199 lines
22 KiB
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199 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.82.e111148" ID-Pensoft-Pub="1864-8312-82-151" ID-Pensoft-UUID="8D187145BFCA55A1A73960CDE5BD124B" ID-ZooBank="43AC036E93CC4D79939A07DF54BE1A2D" ModsDocID="1864-8312-82-151" checkinTime="1711188240520" checkinUser="pensoft" docAuthor="Brazidec, Manuel, Vilhelmsen, Lars, Boudinot, Brendon E., Richter, Adrian, Hammel, Joerg U., Perkovsky, Evgeny E., Fan, Yong, Wang, Zhen, Wu, Qiong, Wang, Bo & Perrichot, Vincent" docDate="2024" docId="6586EC0085A85D39BE714BC02321F92A" docLanguage="en" docName="ArthropodSystPhyl 82: 151-181" docOrigin="Arthropod Systematics & amp; Phylogeny 82" docPubDate="2024-03-22" docSource="http://dx.doi.org/10.3897/asp.82.e111148" docTitle="Kamyristi exfrigore Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot 2024, sp. nov." docType="treatment" docUuid="2141EC6A-4F85-4ECA-A118-0F3F8CBF1D34" docUuidSource="ZooBank" docVersion="1" id="8D187145BFCA55A1A73960CDE5BD124B" lastPageNumber="151" masterDocId="8D187145BFCA55A1A73960CDE5BD124B" masterDocTitle="Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities" masterLastPageNumber="181" masterPageNumber="151" pageNumber="151" updateTime="1711188240520" updateUser="pensoft">
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<mods:title id="C241F6A8ECDF732BADBBB3A5649D4065">Unveiling ancient diversity of long-tailed wasps (Hymenoptera: Megalyridae): new taxa from Cretaceous Kachin and Taimyr ambers and their phylogenetic affinities</mods:title>
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<mods:namePart id="BD3251EE5DB4B25A2050CB7A4F488FE9">Brazidec, Manuel</mods:namePart>
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<mods:affiliation id="D9163699A9DF1335CB4BFEEB4CBA570B">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France & Institut de Syste ́ matique, E ́ volution, Biodiversite ́ (ISYEB), Muse ́ um national d'Histoire naturelle, CNRS, Sorbonne Universite ́, EPHE, Universite ́ des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France & State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:nameIdentifier id="C1CCA387964352327EB8F4256748AEDB" type="email">manuel.brazidec@gmail.com</mods:nameIdentifier>
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<mods:namePart id="E3A8D737C213D8BD039D918B4D853512">Vilhelmsen, Lars</mods:namePart>
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<mods:affiliation id="D73793E881A107AAC5163A70945A9746">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark</mods:affiliation>
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<mods:namePart id="11EACBBB95ABA5E073D8396D33FD7CA6">Boudinot, Brendon E.</mods:namePart>
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<mods:affiliation id="ADCCABF10C3E4D183E20E63D5B5A6F40">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & National Museum of Natural History, Smithsonian Institution, 10 th & Constitution Ave. NW, Washington, DC, USA & Senckenberg Gesellschaft fuer Naturforschung und Naturmuseum, Frankfurt am Main, 60325 Germany</mods:affiliation>
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<mods:namePart id="0940138A4732F585826C4C2B2895C342">Richter, Adrian</mods:namePart>
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<mods:nameIdentifier id="A25AE796CA0D3D0C5F232B96C701B9A7" type="ORCID">https://orcid.org/0000-0001-5627-2302</mods:nameIdentifier>
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<mods:affiliation id="A769E39CDB6DEEDCD51BFDEC053E8563">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, Vor dem Neutor 1, 07743 Jena, Germany & Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa, Japan</mods:affiliation>
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<mods:namePart id="4ACA126628EC79A8132651C1B06E63E9">Hammel, Joerg U.</mods:namePart>
