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<document id="2AEF0EAFA7D19031AA23322B6F320AAB" ID="10.11646/zootaxa.4175.3.5" ID-DOI="10.11646/zootaxa.4175.3.5" ID-GBIF-Dataset="bc98e8c5-0ebd-46e7-b105-635a9b47c397" ID-ISSN="1175-5326" ID-Zenodo-Dep="265766" ID-ZooBank="18C4A40C-C587-4E67-A769-FEBAD92EBCC8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476429581541" checkinUser="plazi" docAuthor="Khaustov, Alexander A. &amp; Sazhnev, Alexey S." docDate="2016" docId="03EB87B3FFCD115FFF76FE88CB1EF8B8" docLanguage="en" docName="zootaxa.4175.3.5.pdf" docOrigin="Zootaxa 4175 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protoallopygmephorus Khaustov &amp; Sazhnev, 2016, gen. nov." docType="treatment" docVersion="7" lastPageNumber="262" masterDocId="FFD2FFCBFFCC115EFFE1FF92CF6FFF95" masterDocTitle="Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan" masterLastPageNumber="273" masterPageNumber="261" pageNumber="262" updateTime="1698389850606" updateUser="plazi">
<mods:mods id="101D6CA58C6AEFBF7A0C9045154C411A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7F1A60AEFE5AEA663D2D1BE707C5522C">
<mods:title id="35BF01B97714692BFCBF0F3D33DA77FD">Mites of the families Neopygmephoridae and Scutacaridae associated with variegated mud-loving beetles (Coleoptera: Heteroceridae) from Russia and Kazakhstan</mods:title>
</mods:titleInfo>
<mods:name id="BCE64D41BE7EA1674E45914EA0A53BDD" type="personal">
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<mods:namePart id="3474EBBBC53DD4BC1C077F818FB36746">Khaustov, Alexander A.</mods:namePart>
</mods:name>
<mods:name id="85A5BB7607FDA7317F7E24EAD4DED9E4" type="personal">
<mods:role id="CAE8105F8156B3B6281D78E6D34E4A44">
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<mods:namePart id="2802F73D8114C965CE749019E1E9265F">Sazhnev, Alexey S.</mods:namePart>
</mods:name>
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<mods:title id="F0FBD7AD8BA826D7EA97EF033A72912D">Zootaxa</mods:title>
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<mods:part id="B6671316872264422D204A1970CD32B9">
<mods:date id="E5B05727AB61C8B8A630B30330147D6A">2016</mods:date>
<mods:detail id="5703D0294E59EF5813052B2851409A08" type="volume">
<mods:number id="174899463D52B86A7DD4DC8F09115A9B">4175</mods:number>
</mods:detail>
<mods:detail id="DE977990A5352C4A41CB60F50CEDFDE0" type="issue">
<mods:number id="AC2B39CAB902F3F5F93800A736223074">3</mods:number>
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<mods:classification id="236407E99BABDB83B00121963FEB1E4D">journal article</mods:classification>
<mods:identifier id="92EC6C1166C810162B5B5D0268C6D6D7" type="DOI">10.11646/zootaxa.4175.3.5</mods:identifier>
<mods:identifier id="9C86880A256ADC8FF7822264E00CE9F2" type="GBIF-Dataset">bc98e8c5-0ebd-46e7-b105-635a9b47c397</mods:identifier>
<mods:identifier id="E9AE301FA01AE704AFF4EC29477461B2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8BE5855B854A63CEB673247ED68CEE35" type="Zenodo-Dep">265766</mods:identifier>
<mods:identifier id="E343E6561D89A3DFFF4F19756BFBF7E7" type="ZooBank">18C4A40C-C587-4E67-A769-FEBAD92EBCC8</mods:identifier>
</mods:mods>
<treatment id="03EB87B3FFCD115FFF76FE88CB1EF8B8" ID-DOI="http://doi.org/10.5281/zenodo.6087522" ID-GBIF-Taxon="124996769" ID-Zenodo-Dep="6087522" LSID="urn:lsid:plazi:treatment:03EB87B3FFCD115FFF76FE88CB1EF8B8" httpUri="http://treatment.plazi.org/id/03EB87B3FFCD115FFF76FE88CB1EF8B8" lastPageNumber="262" pageId="1" pageNumber="262">
<subSubSection id="C358652EFFCD115FFF76FE88CD19FEA1" box="[151,630,282,308]" pageId="1" pageNumber="262" type="nomenclature">
<paragraph id="8BFD36A5FFCD115FFF76FE88CD19FEA1" blockId="1.[151,630,282,308]" box="[151,630,282,308]" pageId="1" pageNumber="262">
<heading id="D0B581C9FFCD115FFF76FE88CD19FEA1" bold="true" box="[151,630,282,308]" fontSize="11" level="1" pageId="1" pageNumber="262" reason="1">
<emphasis id="B936EAB7FFCD115FFF76FE88CD19FEA1" bold="true" box="[151,630,282,308]" pageId="1" pageNumber="262">
Genus
