D4F7A86B761D4200C5D11EF070FCB8" box="[1000,1070,814,833]" captionStart="Fig" captionStartId="4.[113,138,983,997]" captionTargetBox="[275,1336,143,946]" captionTargetId="figure-478@4.[274,1336,142,947]" captionTargetPageId="4" captionText="Fig.4. Examples of plants from six species of the Schoenus cuspidatus andallies group found in southern Africa,showing the corresponding inflorescences and spikelets beneath each plant: A,G and M—S.crassiculmis; B,H and N—S.quartziticus, C,I andO—S.limosus; D,J,and P—S.purpurascens; E,K andQ—S.prophyllus; and F,L and R—S.calceolus. The black scale bar adjacent to the inflorescences in G—L represents 10 mm, while the scale bars adjacent to the spikelets in M—R represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496604" httpUri="https://zenodo.org/record/10496604/files/figure.png" pageId="2" pageNumber="521">Fig. 4N</figureCitation>
); plants endemic to quartz patches on shale-derived soils in the Overberg region of the
<collectingRegion id="222B25CF6B761D4207DCD17AF187FCA4" box="[1265,1497,842,861]" country="South Africa" name="Western Cape" pageId="2" pageNumber="521">Western Cape Province</collectingRegion>
of
<collectingCountry id="98F8ABBD6B761D420079D155F795FC81" box="[852,971,869,888]" name="South Africa" pageId="2" pageNumber="521">South Africa</collectingCountry>
:…..…...……………………....…....
<emphasis id="D29B373F6B761D4207C2D155F187FC81" bold="true" box="[1263,1497,868,888]" pageId="2" pageNumber="521">
8.
<taxonomicName id="27EF90AE6B761D42062BD154F187FC81" ID-CoL="8XW53" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[1286,1497,868,888]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="quartziticus">
<emphasis id="D29B373F6B761D42062BD154F187FC81" bold="true" box="[1286,1497,868,888]" italics="true" pageId="2" pageNumber="521">Schoenus quartziticus</emphasis>
</taxonomicName>
</emphasis>
9b. Spikelets usually&gt;4.0 mm in length; glumes firm (
<figureCitation id="78D4F7A86B761D4206A9D1B1F190FC6C" box="[1412,1486,897,917]" captionStart="Fig" captionStartId="4.[113,138,983,997]" captionTargetBox="[275,1336,143,946]" captionTargetId="figure-478@4.[274,1336,142,947]" captionTargetPageId="4" captionText="Fig.4. Examples of plants from six species of the Schoenus cuspidatus andallies group found in southern Africa,showing the corresponding inflorescences and spikelets beneath each plant: A,G and M—S.crassiculmis; B,H and N—S.quartziticus, C,I andO—S.limosus; D,J,and P—S.purpurascens; E,K andQ—S.prophyllus; and F,L and R—S.calceolus. The black scale bar adjacent to the inflorescences in G—L represents 10 mm, while the scale bars adjacent to the spikelets in M—R represent 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496604" httpUri="https://zenodo.org/record/10496604/files/figure.png" pageId="2" pageNumber="521">Fig. 4O</figureCitation>
); plants endemic to grasslands and wetlands of the
<collectingRegion id="222B25CF6B761D420603D1AEF1F0FC49" box="[1326,1454,925,945]" country="South Africa" name="Eastern Cape" pageId="2" pageNumber="521">Eastern Cape</collectingRegion>
and
<collectingRegion id="222B25CF6B761D420011D189F795FC35" box="[828,971,953,972]" country="South Africa" name="KwaZulu-Natal" pageId="2" pageNumber="521">KwaZulu-Natal</collectingRegion>
Provinces of
<collectingCountry id="98F8ABBD6B761D42076BD189F09EFC35" box="[1094,1216,953,972]" name="South Africa" pageId="2" pageNumber="521">South Africa</collectingCountry>
):……...….
<emphasis id="D29B373F6B761D420637D189F187FC35" bold="true" box="[1306,1497,952,972]" pageId="2" pageNumber="521">
9.