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<mods:affiliation id="A732C843A437D6F3FD35A9382BD7175F">Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Str. 1, 21502 Geesthacht, Germany</mods:affiliation>
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<mods:namePart id="89959F1A30866D761B0800F70A359228">Perkovsky, Evgeny E.</mods:namePart>
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<mods:nameIdentifier id="FFA6B85B50496CD4947CEF0CEA5786F1" type="ORCID">https://orcid.org/0000-0002-7959-4379</mods:nameIdentifier>
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<mods:affiliation id="69EBBB8295BFE9741F3DB0AD1B163F26">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & I. I. Schmalhausen Institut of Zoology, National Academy of Sciences of Ukraine, B. Khmelnitskogo 15, Kiev 01030, Ukraine</mods:affiliation>
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<mods:namePart id="7D7C051299D37F479A1EF50DF545B433">Fan, Yong</mods:namePart>
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<mods:affiliation id="5CE5C77296E2F7B811519DAE592E336F">Fushun Amber Institute, Fushun 113005, China</mods:affiliation>
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<mods:namePart id="DC5501BFDC9A06F26B1BA5F880E8EF6D">Wang, Zhen</mods:namePart>
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<mods:affiliation id="807D0A917489634C21F59D419005BCFA">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="A971A670FF67916692E3614BD64675D1">Wu, Qiong</mods:namePart>
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<mods:affiliation id="6A8CB9AFD57DED6E37C3CE3939830F75">Natural History Museum of Denmark, SCIENCE, University of Copenhagen, Universitetsparken 15, DK- 2100, Denmark & College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China</mods:affiliation>
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<mods:namePart id="4F86D76E020A5414DA2CAE0C92E1771E">Wang, Bo</mods:namePart>
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<mods:affiliation id="4DAD2A698E775E2A703419BCD432BE26">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
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<mods:namePart id="D092377ADA5ECA821B2F5F4544FC4C7C">Perrichot, Vincent</mods:namePart>
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<mods:nameIdentifier id="B0B1B78F11E7B31DA29B14912C6F1674" type="ORCID">https://orcid.org/0000-0002-7973-0430</mods:nameIdentifier>
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<mods:affiliation id="F3A908B2384C04E1ADCF147F772742F6">Univ Rennes, CNRS, Geosciences Rennes, UMR 6118, 35000, Rennes, France</mods:affiliation>
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<mods:typeOfResource id="F49D08784AA376333856E192B98DB0D3">text</mods:typeOfResource>
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<mods:title id="756C259F4EEC142A9290C13E79135614">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part id="35DE212376DC98260132A5B303C15704">
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<mods:date id="B94C5B8FCE5C33C6070AEC06B40EC0FB">2024</mods:date>
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<mods:number id="CF580886862842DA950A59E4E5EA14C6">2024-03-22</mods:number>
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<mods:detail id="890374EDEBE20C7B46A5367E63888E4A" type="volume">
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<mods:number id="D0BEBF49EA78FFF07712023362773338">82</mods:number>
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<mods:start id="22F35E7D16708B616AFAA0A7AB594417">151</mods:start>
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<mods:url id="8B24220A3F9101A1408F9EF869FBEDB5">http://dx.doi.org/10.3897/asp.82.e111148</mods:url>
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<mods:identifier id="9F2E47AA6D3D5703D5ADAEE44D102C5F" type="DOI">http://dx.doi.org/10.3897/asp.82.e111148</mods:identifier>
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<treatment id="6586EC0085A85D39BE714BC02321F92A" LSID="urn:lsid:zoobank.org:act:2141EC6A-4F85-4ECA-A118-0F3F8CBF1D34" httpUri="http://treatment.plazi.org/id/6586EC0085A85D39BE714BC02321F92A" lastPageNumber="151" pageId="0" pageNumber="151">