<taxonomicName id="4C424D26FFCD115FFF0FFE88CD6EFEA1" box="[238,513,282,308]" class="Insecta" family="Heteroceridae" genus="Protoallopygmephorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B936EAB7FFCD115FFF0FFE88CD6EFEA1" bold="true" box="[238,513,282,308]" italics="true" pageId="1" pageNumber="262">Protoallopygmephorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557CCFFCD115FFDE9FE88CD19FEA1" box="[520,630,282,308]" pageId="1" pageNumber="262" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C358652EFFCD115FFF76FEF0CDBDFEE2" box="[151,722,353,375]" pageId="1" pageNumber="262" type="materials_examined">
<paragraph id="8BFD36A5FFCD115FFF76FEF0CDBDFEE2" blockId="1.[151,722,353,375]" box="[151,722,353,375]" pageId="1" pageNumber="262">
Type species:
<taxonomicName id="4C424D26FFCD115FFEC7FEF3CD13FEE2" box="[294,636,353,375]" class="Insecta" family="Heteroceridae" genus="Protoallopygmephorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="heteroceri" status="sp. nov.">
<emphasis id="B936EAB7FFCD115FFEC7FEF3CD13FEE2" box="[294,636,353,375]" italics="true" pageId="1" pageNumber="262">Protoallopygmephorus heteroceri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557CCFFCD115FFD63FEF3CDBDFEE2" box="[642,722,353,375]" pageId="1" pageNumber="262" rank="species">
<emphasis id="B936EAB7FFCD115FFD63FEF3CDA3FEE3" bold="true" box="[642,716,353,374]" pageId="1" pageNumber="262">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C358652EFFCD115FFF76FE36CFA8F908" pageId="1" pageNumber="262" type="diagnosis">
<paragraph id="8BFD36A5FFCD115FFF76FE36CD2CFA9C" blockId="1.[151,1436,420,1838]" pageId="1" pageNumber="262">
<emphasis id="B936EAB7FFCD115FFF76FE36CE65FE28" bold="true" box="[151,266,420,445]" pageId="1" pageNumber="262">Diagnosis</emphasis>
. FEMALE. Body oval. Only the posterior part of the prodorsum covered by tergite C. Gnathosomal capsule oval, its length about equal to its width, dorsally with two pairs of cheliceral setae (
<emphasis id="B936EAB7FFCD115FFB6DFE5ACBDBFE75" box="[1164,1204,456,480]" italics="true" pageId="1" pageNumber="262">cha</emphasis>
,
<emphasis id="B936EAB7FFCD115FFB23FE5ACB85FE75" box="[1218,1258,456,480]" italics="true" pageId="1" pageNumber="262">chb</emphasis>
) and postpalpal setae situated anteriorly to setae
<emphasis id="B936EAB7FFCD115FFDE7FE7ECD41FD91" box="[518,558,492,516]" italics="true" pageId="1" pageNumber="262">cha</emphasis>
; palps prominent, with two pairs of dorsolateral setae (
<emphasis id="B936EAB7FFCD115FFB41FE7ECBA3FD91" box="[1184,1228,492,516]" italics="true" pageId="1" pageNumber="262">dFe</emphasis>
,
<emphasis id="B936EAB7FFCD115FFB3BFE7ECA67FD91" box="[1242,1288,492,516]" italics="true" pageId="1" pageNumber="262">dGe</emphasis>
) and distinct tibial claw distally. Subcapitulum with one pair of setae
<emphasis id="B936EAB7FFCD115FFCC4FD83CC56FDBD" box="[805,825,529,552]" italics="true" pageId="1" pageNumber="262">m</emphasis>
and one pair of oval pits in the posterior half. Palps ventrally with mushroom-like accessory setigenous structure and small solenidion. Prodorsum with two pair of simple setae (
<date id="FFFC1065FFCD115FFED5FDCBCE26FDE1" box="[308,329,601,628]" pageId="1" pageNumber="262">
<emphasis id="B936EAB7FFCD115FFED5FDCBCE2FFDE5" box="[308,320,601,624]" italics="true" pageId="1" pageNumber="262">v</emphasis>
2
</date>
,
<emphasis id="B936EAB7FFCD115FFEB9FDCBCE00FDE5" box="[344,367,601,624]" italics="true" pageId="1" pageNumber="262">sc</emphasis>
2), one pair of clavate trichobothria (
<emphasis id="B936EAB7FFCD115FFCF1FDCBCC48FDE5" box="[784,807,601,624]" italics="true" pageId="1" pageNumber="262">sc</emphasis>
1), and circular stigmata. Dorsal hysterosomal setae not modified. Tergite C with two pairs of setae (
<emphasis id="B936EAB7FFCD115FFD60FD12CDE2FD02" box="[641,653,640,663]" italics="true" pageId="1" pageNumber="262">c</emphasis>
1,
<emphasis id="B936EAB7FFCD115FFD42FD12CDC0FD02" box="[675,687,640,663]" italics="true" pageId="1" pageNumber="262">c</emphasis>
2); tergite D with one pair of setae (
<emphasis id="B936EAB7FFCD115FFBD5FDEDCB2DFD02" box="[1076,1090,639,663]" italics="true" pageId="1" pageNumber="262">d</emphasis>
) and one pair of round cupules
