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<document ID-DOI="10.5281/zenodo.439745" ID-GBIF-Dataset="e7d164f6-b22c-4120-b1f6-6464cfc6de0b" ID-GBIF-Taxon="128090364" ID-ISSN="0005 6219" ID-Zenodo-Dep="439745" checkinTime="1488976263354" checkinUser="plazi" docAuthor="Ruud, A. Bank, Henk P. M. G Menkhorst &amp; Eike Neubert" docDate="2016" docId="29708788FFCDFFDEFF32FC8CFAE9FD6F" docLanguage="en" docName="basteria.80.1-3.5.2016 Bank et al Lauriidae Enidae Vitrinidae Turkey.pdf" docOrigin="Basteria 80 (1)" docStyle="DocumentStyle{}" docTitle="Imparietula inflexa Ruud, Menkhorst &amp; Neubert, 2016, spec. nov." docType="treatment" docVersion="9" lastPageNumber="14" masterDocId="D549FFF0FFC6FFD5FFDDFFA1FFAEFFF7" masterDocTitle="Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)" masterLastPageNumber="30" masterPageNumber="5" pageId="11" pageNumber="14" updateTime="1636507312436" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ruud, A. Bank</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Henk P. M. G Menkhorst</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eike Neubert</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Basteria</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>5</mods:start>
<mods:end>30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.439745</mods:identifier>
<mods:identifier type="GBIF-Dataset">e7d164f6-b22c-4120-b1f6-6464cfc6de0b</mods:identifier>
<mods:identifier type="ISSN">00056219</mods:identifier>
<mods:identifier type="Zenodo-Dep">439745</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="128090364" LSID="urn:lsid:plazi:treatment:29708788FFCDFFDEFF32FC8CFAE9FD6F" httpUri="http://treatment.plazi.org/id/29708788FFCDFFDEFF32FC8CFAE9FD6F" lastPageNumber="14" pageId="11" pageNumber="14">
<subSubSection box="[239,675,813,838]" pageId="11" pageNumber="14" type="nomenclature">
<paragraph blockId="11.[239,675,813,838]" box="[239,675,813,838]" pageId="11" pageNumber="14">
<heading bold="true" box="[239,675,813,838]" centered="true" fontSize="10" level="4" pageId="11" pageNumber="14" reason="0">
<emphasis bold="true" box="[239,675,813,838]" pageId="11" pageNumber="14">
<taxonomicName box="[239,457,813,837]" class="Gastropoda" family="Enidae" genus="Imparietula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="inflexa" status="spec. nov.">
<emphasis bold="true" box="[239,457,813,837]" italics="true" pageId="11" pageNumber="14">Imparietula inflexa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[464,577,814,838]" pageId="11" pageNumber="14" rank="species">spec. nov.</taxonomicNameLabel>
(
<figureCitation box="[591,669,814,838]" captionStart="Figs 16 - 20" captionStartId="21.[151,191,1858,1877]" captionTargetBox="[151,1436,155,1833]" captionTargetId="figure@21.[151,1436,155,1833]" captionTargetPageId="21" captionText="Figs 16 - 20. Multidentula and Imparietula taxa from Turkey. 16, Multidentula squalina squalina (L. Pfeiffer, 1848), NMBE 548313 (H = 7.8 mm), Turkey, Vil. Karabük, eastern part Eskipazar, leg. Hausdorf, 27. ix. 1987. 17, Multidentula squalina eudoxina (Nägele, 1894), paralectotype SMF 14195 a coll. Kobelt ex Nägele (H = 6.3 mm), Turkey, Vil. Tokat, Tokat. 18, Multidentula squalina acutior (Lindholm, 1922), NMBE 544678 (H = 5.7 mm), Turkey, Vil. Artvin, N. Kınalıçam (= NNW. Cağliyan). 19, Multidentula reducta spec. nov., holotype NMBE 544681 (H = 6.2 mm), Turkey, Vil. Sivas, Zara, 16 km towards Imranlı. 20, Imparietula inflexa spec. nov., holotype NMBE 544683 (H = 12, 4 mm), Turkey, Vil. Giresun, 5 km N. Tamdere (= S. Yavuzkemal). All photographs Bochud &amp; Neubert, × 8." httpUri="https://zenodo.org/record/439752/files/figure.png" pageId="11" pageNumber="14">Fig. 20</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="14" type="materials_examined">
<paragraph blockId="11.[151,756,887,976]" pageId="11" pageNumber="14">
Type locality &amp; type specimens.
<materialsCitation ID-GBIF-Occurrence="1455386813" collectingDate="1992-08-06" collectionCode="NMBE" collectorName="H. P. M. G. Menkhorst" country="Turkey" elevation="1430" latitude="40.5397" location="5 km N. Tamdere (= S. Yavuzkemal)" longLatPrecision="6" longitude="38.3575" pageId="11" pageNumber="14" specimenCode="544683" specimenCount="1" stateProvince="Vilayet Giresun" typeStatus="holotype">
<collectingCountry box="[473,540,887,907]" name="Turkey" pageId="11" pageNumber="14">Turkey</collectingCountry>
,
<collectingRegion box="[550,697,887,907]" pageId="11" pageNumber="14">Vilayet Giresun</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:29708788FFCDFFDEFF32FC8CFAE9FD6F:A4066045FFCDFFDEFD18FCD6FE1AFC59" country="Turkey" latitude="40.5397" longLatPrecision="6" longitude="38.3575" name="5 km N. Tamdere (= S. Yavuzkemal)" pageId="11" pageNumber="14" stateProvince="Vilayet Giresun">
<quantity box="[709,756,887,906]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="14" unit="km" value="5.0">5 km</quantity>
N. Tamdere (= S. Yavuzkemal)
</location>
,
<quantity box="[446,513,922,941]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="11" pageNumber="14" unit="m" value="1430.0">
<elevation box="[446,513,922,941]" metricMagnitude="3" metricUnit="m" metricValue="1.43" pageId="11" pageNumber="14" unit="m" value="1430.0">1430 m</elevation>
</quantity>
(
<geoCoordinate box="[525,621,922,941]" direction="north" orientation="latitude" pageId="11" pageNumber="14" precision="5" value="40.5397">40.5397°N</geoCoordinate>
<geoCoordinate box="[627,719,922,942]" direction="east" orientation="longitude" pageId="11" pageNumber="14" precision="5" value="38.3575">38.3575°E</geoCoordinate>
),
<paragraph blockId="11.[151,756,887,976]" box="[151,737,956,976]" pageId="11" pageNumber="14">
<collectorName box="[151,348,956,976]" pageId="11" pageNumber="14">H.P.M.G. Menkhorst</collectorName>
leg.,
<date box="[397,495,956,975]" pageId="11" pageNumber="14" value="1992-08-06">
<collectingDate box="[397,495,956,975]" pageId="11" pageNumber="14" value="1992-08-06">6.viii.1992</collectingDate>
</date>
.
<typeStatus box="[503,592,956,975]" pageId="11" pageNumber="14">Holotype</typeStatus>
<collectionCode box="[597,661,957,976]" collectionName="Switzerland, Bern, Naturhistorische Museums" pageId="11" pageNumber="14">NMBE</collectionCode>
<specimenCode box="[667,734,956,975]" pageId="11" pageNumber="14">544683</specimenCode>
.
</paragraph>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="14" type="diagnosis">
<paragraph blockId="11.[151,764,1022,2051]" pageId="11" pageNumber="14">
Diagnosis. A small, hornbrown coloured
<taxonomicName box="[631,751,1022,1046]" class="Gastropoda" family="Enidae" genus="Imparietula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis box="[631,751,1022,1046]" italics="true" pageId="11" pageNumber="14">Imparietula</emphasis>
</taxonomicName>
species with a granulated sculpture, a strong
<taxonomicName box="[658,752,1057,1081]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior and a biarcuate last whorl.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="14" type="description">
<paragraph blockId="11.[151,764,1022,2051]" pageId="11" pageNumber="14">
Description. Shell dextral, spindle-shaped in outline, with an open, slit-like umbilicus. The 7.9-8.1 whorls are convex with a rather deep suture. Teleoconch with irregular, dense, rather pronounced, oblique striae crossed by irregular, indistinct, incised spiral striae producing a granulated sculpture. Shell fairly solid, not translucent, dull, hornbrown coloured, with a whitish band behind the peristome. Peristome reflected, thickened by a labial callus, the columellar and palatal insertion connected by a thin callus, which is thickened near its ends. The thickening of the parietal callus at the palatal insertion of the peristome resembles
<taxonomicName box="[247,414,1542,1566]" class="Gastropoda" family="Vitrinidae" genus="Amphiscopus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="subangularis">a subangularis</taxonomicName>
. A prominent
<taxonomicName box="[574,668,1542,1566]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior is present. The last whorl is biarcuate, being strongly looped inward the middle; there is a prominent impression at the
<taxonomicName box="[407,501,1646,1670]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior. The columellar ledge reaches halfway of the columellar side of the aperture. There is an additional thickening running from the insertion of the columellar ledge to the basal part of the peristome.
</paragraph>
<paragraph blockId="11.[151,764,1022,2051]" pageId="11" pageNumber="14">Measurements (n = 3). H = 12.3-12.8 (mean 12.6); LWH = 5.9-6.5 (mean 6.3); MH = 3.8-4.3 (mean 4.0); LWD = 4.3-4.7 (mean 4.4); LWM = 4.2-4.6 (mean 4.4); MD = 2.9-3.1 (mean 3.0); NW = 7.9-8.1 (mean 8.0).</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="14" type="distribution">
<paragraph blockId="11.[151,764,1022,2051]" pageId="11" pageNumber="14">Localities. Vilayet Giresun: type locality (see above); 6 km N. Tamdere (HMK/1); Giresun, 75 km towards Şebinkarahisar, 1500 m (NMBE/1).</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="14" type="etymology">
<paragraph blockId="11.[824,1434,155,664]" pageId="11" pageNumber="14">
Derivatio nominis. Named after the inwards curved aperture near the
<taxonomicName box="[1023,1117,190,214]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="14" type="diagnosis">
<paragraph blockId="11.[824,1434,155,664]" pageId="11" pageNumber="14">
Differentiation.
<taxonomicName box="[1059,1149,224,248]" class="Gastropoda" family="Enidae" genus="Imparietula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="inflexa">
<emphasis box="[1059,1149,224,248]" italics="true" pageId="11" pageNumber="14">I. inflexa</emphasis>
</taxonomicName>
differs from
<taxonomicName box="[1295,1389,224,248]" class="Gastropoda" family="Enidae" genus="Imparietula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="pelidne">
<emphasis box="[1295,1389,224,248]" italics="true" pageId="11" pageNumber="14">I. pelidne</emphasis>
</taxonomicName>
by its dimensions, the stronger
<taxonomicName box="[1140,1234,259,283]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior, the biarcuate last whorl, and the additional thickening inside the aperture at the columellar-basal margin.
<taxonomicName authority="Scopoli, 1777" authorityName="Scopoli" authorityYear="1777" class="Gastropoda" family="Bulimulidae" genus="Bulimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="leucodon">
<emphasis italics="true" pageId="11" pageNumber="14">I. leucodon</emphasis>
</taxonomicName>
also has a prominent
<taxonomicName box="[1129,1223,363,387]" class="Gastropoda" family="Enidae" genus="Clausilioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="palatalis">palatalis</taxonomicName>
superior, but it differs from
<taxonomicName box="[964,1054,398,422]" class="Gastropoda" family="Enidae" genus="Imparietula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="14" phylum="Mollusca" rank="species" species="inflexa">
<emphasis box="[964,1054,398,422]" italics="true" pageId="11" pageNumber="14">I. inflexa</emphasis>
</taxonomicName>
by its dimensions, the less biarcuate last whorl, the less pronounced oblique striae and the less pronounced granulation, the much stronger developed parietal callus, and the absence of an additional thickening in the aperture at the columellar-basal margin.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="14" type="discussion">
<paragraph blockId="11.[824,1434,155,664]" pageId="11" pageNumber="14">Remarks. The description is based on only three specimens, none of them being well-preserved.</paragraph>
</subSubSection>
</treatment>
</document>