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<subSubSection id="18127FE0407072497E94B183664C768A" pageId="0" pageNumber="151" type="nomenclature">
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<paragraph id="EF81BC6455D5FCDF7E86FF988E6F7AAD" pageId="0" pageNumber="151">
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3.2.1.1.1. †
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<taxonomicName id="7CD3F3DEE643DB495ECF26908DA2DAB7" LSID="https://zoobank.org/2141EC6A-4F85-4ECA-A118-0F3F8CBF1D34" authority="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot, 2024" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Kamyristi" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Kamyristi exfrigore" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="exfrigore" status="sp. nov.">Kamyristi exfrigore</taxonomicName>
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<taxonomicNameLabel id="519525CA547672F473BD2FBF2C11B5E0" pageId="0" pageNumber="151">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="06B27F097E7C6100F71EDDBE74078A87" pageId="0" pageNumber="151" type="description">
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<paragraph id="0C6110BBA2B0B5A4409F3D55A45436F6" pageId="0" pageNumber="151">
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<figureCitation id="A3801934A5F11B61B9CC35167B2D4265" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of fore wing venation for the taxa described in the present paper. A † Cretolyra noijebumensis gen. et sp. nov.; B † Cretolyra shawi gen. et sp. nov.; C † Genkyhag innebula gen. et sp. nov.; D † Megacoxa chandrahrasa gen. et sp. nov.; E † Megacoxa janzeni gen. et sp. nov.; F † Megacoxa synchrotron gen. et sp. nov.; G † Kamyristi exfrigore gen. et sp. nov.; H † Kamyristi yantardakhensis gen. et sp. nov. Scale bars: 0.5 mm (A - H)." figureDoi="10.3897/asp.82.e111148.figure2" httpUri="https://binary.pensoft.net/fig/1007131" pageId="0" pageNumber="151">Figures 2G</figureCitation>
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<figureCitation id="B1F6DBA7F9F0EBC6943D07611A6F646F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. † Kamyristi gen. nov. A - C † Kamyristi exfrigore gen. et sp. nov., holotype female PIN 3730 / 411, Albian-Cenomanian, Taimyr amber. A habitus in lateral view; B habitus in dorsal view; C head and mesosoma in lateral view; D - F † Kamyristi yantardakhensis gen. et sp. nov., holotype male 3311 / 2718, Santonian, Taimyr amber; D habitus in lateral view; E habitus in ventral view; E habitus in dorsal view. Scale bars: 0.5 mm (A, B, D, E); 0.25 mm (C). AG = inner axillar groove; ce = cercus; MMS = median mesoscutal sulcus; OC = occipital carina; OvS = ovipositor sheaths; pa = parameres; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure9" httpUri="https://binary.pensoft.net/fig/1007138" pageId="0" pageNumber="151">, 9A-C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="200288D3DF1B63ED6AF0BFF1D7FF0B5F" pageId="0" pageNumber="151" type="etymology">
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<paragraph id="C0FB0E006AFC7A61BFFF0B4FD03E7C99" pageId="0" pageNumber="151">Etymology.</paragraph>
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<paragraph id="3655FDD95B9D69654E40F2DA7EA693DD" pageId="0" pageNumber="151">
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"
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<emphasis id="A956F03E2671E989A2FA238E51F67ACE" italics="true" pageId="0" pageNumber="151">exfrigore</emphasis>
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" means "from the cold" in Latin as this species is the northernmost record of a megalyrid, extant and extinct, in a region where the temperature is currently below zero most of the year. The specific epithet is to be treated as an adjective.