<emphasis id="B936EAB7FFCD115FFF76FD34CFC2FD28" box="[151,173,678,701]" italics="true" pageId="1" pageNumber="262">ia</emphasis>
; tergite EF with two pairs of setae (
<emphasis id="B936EAB7FFCD115FFDDBFD34CD29FD28" box="[570,582,678,701]" italics="true" pageId="1" pageNumber="262">e</emphasis>
,
<emphasis id="B936EAB7FFCD115FFDB5FD34CD33FD2B" box="[596,604,678,702]" italics="true" pageId="1" pageNumber="262">f</emphasis>
); tergite H with two pairs of setae (
<emphasis id="B936EAB7FFCD115FFC07FD37CC9BFD28" box="[998,1012,677,701]" italics="true" pageId="1" pageNumber="262">h</emphasis>
1,
<emphasis id="B936EAB7FFCD115FFBE8FD37CB78FD28" box="[1033,1047,677,701]" italics="true" pageId="1" pageNumber="262">h</emphasis>
2) and one pair of round cupules
<emphasis id="B936EAB7FFCD115FFA9EFD37CAFAFD28" box="[1407,1429,677,701]" italics="true" pageId="1" pageNumber="262">ih</emphasis>
. Coxal fields I with two pairs of setae (1
<emphasis id="B936EAB7FFCD115FFDB6FD5FCD0AFD71" box="[599,613,717,740]" italics="true" pageId="1" pageNumber="262">a</emphasis>
, 1
<emphasis id="B936EAB7FFCD115FFD60FD5ECDE0FD71" box="[641,655,716,740]" italics="true" pageId="1" pageNumber="262">b</emphasis>
); coxal fields II with two pairs of setae (2
<emphasis id="B936EAB7FFCD115FFB8AFD5FCB16FD71" box="[1131,1145,717,740]" italics="true" pageId="1" pageNumber="262">a</emphasis>
, 2
<emphasis id="B936EAB7FFCD115FFB77FD5ECBCBFD71" box="[1174,1188,716,740]" italics="true" pageId="1" pageNumber="262">b</emphasis>
); coxal fields III with three pairs of setae (3
<emphasis id="B936EAB7FFCD115FFE69FD63CEF9FC9D" box="[392,406,753,776]" italics="true" pageId="1" pageNumber="262">a</emphasis>
, 3
<emphasis id="B936EAB7FFCD115FFE53FD62CEAFFC9D" box="[434,448,752,776]" italics="true" pageId="1" pageNumber="262">b</emphasis>
, 3
<emphasis id="B936EAB7FFCD115FFE3DFD63CE87FC9D" box="[476,488,753,776]" italics="true" pageId="1" pageNumber="262">c</emphasis>
); coxal fields IV with three pairs of setae (4
<emphasis id="B936EAB7FFCD115FFC36FD63CC8AFC9D" box="[983,997,753,776]" italics="true" pageId="1" pageNumber="262">a</emphasis>
, 4
<emphasis id="B936EAB7FFCD115FFBE0FD62CB60FC9D" box="[1025,1039,752,776]" italics="true" pageId="1" pageNumber="262">b</emphasis>
, 4
<emphasis id="B936EAB7FFCD115FFBCAFD63CB58FC9D" box="[1067,1079,753,776]" italics="true" pageId="1" pageNumber="262">c</emphasis>
). Pseudanal segment with three pairs of setae (
<emphasis id="B936EAB7FFCD115FFEA3FC87CE34FCB9" box="[322,347,789,812]" italics="true" pageId="1" pageNumber="262">ps</emphasis>
1-3). Apodemes 1 (ap1) well-developed, apodemes 2 (ap2) well-developed and fused with welldeveloped prosternal apodeme (appr); appr joined with sejugal apodeme (apsej); apodemes 3 (ap3) straight, diffuse, apodemes 4 (ap4) long, reaching to bases of legs III. Apodemes 5 absent. Secondary transverse apodeme absent. Posterior margin of posterior sternal plate entire. Anterior genital sclerite (ags) small, triangular, posterior genital sclerite (pgs) large, triangular, median genital sclerite (mgs) present, oval. Ventral idiosomal setae long, sparsely barbed. Legs I subequal to leg II. Tibiotarsus I slightly enlarged, with well-developed claw; unguinal setae (
<emphasis id="B936EAB7FFCD115FFF41FC62CFC1FB92" box="[160,174,1008,1031]" italics="true" pageId="1" pageNumber="262">u</emphasis>
) spiniform, separated, solenidion
<emphasis id="B936EAB7FFCD115FFDCEFC62CD2CFB92" box="[559,579,1008,1031]" italics="true" pageId="1" pageNumber="262">ω</emphasis>
1 fused with tibiotarsus. Seta
<emphasis id="B936EAB7FFCD115FFC6AFC7DCCF6FB92" box="[907,921,1007,1031]" italics="true" pageId="1" pageNumber="262">d</emphasis>
of femur hook-like. Legs IIIII with one pair of basally thickened, slightly asymmetrical claws and pointed distally empodium; claws on tarsus IV simple, as large as those on tarsi II and III, empodium pointed, but smaller than on tarsi II-III. Tarsus and pretarsus IV short. Femora IIIIV divided into basi- and telofemur. Leg setation: leg I; tr 1 (
<emphasis id="B936EAB7FFCD115FFC27FBCCCCB4FBE0" box="[966,987,1118,1141]" italics="true" pageId="1" pageNumber="262">v</emphasis>
), fe 3 (