<taxonomicName id="27EF90AE6B761D42061DD188F187FC35" ID-CoL="8XW52" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[1328,1497,952,972]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="limosus">
<emphasis id="D29B373F6B761D42061DD188F187FC35" bold="true" box="[1328,1497,952,972]" italics="true" pageId="2" pageNumber="521">Schoenus limosus</emphasis>
</taxonomicName>
</emphasis>
5b. Spikelets ovate (
<figureCitation id="78D4F7A86B761D420736D1E5F012FC11" box="[1051,1100,981,1000]" captionStart="Fig" captionStartId="2.[113,138,384,398]" captionTargetBox="[351,1261,142,346]" captionTargetId="figure-573@2.[350,1261,142,349]" captionTargetPageId="2" captionText="Fig. 1. Spikelet variation in the Schoenus cuspidatus and allies group found in southern Africa: A—S. cuspidatus (lanceolate spikelet), B—S. crassus (lanceolate spikelet), C—S. auritus (lanceolate spikelet), D—S. graminifolius (lanceolate spikelet), E—S. calceolus (ovate spikelet) and F—S. ligulatus (linear-lanceolate spikelet).The black scale bar next to the spikelets represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10496598" httpUri="https://zenodo.org/record/10496598/files/figure.png" pageId="2" pageNumber="521">Fig. 1</figureCitation>
): 10a. Sheaths loose; leaves appearing flat as they are usually not-channelled above ligule; plants endemic to mountainous regions of the southern
<collectingRegion id="222B25CF6B761D420725D619F152FBC5" box="[1032,1292,1065,1084]" country="South Africa" name="Western Cape" pageId="2" pageNumber="521">Western Cape Province</collectingRegion>
of
<collectingCountry id="98F8ABBD6B761D420667D619F189FBC5" box="[1354,1495,1065,1084]" name="South Africa" pageId="2" pageNumber="521">South Africa</collectingCountry>
: ……………………………………...….………...
<emphasis id="D29B373F6B761D4207F8D675F187FBA1" bold="true" box="[1237,1497,1091,1112]" pageId="2" pageNumber="521">
10.
<taxonomicName id="27EF90AE6B761D4207DAD673F187FBA1" ID-CoL="8Y9FB" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[1271,1497,1091,1112]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="purpurascens">
<emphasis id="D29B373F6B761D4207DAD673F187FBA1" bold="true" box="[1271,1497,1091,1112]" italics="true" pageId="2" pageNumber="521">Schoenus purpurascens</emphasis>
</taxonomicName>
</emphasis>
10b. Sheaths firm; leaves appearing terete as they are channelled above ligule; plants endemic to the Agulhas Plains region of
<collectingCountry id="98F8ABBD6B761D42068CD64CF729FB52" name="South Africa" pageId="2" pageNumber="521">South Africa</collectingCountry>
: 11a. Spikelets with firm prophylls that often shine; prophyll mucros&gt;3.0 mm; base of culms terete and with sheath not holding several leaf bases together:………………………………….….
<emphasis id="D29B373F6B761D4207D8D6DCF184FB06" bold="true" box="[1269,1498,1259,1279]" pageId="2" pageNumber="521">
11.
<taxonomicName id="27EF90AE6B761D42063BD6DBF184FB06" ID-CoL="8Y9F9" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[1302,1498,1259,1279]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="prophyllus">
<emphasis id="D29B373F6B761D42063BD6DBF184FB06" bold="true" box="[1302,1498,1259,1279]" italics="true" pageId="2" pageNumber="521">Schoenus prophyllus</emphasis>
</taxonomicName>
</emphasis>
11b. Spikelets with prophylls that are not firm and shining; prophyll mucros &lt;
<quantity id="271746C86B761D4200BBD714F781FACE" box="[918,991,1316,1335]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="521" unit="mm" value="2.4">2.4 mm</quantity>
long; base of culm flattened and with sheath holding several leaf bases together (
<figureCitation id="78D4F7A86B761D420710D770F030FAAA" box="[1085,1134,1344,1363]" captionStart="Fig" captionStartId="4.[113,138,1949,1963]" captionTargetBox="[334,564,1306,1913]" captionTargetId="figure-590@4.[333,564,1305,1914]" captionTargetPageId="4" captionText="Fig. 5. Culm base of Schoenus calceolus showing culms grouped together by a translucent leaf sheath.The black scale bar adjacent to the base of the culms represents 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10496606" httpUri="https://zenodo.org/record/10496606/files/figure.png" pageId="2" pageNumber="521">Fig. 5</figureCitation>
):………………..
<emphasis id="D29B373F6B761D420629D770F142FAAA" bold="true" box="[1284,1308,1344,1363]" pageId="2" pageNumber="521">12</emphasis>
.
<taxonomicName id="27EF90AE6B761D420608D70EF184FAAA" ID-CoL="8XW4X" authorityName="T. L. Elliott &amp; Muasya" authorityYear="2019" box="[1317,1498,1342,1363]" class="Liliopsida" family="Cyperaceae" genus="Schoenus" kingdom="Plantae" order="Poales" pageId="2" pageNumber="521" phylum="Tracheophyta" rank="species" species="calceolus">
<emphasis id="D29B373F6B761D420608D70EF184FAAA" bold="true" box="[1317,1498,1342,1363]" italics="true" pageId="2" pageNumber="521">Schoenus calceolus</emphasis>
</taxonomicName>
</paragraph>
</key>
</subSubSection>
</treatment>
</document>