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</paragraph>
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</subSubSection>
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<subSubSection id="67ECA4F23C0356A3974A2E3B5D1D2CB7" pageId="0" pageNumber="151" type="materials_examined">
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<paragraph id="E6EA122027F0AB1FBEC24EBD64AE381C" pageId="0" pageNumber="151">Material studied.</paragraph>
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<paragraph id="E44AFAD8C72F02EC5CE1AEF7FED508F9" pageId="0" pageNumber="151">
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<materialsCitation id="B81364BFD36A7105AA57E32B9317FA21" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" location="Paleontological Institute" municipality="Russian Academy of Sciences" specimenCount="1" typeStatus="Holotype">
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<emphasis id="643E44EB68AD8474375D29A37A18C7C8" bold="true" italics="true" pageId="0" pageNumber="151">
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<typeStatus id="1A620EA3D9F802F67C52577C7EC7B55A" type="holotype">Holotype</typeStatus>
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</emphasis>
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female PIN 3730/411; housed in the collection of the
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<location id="2F9ECE2C62FB4E5525C481E8679942E6" LSID="urn:lsid:plazi:treatment:6586EC0085A85D39BE714BC02321F92A:2F9ECE2C62FB4E5525C481E8679942E6" municipality="Russian Academy of Sciences" name="Paleontological Institute">Paleontological Institute</location>
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of the
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<collectingMunicipality id="790E3A7B3B03FB6D38C87E2654D54937">Russian Academy of Sciences</collectingMunicipality>
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(PIN).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="EEA041CFB198AE28A0BC967BA22CE718" pageId="0" pageNumber="151" type="type locality">
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<paragraph id="34AD167316F5FC88FFF66D74FF764CB0" pageId="0" pageNumber="151">Type locality.</paragraph>
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<paragraph id="2F04EFB989EDE295593406EC7B7230F2" pageId="0" pageNumber="151">Baikura, Russia.</paragraph>
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</subSubSection>
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<subSubSection id="4461F487047EC0DFA81B019EAEAE8F96" pageId="0" pageNumber="151" type="age">
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<paragraph id="E0B00DA6232427D8B9FE7DBF12FBB248" pageId="0" pageNumber="151">Age.</paragraph>
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<paragraph id="01CB47F90ECE0992E559A0970DE32CE9" pageId="0" pageNumber="151">Upper Albian to lower Cenomanian, mid-Cretaceous.</paragraph>
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</subSubSection>
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<subSubSection id="3E9F465C63436267A6BC9D3EE4F2BC63" pageId="0" pageNumber="151" type="diagnosis">
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<paragraph id="2DBD77EFBA581637073E57C4285FA773" pageId="0" pageNumber="151">Diagnosis.</paragraph>
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<paragraph id="E0B5F29F6E9AFD3A4500574E6ED511F4" pageId="0" pageNumber="151">
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Eye nearly round (vs. oval, higher than long in †
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<taxonomicName id="81B20EDB51E0348FE4D7143D0458CC0A" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Kamyristi" kingdom="Animalia" lsidName="Kamyristi yantardakhensis" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="yantardakhensis">
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<emphasis id="C1C525241A4CC0A5125F02EB134C6C97" italics="true" pageId="0" pageNumber="151">Kamyristi yantardakhensis</emphasis>
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</taxonomicName>
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<emphasis id="A0638C8C17DAA51888CE3E507002AAD1" bold="true" pageId="0" pageNumber="151">gen. et sp. nov.</emphasis>
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); axillae contiguous medially with axillar groove smooth (Fig.