<emphasis id="B936EAB7FFCD115FFBCFFBCCCB53FBE3" box="[1070,1084,1118,1142]" italics="true" pageId="1" pageNumber="262">d</emphasis>
,
<emphasis id="B936EAB7FFCD115FFBABFBCCCB34FBE3" box="[1098,1115,1118,1142]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFB88FBCCCBEAFBE0" box="[1129,1157,1118,1141]" italics="true" pageId="1" pageNumber="262">v”</emphasis>
), ge 4 (
<emphasis id="B936EAB7FFCD115FFB3DFBCCCB82FBE3" box="[1244,1261,1118,1142]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFB1DFBCCCA7CFBE3" box="[1276,1299,1118,1142]" italics="true" pageId="1" pageNumber="262">l”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFAC0FBCCCA58FBE0" box="[1313,1335,1118,1141]" italics="true" pageId="1" pageNumber="262">v</emphasis>
,
<emphasis id="B936EAB7FFCD115FFAA4FBCCCA0FFBE0" box="[1349,1376,1118,1141]" italics="true" pageId="1" pageNumber="262">v”</emphasis>
), tita 18(4) (
<emphasis id="B936EAB7FFCD115FFF04FB10CF9CFB0F" box="[229,243,1154,1178]" italics="true" pageId="1" pageNumber="262">d</emphasis>
,
<emphasis id="B936EAB7FFCD115FFEE3FB10CE7CFB0F" box="[258,275,1154,1178]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFEC3FB10CE56FB0F" box="[290,313,1154,1178]" italics="true" pageId="1" pageNumber="262">l”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFEA9FB10CE31FB0C" box="[328,350,1154,1177]" italics="true" pageId="1" pageNumber="262">v</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE8CFB10CEE7FB0C" box="[365,392,1154,1177]" italics="true" pageId="1" pageNumber="262">v”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE79FB10CECBFB0F" box="[408,420,1154,1178]" italics="true" pageId="1" pageNumber="262">k</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE52FB10CEBFFB0C" box="[435,464,1154,1177]" italics="true" pageId="1" pageNumber="262">tc</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE3EFB10CD6CFB0C" box="[479,515,1154,1177]" italics="true" pageId="1" pageNumber="262">tc”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFDF3FB10CD5FFB0C" box="[530,560,1154,1177]" italics="true" pageId="1" pageNumber="262">p”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFDDEFB10CD37FB0F" box="[575,600,1154,1178]" italics="true" pageId="1" pageNumber="262">ft</emphasis>
,
<emphasis id="B936EAB7FFCD115FFD86FB10CDEAFB0F" box="[615,645,1154,1178]" italics="true" pageId="1" pageNumber="262">ft”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFD74FB10CDDBFB0F" box="[661,692,1154,1178]" italics="true" pageId="1" pageNumber="262">pl</emphasis>
,
<emphasis id="B936EAB7FFCD115FFD22FB10CD88FB0F" box="[707,743,1154,1178]" italics="true" pageId="1" pageNumber="262">pl”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFD16FB10CC74FB0C" box="[759,795,1154,1177]" italics="true" pageId="1" pageNumber="262">pv</emphasis>
,
<emphasis id="B936EAB7FFCD115FFCCBFB10CC3CFB0C" box="[810,851,1154,1177]" italics="true" pageId="1" pageNumber="262">pv”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFC82FB11CC01FB0F" box="[867,878,1155,1178]" italics="true" pageId="1" pageNumber="262">s</emphasis>
,
<emphasis id="B936EAB7FFCD115FFC9DFB10CCFCFB0C" box="[892,915,1154,1177]" italics="true" pageId="1" pageNumber="262">u</emphasis>
,
<emphasis id="B936EAB7FFCD115FFC42FB10CCAFFB0C" box="[931,960,1154,1177]" italics="true" pageId="1" pageNumber="262">u”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFC2EFB11CC8CFB0F" box="[975,995,1155,1178]" italics="true" pageId="1" pageNumber="262">ω</emphasis>
1,
<emphasis id="B936EAB7FFCD115FFC1BFB11CB61FB0F" box="[1018,1038,1155,1178]" italics="true" pageId="1" pageNumber="262">ω</emphasis>
2,
<emphasis id="B936EAB7FFCD115FFBC0FB11CB5FFB0F" box="[1057,1072,1155,1178]" italics="true" pageId="1" pageNumber="262">φ</emphasis>
1,
<emphasis id="B936EAB7FFCD115FFBA6FB11CB39FB0F" box="[1095,1110,1155,1178]" italics="true" pageId="1" pageNumber="262">φ</emphasis>
2); leg II: tr 1 (
<emphasis id="B936EAB7FFCD115FFB1DFB10CA7DFB0C" box="[1276,1298,1154,1177]" italics="true" pageId="1" pageNumber="262">v</emphasis>
), fe 3 (
<emphasis id="B936EAB7FFCD115FFA86FB10CA1AFB0F" box="[1383,1397,1154,1178]" italics="true" pageId="1" pageNumber="262">d</emphasis>
,
<emphasis id="B936EAB7FFCD115FFA65FB10CAFAFB0F" box="[1412,1429,1154,1178]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFF76FB3BCFDDFB55" box="[151,178,1193,1216]" italics="true" pageId="1" pageNumber="262">v”</emphasis>
), ge 3 (
<emphasis id="B936EAB7FFCD115FFEEAFB3BCE73FB54" box="[267,284,1193,1217]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFECBFB3BCE33FB55" box="[298,348,1193,1217]" italics="true" pageId="1" pageNumber="262">l”, v</emphasis>
), ti 4(1) (
<emphasis id="B936EAB7FFCD115FFE35FB3BCE8DFB54" box="[468,482,1193,1217]" italics="true" pageId="1" pageNumber="262">d</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE11FB3BCD6EFB54" box="[496,513,1193,1217]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFDEEFB3BCD4AFB55" box="[527,549,1193,1216]" italics="true" pageId="1" pageNumber="262">v</emphasis>
,
<emphasis id="B936EAB7FFCD115FFDD2FB3BCD21FB55" box="[563,590,1193,1216]" italics="true" pageId="1" pageNumber="262">v”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFDBFFB3BCD02FB55" box="[606,621,1193,1216]" italics="true" pageId="1" pageNumber="262">φ</emphasis>
), ta 6(1) (
<emphasis id="B936EAB7FFCD115FFD01FB3BCD92FB55" box="[736,765,1193,1216]" italics="true" pageId="1" pageNumber="262">tc</emphasis>
,
<emphasis id="B936EAB7FFCD115FFCEDFB3BCC40FB55" box="[780,815,1193,1216]" italics="true" pageId="1" pageNumber="262">tc”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFCDCFB3BCC0DFB54" box="[829,866,1193,1217]" italics="true" pageId="1" pageNumber="262">pl”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFC90FB3BCCFAFB55" box="[881,917,1193,1216]" italics="true" pageId="1" pageNumber="262">pv</emphasis>
,
<emphasis id="B936EAB7FFCD115FFC42FB3BCCA2FB55" box="[931,973,1193,1216]" italics="true" pageId="1" pageNumber="262">pv”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFC3DFB3BCC9CFB55" box="[988,1011,1193,1216]" italics="true" pageId="1" pageNumber="262">u</emphasis>
,
<emphasis id="B936EAB7FFCD115FFBE0FB3BCB7AFB55" box="[1025,1045,1193,1216]" italics="true" pageId="1" pageNumber="262">ω</emphasis>
); leg III: tr 1 (
<emphasis id="B936EAB7FFCD115FFB58FB3BCBA0FB55" box="[1209,1231,1193,1216]" italics="true" pageId="1" pageNumber="262">v</emphasis>
), fe 2 (
<emphasis id="B936EAB7FFCD115FFAC3FB3BCA5FFB54" box="[1314,1328,1193,1217]" italics="true" pageId="1" pageNumber="262">d</emphasis>
,
<emphasis id="B936EAB7FFCD115FFADEFB3BCA3AFB55" box="[1343,1365,1193,1216]" italics="true" pageId="1" pageNumber="262">v</emphasis>
), ge 2 (
<emphasis id="B936EAB7FFCD115FFF41FB5FCFDEFB70" box="[160,177,1229,1253]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFF5EFB5FCFBAFB71" box="[191,213,1229,1252]" italics="true" pageId="1" pageNumber="262">v</emphasis>
), ti 4(1) (
<emphasis id="B936EAB7FFCD115FFEA1FB5FCE21FB70" box="[320,334,1229,1253]" italics="true" pageId="1" pageNumber="262">d</emphasis>
,
<emphasis id="B936EAB7FFCD115FFEBDFB5FCE02FB70" box="[348,365,1229,1253]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE9BFB5FCEFFFB71" box="[378,400,1229,1252]" italics="true" pageId="1" pageNumber="262">v</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE7CFB5FCED6FB71" box="[413,441,1229,1252]" italics="true" pageId="1" pageNumber="262">v”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE26FB5FCEB9FB71" box="[455,470,1229,1252]" italics="true" pageId="1" pageNumber="262">φ</emphasis>
), ta 6 (
<emphasis id="B936EAB7FFCD115FFDC7FB5FCD2CFB71" box="[550,579,1229,1252]" italics="true" pageId="1" pageNumber="262">tc</emphasis>
,
<emphasis id="B936EAB7FFCD115FFDB0FB5FCD1AFB71" box="[593,629,1229,1252]" italics="true" pageId="1" pageNumber="262">tc”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFD62FB5FCDC8FB70" box="[643,679,1229,1253]" italics="true" pageId="1" pageNumber="262">pl”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFD54FB5FCDB7FB71" box="[693,728,1229,1252]" italics="true" pageId="1" pageNumber="262">pv</emphasis>
,
<emphasis id="B936EAB7FFCD115FFD06FB5FCC7FFB71" box="[743,784,1229,1252]" italics="true" pageId="1" pageNumber="262">pv”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFCFFFB5FCC5AFB71" box="[798,821,1229,1252]" italics="true" pageId="1" pageNumber="262">u</emphasis>
); leg IV: tr 1 (
<emphasis id="B936EAB7FFCD115FFC34FB5FCC85FB71" box="[981,1002,1229,1252]" italics="true" pageId="1" pageNumber="262">v</emphasis>
), fe 2 (
<emphasis id="B936EAB7FFCD115FFBDDFB5FCB25FB70" box="[1084,1098,1229,1253]" italics="true" pageId="1" pageNumber="262">d</emphasis>
,
<emphasis id="B936EAB7FFCD115FFBB9FB5FCB01FB71" box="[1112,1134,1229,1252]" italics="true" pageId="1" pageNumber="262">v</emphasis>
), ge 1 (
<emphasis id="B936EAB7FFCD115FFB22FB5FCBB6FB71" box="[1219,1241,1229,1252]" italics="true" pageId="1" pageNumber="262">v</emphasis>
), ti 4(1) (
<emphasis id="B936EAB7FFCD115FFAA4FB5FCA3CFB70" box="[1349,1363,1229,1253]" italics="true" pageId="1" pageNumber="262">d</emphasis>
,
<emphasis id="B936EAB7FFCD115FFA81FB5FCA1EFB70" box="[1376,1393,1229,1253]" italics="true" pageId="1" pageNumber="262">l</emphasis>
,
<emphasis id="B936EAB7FFCD115FFA9EFB5FCAFAFB71" box="[1407,1429,1229,1252]" italics="true" pageId="1" pageNumber="262">v</emphasis>
,
<emphasis id="B936EAB7FFCD115FFF76FB63CFDDFA9D" box="[151,178,1265,1288]" italics="true" pageId="1" pageNumber="262">v”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFF20FB63CFBFFA9D" box="[193,208,1265,1288]" italics="true" pageId="1" pageNumber="262">φ</emphasis>
), ta 6 (
<emphasis id="B936EAB7FFCD115FFEC0FB63CE29FA9C" box="[289,326,1265,1289]" italics="true" pageId="1" pageNumber="262">pl”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFEB5FB63CE1EFA9D" box="[340,369,1265,1288]" italics="true" pageId="1" pageNumber="262">tc</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE61FB63CECBFA9D" box="[384,420,1265,1288]" italics="true" pageId="1" pageNumber="262">tc”</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE53FB63CEA6FA9D" box="[434,457,1265,1288]" italics="true" pageId="1" pageNumber="262">u</emphasis>
,
<emphasis id="B936EAB7FFCD115FFE36FB63CE94FA9D" box="[471,507,1265,1288]" italics="true" pageId="1" pageNumber="262">pv</emphasis>
,
<emphasis id="B936EAB7FFCD115FFDE8FB63CD5DFA9D" box="[521,562,1265,1288]" italics="true" pageId="1" pageNumber="262">pv”</emphasis>
).
</paragraph>
<paragraph id="8BFD36A5FFCD115FFF26FA86CEFFF9C3" blockId="1.[151,1436,420,1838]" pageId="1" pageNumber="262">
<emphasis id="B936EAB7FFCD115FFF26FA86CEA6FAB8" bold="true" box="[199,457,1300,1325]" pageId="1" pageNumber="262">Differential diagnosis</emphasis>
. The new genus is most similar to
<taxonomicName id="4C424D26FFCD115FFC88FA87CBABFAB8" authority="Cross, 1965" authorityName="Cross" authorityYear="1965" box="[873,1220,1300,1325]" class="Arachnida" family="Pygmephoridae" genus="Allopygmephorus" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFC88FA87CB41FAB8" box="[873,1070,1301,1325]" italics="true" pageId="1" pageNumber="262">Allopygmephorus</emphasis>
<bibRefCitation id="EFD34B54FFCD115FFBD9FA87CBABFAB8" author="Cross" box="[1080,1220,1300,1325]" pageId="1" pageNumber="262" refString="Cross, E. A. (1965) The generic relationships of the family Pyemotidae (Acarina, Trombidiformes). The University of Kansas Science Bulletin, 45, 29 - 215." type="journal article" year="1965">Cross, 1965</bibRefCitation>
</taxonomicName>
. Both genera have characteristic pointed empodia on tarsi II-IV, separated spiniform unguinal setae on tibiotarsus I, the same leg setation, and similarly shaped claws on the legs. The new genus differs from
<taxonomicName id="4C424D26FFCD115FFBE1FACFCBAAFAE0" box="[1024,1221,1373,1397]" class="Arachnida" family="Pygmephoridae" genus="Allopygmephorus" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFBE1FACFCBAAFAE0" box="[1024,1221,1373,1397]" italics="true" pageId="1" pageNumber="262">Allopygmephorus</emphasis>
</taxonomicName>
by the presence of setae
<date id="FFFC1065FFCD115FFF37FA13CF85FA08" box="[214,234,1409,1437]" pageId="1" pageNumber="262">
<emphasis id="B936EAB7FFCD115FFF37FA13CF8DFA0D" box="[214,226,1409,1432]" italics="true" pageId="1" pageNumber="262">v</emphasis>
2
</date>
(absent in
<taxonomicName id="4C424D26FFCD115FFE87FA13CD43FA0C" box="[358,556,1409,1433]" class="Arachnida" family="Pygmephoridae" genus="Allopygmephorus" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFE87FA13CD43FA0C" box="[358,556,1409,1433]" italics="true" pageId="1" pageNumber="262">Allopygmephorus</emphasis>
</taxonomicName>
), the presence of the median genital sclerite (absent in
<taxonomicName id="4C424D26FFCD115FFB77FA13CA34FA0C" box="[1174,1371,1409,1433]" class="Arachnida" family="Pygmephoridae" genus="Allopygmephorus" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFB77FA13CA34FA0C" box="[1174,1371,1409,1433]" italics="true" pageId="1" pageNumber="262">Allopygmephorus</emphasis>
</taxonomicName>
), and by solenidion
<emphasis id="B936EAB7FFCD115FFEDBFA3ACE21FA2A" box="[314,334,1448,1471]" italics="true" pageId="1" pageNumber="262">ω</emphasis>
1 fused with tibiotarsus I (prominent in
<taxonomicName id="4C424D26FFCD115FFCF5FA35CCB6FA2A" box="[788,985,1447,1471]" class="Arachnida" family="Pygmephoridae" genus="Allopygmephorus" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFCF5FA35CCB6FA2A" box="[788,985,1447,1471]" italics="true" pageId="1" pageNumber="262">Allopygmephorus</emphasis>
</taxonomicName>
). Among neopygmephorid genera with median genital sclerite, the new genus is most similar to
<taxonomicName id="4C424D26FFCD115FFCC5FA5CCBCFFA72" authority="Cross, 1965" authorityName="Cross" authorityYear="1965" box="[804,1184,1486,1511]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFCC5FA5CCB62FA73" box="[804,1037,1486,1510]" italics="true" pageId="1" pageNumber="262">Pseudopygmephorus</emphasis>
<bibRefCitation id="EFD34B54FFCD115FFBF7FA5DCBCFFA72" author="Cross" box="[1046,1184,1486,1511]" pageId="1" pageNumber="262" refString="Cross, E. A. (1965) The generic relationships of the family Pyemotidae (Acarina, Trombidiformes). The University of Kansas Science Bulletin, 45, 29 - 215." type="journal article" year="1965">Cross, 1965</bibRefCitation>
</taxonomicName>
, by having solenidion
<emphasis id="B936EAB7FFCD115FFF76FA61CFC4F99F" box="[151,171,1523,1546]" italics="true" pageId="1" pageNumber="262">ω</emphasis>
1fused with tibiotarsus I. The new genus differs from
<taxonomicName id="4C424D26FFCD115FFCECFA60CC99F99F" box="[781,1014,1522,1546]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFCECFA60CC99F99F" box="[781,1014,1522,1546]" italics="true" pageId="1" pageNumber="262">Pseudopygmephorus</emphasis>
</taxonomicName>
by the characteristic distally-pointed empodia on tarsi IIIV (flipper-like in
<taxonomicName id="4C424D26FFCD115FFD74F98BCC11F9A4" box="[661,894,1561,1585]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFD74F98BCC11F9A4" box="[661,894,1561,1585]" italics="true" pageId="1" pageNumber="262">Pseudopygmephorus</emphasis>
</taxonomicName>
), and non-bifurcate setae 1
<emphasis id="B936EAB7FFCD115FFB05F98BCB9DF9A4" box="[1252,1266,1561,1585]" italics="true" pageId="1" pageNumber="262">b</emphasis>
(bifurcate in
<taxonomicName id="4C424D26FFCD115FFF76F9AFCEEFF9C0" box="[151,384,1597,1621]" class="Arachnida" family="Pygmephoridae" genus="Pseudopygmephorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFF76F9AFCEEFF9C0" box="[151,384,1597,1621]" italics="true" pageId="1" pageNumber="262">Pseudopygmephorus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BFD36A5FFCD115FFF26F9F3CFA8F908" blockId="1.[151,1436,420,1838]" pageId="1" pageNumber="262">
<emphasis id="B936EAB7FFCD115FFF26F9F3CEE1F9EF" bold="true" box="[199,398,1633,1658]" pageId="1" pageNumber="262">Species included</emphasis>
. The genus
<taxonomicName id="4C424D26FFCD115FFDC1F9F3CC4EF9EC" box="[544,801,1633,1657]" class="Insecta" family="Heteroceridae" genus="Protoallopygmephorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B936EAB7FFCD115FFDC1F9F3CC4EF9EC" box="[544,801,1633,1657]" italics="true" pageId="1" pageNumber="262">Protoallopygmephorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557CCFFCD115FFCCAF9F3CCF8F9EC" box="[811,919,1633,1658]" pageId="1" pageNumber="262" rank="genus">
<emphasis id="B936EAB7FFCD115FFCCAF9F3CCFFF9EF" bold="true" box="[811,912,1633,1658]" pageId="1" pageNumber="262">gen. nov</emphasis>
.
</taxonomicNameLabel>
includes only type species,
<taxonomicName id="4C424D26FFCD115FFB01F9F0CA1FF9EC" box="[1248,1392,1633,1657]" class="Insecta" family="Heteroceridae" genus="Protoallopygmephorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="heteroceri" status="sp. nov.">
<emphasis id="B936EAB7FFCD115FFB01F9F0CA1FF9EC" box="[1248,1392,1633,1657]" italics="true" pageId="1" pageNumber="262">P. heteroceri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557CCFFCD115FFA9BF9F3CFA8F908" pageId="1" pageNumber="262" rank="species">
<emphasis id="B936EAB7FFCD115FFA9BF9F3CFAEF90B" bold="true" pageId="1" pageNumber="262">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C358652EFFCD115FFF26F93BCCD6F973" pageId="1" pageNumber="262" type="distribution">
<paragraph id="8BFD36A5FFCD115FFF26F93BCCD6F973" blockId="1.[151,1436,420,1838]" pageId="1" pageNumber="262">
<emphasis id="B936EAB7FFCD115FFF26F93BCE81F957" bold="true" box="[199,494,1705,1730]" pageId="1" pageNumber="262">Distribution and habitat</emphasis>
.
<taxonomicName id="4C424D26FFCD115FFE1FF93BCC15F954" box="[510,890,1705,1729]" class="Insecta" family="Heteroceridae" genus="Protoallopygmephorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="heteroceri" status="sp. nov.">
<emphasis id="B936EAB7FFCD115FFE1FF93BCC15F954" box="[510,890,1705,1729]" italics="true" pageId="1" pageNumber="262">Protoallopygmephorus heteroceri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20557CCFFCD115FFC65F93BCCB0F954" box="[900,991,1705,1730]" pageId="1" pageNumber="262" rank="species">
<emphasis id="B936EAB7FFCD115FFC65F93BCCB7F957" bold="true" box="[900,984,1705,1730]" pageId="1" pageNumber="262">sp. nov</emphasis>
.
</taxonomicNameLabel>
is phoretic on
<taxonomicName id="4C424D26FFCD115FFB6FF938CE79F973" authority="Thunberg" authorityName="Thunberg" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="fenestratus">
<emphasis id="B936EAB7FFCD115FFB6FF938CAF3F954" box="[1166,1436,1705,1729]" italics="true" pageId="1" pageNumber="262">Heterocerus fenestratus</emphasis>
(Thunberg)
</taxonomicName>
(
<taxonomicName id="4C424D26FFCD115FFEC5F95FCECAF973" box="[292,421,1741,1766]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C424D26FFCD115FFE51F95FCD20F973" box="[432,591,1741,1766]" class="Insecta" family="Heteroceridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="262" phylum="Arthropoda" rank="family">Heteroceridae</taxonomicName>
) in the European part of
<collectingCountry id="F3557635FFCD115FFC87F95FCCDAF973" box="[870,949,1741,1766]" name="Russia" pageId="1" pageNumber="262">Russia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C358652EFFCD115FFF26F963CB1EF8B8" pageId="1" pageNumber="262" type="etymology">
<paragraph id="8BFD36A5FFCD115FFF26F963CB1EF8B8" blockId="1.[151,1436,420,1838]" pageId="1" pageNumber="262">
<emphasis id="B936EAB7FFCD115FFF26F963CE29F89F" bold="true" box="[199,326,1777,1802]" pageId="1" pageNumber="262">Etymology</emphasis>
. The name of the new genus is combined from Greek
<emphasis id="B936EAB7FFCD115FFC45F963CC84F89C" box="[932,1003,1777,1801]" italics="true" pageId="1" pageNumber="262">πρωτΟ</emphasis>
meaning
<emphasis id="B936EAB7FFCD115FFBB8F963CBE2F89C" box="[1113,1165,1777,1801]" italics="true" pageId="1" pageNumber="262">first,</emphasis>
and the name of closely related genus
<taxonomicName id="4C424D26FFCD115FFED3F887CE98F8B8" box="[306,503,1813,1837]" class="Arachnida" family="Pygmephoridae" genus="Allopygmephorus" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B936EAB7FFCD115FFED3F887CE98F8B8" box="[306,503,1813,1837]" italics="true" pageId="1" pageNumber="262">Allopygmephorus</emphasis>
</taxonomicName>
. It refers to its similarity to the genus
<emphasis id="B936EAB7FFCD115FFC45F887CB1EF8B8" box="[932,1137,1813,1837]" italics="true" pageId="1" pageNumber="262">
<taxonomicName id="4C424D26FFCD115FFC45F887CB02F8B8" box="[932,1133,1813,1837]" class="Arachnida" family="Pygmephoridae" genus="Allopygmephorus" kingdom="Animalia" order="Prostigmata" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">Allopygmephorus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>