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<figureCitation id="62A54D3A84815A95903C97AF67BB0E0A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. † Kamyristi gen. nov. A - C † Kamyristi exfrigore gen. et sp. nov., holotype female PIN 3730 / 411, Albian-Cenomanian, Taimyr amber. A habitus in lateral view; B habitus in dorsal view; C head and mesosoma in lateral view; D - F † Kamyristi yantardakhensis gen. et sp. nov., holotype male 3311 / 2718, Santonian, Taimyr amber; D habitus in lateral view; E habitus in ventral view; E habitus in dorsal view. Scale bars: 0.5 mm (A, B, D, E); 0.25 mm (C). AG = inner axillar groove; ce = cercus; MMS = median mesoscutal sulcus; OC = occipital carina; OvS = ovipositor sheaths; pa = parameres; sp = anterior thoracic spiracle." figureDoi="10.3897/asp.82.e111148.figure9" httpUri="https://binary.pensoft.net/fig/1007138" pageId="0" pageNumber="151">9B</figureCitation>
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; vs. separated medially with axillar groove crenulated in †
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<taxonomicName id="AA9931AA8BC720EB5F6EBDD9C0C995B7" authorityName="Brazidec & Vilhelmsen & Boudinot & Richter & Hammel & Perkovsky & Fan & Wang & Wu & Wang & Perrichot" authorityYear="2024" family="Megalyridae" genus="Kamyristi" kingdom="Animalia" lsidName="Kamyristi yantardakhensis" order="Hymenoptera" pageId="0" pageNumber="151" rank="species" species="yantardakhensis">
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<emphasis id="DB58FB25F81945B17BCDB0193440FEB8" italics="true" pageId="0" pageNumber="151">Kamyristi yantardakhensis</emphasis>
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</taxonomicName>
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<emphasis id="550797C5980E47CD414A9300671C1715" bold="true" pageId="0" pageNumber="151">gen. et sp. nov.</emphasis>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="BDEFBCFD48A093AEAF255D3BFBA075C5" pageId="0" pageNumber="151" type="description">
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<paragraph id="46C5CA3372CD3EFF4BED199DA6A9E99E" pageId="0" pageNumber="151">Description.</paragraph>
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<paragraph id="35F58E9947A97819271EF83A5C8BF3BE" pageId="0" pageNumber="151">
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Body length 2.03 mm. Almost complete, except ovipositor. -
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<emphasis id="8252EE9F026D18A5B34EF62BE640AE74" bold="true" italics="true" pageId="0" pageNumber="151">Head</emphasis>
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globular, higher than long (length 0.38 mm; height 0.43); slightly pubescent on vertex and frons; frons convex, shagreened; compound eye rounded, slightly higher than long, not covering head length, postocular carina absent; vertex convex, shagreened; torulus inserted very close to ventral margin of eye; subantennal groove present; mandible with three teeth; scape short and thick (length 0.09 mm); pedicel same length; flagellomeres cylindrical, longer than wide, elongate (length ca. 0.08 mm); flagellomere 12 longest; occipital carina crenulate. -
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<emphasis id="48B19D9F0D68B616A4E3CF37881BBB03" bold="true" italics="true" pageId="0" pageNumber="151">Mesosoma</emphasis>
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shorter than metasoma (length 0.69 mm; height 0.40 mm), almost fully shagreened; mesoscutum shagreened, short (length 0.20 mm; width 0.47 mm), one third of mesosoma length, divided by crenulate median mesoscutal sulcus, with sparse erect setae; axillae continuous medially, with inner axillar groove smooth; mesoscutellum as long as mesoscutum, posterior margin slightly outcurved; pronotum not visible dorsally; anterior thoracic spiracle fully surrounded by pronotal cuticle; propodeum areolate-rugose. -
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<emphasis id="C28846FAC12B4DF4DD8F2335B276908C" bold="true" italics="true" pageId="0" pageNumber="151">Fore wing</emphasis>
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hyaline (length ca. 1.50 mm) and covered with microtrichia; C, Sc+R, Rs +M, basal segments of Rs and M; M+Cu, Cu and 1m-cu spectral; Rs not closing first submarginal cell and pigmented for two thirds between r-rs and Rs+M; M pigmented to level of 2r-m and then spectral to wing tip; pterostigma broad; R1 extending beyond marginal cell; Rs pigmented, arched toward stigma, closing marginal cell. -
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<emphasis id="C8343579B9C753C7D0B67258055F8DCE" bold="true" italics="true" pageId="0" pageNumber="151">Legs</emphasis>
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bearing many scattered erect setae; one mesotibial spur, one metatibial spur present; metafemur swollen, metatibia notched before spur; metabasitarsus three times longer than following, remainder same length. -
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<emphasis id="212C72B59C8CC2B3F6D3D39CADB2A10C" bold="true" italics="true" pageId="0" pageNumber="151">Metasoma</emphasis>
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almost half body length (length 0.96 mm), elongate, smooth; segments subequal in length; hypopygium well-developed, concave; ovipositor incomplete, covered by sheaths only at base; cerci triangular, exserted.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |