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<document id="0DE0388E55347A63F1D601163BEEF236" ID-CLB-Dataset="37216" ID-DOI="10.5852/ejt.2016.252" ID-GBIF-Dataset="7fad12f5-99eb-4308-a83e-53b274aca16b" ID-ISSN="2118-9773" ID-Zenodo-Dep="221250" ID-ZooBank="urn:lsid:zoobank.org:pub:F4F9AFF3-C4D3-4BFE-B4C8-516C14758DAE" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1482705154557" checkinUser="plazi" docAuthor="Horia R. Galea, Cornelia Roder, Christoph Walcher, Marco Warmuth, Eberhard Kohlberg &amp; Philipp F. Fischer" docDate="2016" docId="03EB879BFF9CFFB1FDFC0004FAE0E574" docLanguage="en" docName="376-1923-1-PB.pdf" docOrigin="European Journal of Taxonomy 252" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tesserogastria musculosa Beyer 1959" docType="treatment" docVersion="15" lastPageNumber="17" masterDocId="FFD2FFE3FF93FFA1FFC8060DFF8CE319" masterDocTitle="Glaciambulata neumayeri gen. et sp. nov., a new Antarctic trachymedusa (Cnidaria: Hydrozoa), with a revision of the family Ptychogastriidae" masterLastPageNumber="30" masterPageNumber="1" pageNumber="16" updateTime="1698435296770" updateUser="ExternalLinkService">
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<mods:title id="76B852B247C552DE7C7CB1EE9E2E7CF0">Glaciambulata neumayeri gen. et sp. nov., a new Antarctic trachymedusa (Cnidaria: Hydrozoa), with a revision of the family Ptychogastriidae</mods:title>
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<mods:namePart id="604160F817A651BF7906AFA4CED9F98C">Horia R. Galea</mods:namePart>
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<paragraph id="8BFD368DFF9CFFAEFDFC0004FB8CE53D" blockId="15.[564,1024,1545,1606]" box="[564,1024,1545,1572]" pageId="15" pageNumber="16">
<heading id="D0B581E1FF9CFFAEFDFC0004FB8CE53D" box="[564,1024,1545,1572]" centered="true" fontSize="11" level="3" pageId="15" pageNumber="16" reason="4">
<taxonomicName id="4C424D0EFF9CFFAEFDFC0004FB8CE53D" ID-CoL="55JH5" authority="Beyer, 1959" authorityName="Beyer" authorityYear="1959" box="[564,1024,1545,1572]" class="Hydrozoa" family="Ptychogastriidae" genus="Tesserogastria" kingdom="Animalia" order="Trachymedusae" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="musculosa">
<emphasis id="B936EA9FFF9CFFAEFDFC0004FCE6E53D" bold="true" box="[564,874,1545,1572]" italics="true" pageId="15" pageNumber="16">Tesserogastria musculosa</emphasis>
<bibRefCitation id="EFD34B7CFF9CFFAEFCB90007FB8CE53D" author="Beyer F." box="[881,1024,1545,1572]" journalOrPublisher="Nytt Magasin for Zoologi" pageId="15" pageNumber="16" pagination="121 - 143" part="6" refString="Beyer F. 1959. A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi 6: 121 - 143." title="A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps" type="journal article" year="1959">Beyer, 1959</bibRefCitation>
</taxonomicName>
</heading>
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<subSubSection id="C3586506FF9CFFAEFD150021FCDAE55F" box="[733,854,1580,1606]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BFD368DFF9CFFAEFD150021FCDAE55F" blockId="15.[564,1024,1545,1606]" box="[733,854,1580,1606]" pageId="15" pageNumber="16">
<figureCitation id="13792A08FF9CFFAEFD150021FCADE55F" box="[733,801,1580,1606]" captionStart="Fig. 4" captionStartId="11.[189,231,1788,1814]" captionTargetBox="[195,1394,408,1741]" captionTargetId="figure@11.[189,1398,401,1745]" captionTargetPageId="11" captionText="Fig. 4. — A B. Ptychogastria asteroides (Haeckel, 1879). A. Lateral view of a medusa. B. Portion of umbrella margin showing the arrangement of tentacles. — C E. Ptychogastria polaris Allman, 1878. C. Medusa in lateral view. D. Oral view of half of the medusa, showing the arrangement of the marginal tentacles. E. Portion of the bell margin, greatly enlarged. — F G. Tesserogastria musculosa Beyer, 1959. Medusa in lateral (F) and oral (G) aspects. [A, B redrawn after Gili et al. (1999); C, E redrawn after Panteleeva et al. (1999); D redrawn after Haeckel (1881 b); F, G redrawn after Hesthagen (1971)]. Scale bars: A, B = 500 µm; C = 3 mm; E = 2 mm." httpUri="https://zenodo.org/record/221253/files/figure.png" pageId="15" pageNumber="16">Fig. 4</figureCitation>
FG
</paragraph>
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<paragraph id="8BFD368DFF9CFFAEFF750062FCA2E591" blockId="15.[189,814,1646,1673]" box="[189,814,1646,1673]" pageId="15" pageNumber="16">
<taxonomicName id="4C424D0EFF9CFFAEFF750062FD33E591" ID-CoL="55JH5" authority="Beyer, 1959: 122" authorityName="Beyer" authorityPageNumber="122" authorityYear="1959" box="[189,703,1646,1673]" class="Hydrozoa" family="Ptychogastriidae" genus="Tesserogastria" kingdom="Animalia" order="Trachymedusae" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="musculosa">
<emphasis id="B936EA9FFF9CFFAEFF750062FE67E591" box="[189,491,1646,1672]" italics="true" pageId="15" pageNumber="16">Tesserogastria musculosa</emphasis>
<bibRefCitation id="EFD34B7CFF9CFFAEFE3A0063FD09E590" author="Beyer F." box="[498,645,1646,1673]" journalOrPublisher="Nytt Magasin for Zoologi" pageId="15" pageNumber="16" pagination="121 - 143" part="6" refString="Beyer F. 1959. A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi 6: 121 - 143." title="A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps" type="journal article" year="1959">Beyer, 1959</bibRefCitation>
: 122
</taxonomicName>
, pls 1 2.
</paragraph>
<paragraph id="8BFD368DFF9CFFAEFF7500BDFDECE5F4" blockId="15.[189,1399,1711,1773]" pageId="15" pageNumber="16">
<taxonomicName id="4C424D0EFF9CFFAEFF7500BDFE66E5D0" ID-CoL="55JH5" box="[189,490,1711,1737]" class="Hydrozoa" family="Ptychogastriidae" genus="Tesserogastria" kingdom="Animalia" order="Trachymedusae" pageId="15" pageNumber="16" phylum="Cnidaria" rank="species" species="musculosa">
<emphasis id="B936EA9FFF9CFFAEFF7500BDFE66E5D0" box="[189,490,1711,1737]" italics="true" pageId="15" pageNumber="16">Tesserogastria musculosa</emphasis>
</taxonomicName>
<bibRefCitation id="EFD34B7CFF9CFFAEFDCD00A2FD4AE5D0" author="Hesthagen I. H." box="[517,710,1711,1737]" journalOrPublisher="Norwegian Journal of Zoology" pageId="15" pageNumber="16" pagination="1 - 19" part="19" refString="Hesthagen I. H. 1971. On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer. Norwegian Journal of Zoology 19 (1): 1 - 19." title="On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer" type="journal article" year="1971">Hesthagen 1971</bibRefCitation>
: 3, Fgs 210. — Bouillon 1985: 202. —
<bibRefCitation id="EFD34B7CFF9CFFAEFB5400BDFAFFE5D0" author="Larson R. J. &amp; Matsumoto G. I. &amp; Madin L. P. &amp; Lewis L. M." box="[1180,1395,1711,1738]" journalOrPublisher="Bulletin of Marine Science" pageId="15" pageNumber="16" pagination="277 - 286" part="51" refString="Larson R. J., Matsumoto G. I., Madin L. P. &amp; Lewis L. M. 1992. Deep-sea benthic and benthopelagic medusae: recent observations from submersibles and a remotely operated vehicle. Bulletin of Marine Science 51 (3): 277 - 286." title="Deep-sea benthic and benthopelagic medusae: recent observations from submersibles and a remotely operated vehicle" type="journal article" year="1992">
Larson
<emphasis id="B936EA9FFF9CFFAEFB3C00BDFAAAE5D0" box="[1268,1318,1711,1737]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. 1992
</bibRefCitation>
: 284. —
<bibRefCitation id="EFD34B7CFF9CFFAEFEFB00DFFDAEE5F5" author="Bouillon J. &amp; Gravili C. &amp; Pages F. &amp; Gili J. M. &amp; Boero F." box="[307,546,1746,1773]" journalOrPublisher="Museum national d'Histoire naturelle, Paris" pageId="15" pageNumber="16" pagination="194" publicationUrl="http://dx.doi" refString="Bouillon J., Gravili C., Pages F., Gili J. M. &amp; Boero F. 2006. An introduction to Hydrozoa. Memoires du Museum national d'Histoire naturelle 194, Museum national d'Histoire naturelle, Paris. http: // dx. doi. org / 10.1086 / 586983" title="An introduction to Hydrozoa. Memoires du Museum national d'Histoire naturelle" type="book chapter" year="2006">
Bouillon
<emphasis id="B936EA9FFF9CFFAEFE6900DEFE59E5F5" box="[417,469,1746,1772]" italics="true" pageId="15" pageNumber="16">et al</emphasis>
. 2006
</bibRefCitation>
: 112.
</paragraph>
</subSubSection>
<subSubSection id="C3586506FF9CFFB1FF750119FEB5E1F6" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="8BFD368DFF9CFFAEFF750119FEC2E437" blockId="15.[189,1399,1812,2026]" box="[189,334,1812,1838]" pageId="15" pageNumber="16">
<heading id="D0B581E1FF9CFFAEFF750119FEC2E437" bold="true" box="[189,334,1812,1838]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="6">
<emphasis id="B936EA9FFF9CFFAEFF750119FEC2E437" bold="true" box="[189,334,1812,1838]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFD368DFF9CFFB1FF75014FFEB5E1F6" blockId="15.[189,1399,1812,2026]" lastBlockId="16.[189,1399,267,751]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">Umbrella bell-shaped, about as high as broad (up to 2.5 mm), with a small, blunt, apical projection surrounded by a circular depression. Exumbrella devoid of radial ridges. Mesoglea rather thin, comparatively thickened at margin, accommodating a belt of chordal cells on which the tentacles insert. Over 300 tentacles, inserted at 35 superimposed levels, apparently not forming distinct rows or clusters; solid, core composed of a single row of chordal cells; proximally constricted, leaving tear drop-shaped scars when shed; blunt-ended to pointed distally, and there devoid of adhesive pads; nematocysts evenly dispersed in epidermis, except for the distal tip, where they are comparatively crowded. Manubrium cross-shaped (four perradial lobes are distinguishable), except proximally where it assumes an octagonal shape at the origin of the radial canals; relatively long, though devoid of a peduncle, and extending somewhat beyond the bell margin; mesenteries absent; mouth square, with four simple lips provided with ciliated cells in the epidermis. Eight radial canals with rounded cross sections connect to the broad ring canal, elliptical in cross section, longer axis oral-aboral; centripetal canals absent. A conspicuous nematocyst ring on lower side of umbrella margin. Velum extremely broad and strong. Eight slender, elongated gonads, in four perradial pairs; one pair conFned to each manubrial lobe. Eight small, shortstalked, interradial statocysts with single, spherical statoliths; one statocyst in each octant. Cnidome: almost spherical stenoteles [(78) × (67) µm], tear drop-shaped microbasic euryteles [(47) × (3 5) µm], and atrichous isorhizas (ca 2 µm); the capsules occur in both tentacles and the marginal ring. Color: umbrella translucent, with white to yellow epidermal spots at junction of radial canals with the ring canal.</paragraph>
</subSubSection>
<subSubSection id="C3586506FF83FFB1FF75051AFE1AE63D" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BFD368DFF83FFB1FF75051AFEA3E028" blockId="16.[189,303,791,817]" box="[189,303,791,817]" pageId="16" pageNumber="17">
<heading id="D0B581E1FF83FFB1FF75051AFEA3E028" bold="true" box="[189,303,791,817]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<emphasis id="B936EA9FFF83FFB1FF75051AFEA3E028" bold="true" box="[189,303,791,817]" pageId="16" pageNumber="17">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BFD368DFF83FFB1FF750548FBB8E7A6" blockId="16.[189,1399,837,1216]" pageId="16" pageNumber="17">
Although the tentacles were described as “not arranged in distinct rows, nor in clusters” around the bell margin,
<bibRefCitation id="EFD34B7CFF83FFB1FE9E0564FE7FE09A" author="Beyer F." box="[342,499,872,899]" journalOrPublisher="Nytt Magasin for Zoologi" pageId="16" pageNumber="17" pagination="121 - 143" part="6" refString="Beyer F. 1959. A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi 6: 121 - 143." title="A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps" type="journal article" year="1959">Beyer (1959)</bibRefCitation>
acknowledged that “It is possible, however, to trace some repetition in the arrangement, and this makes 48 groups […]. The pattern is thus not quite unlike that of
<taxonomicName id="4C424D0EFF83FFB1FB1A0586FEFCE0D1" authority="Allman" authorityName="Allman" class="Hydrozoa" family="Ptychogastriidae" genus="Ptychogastria" kingdom="Animalia" order="Trachymedusae" pageId="16" pageNumber="17" phylum="Cnidaria" rank="species" species="polaris">
<emphasis id="B936EA9FFF83FFB1FB1A0586FE9CE0D1" italics="true" pageId="16" pageNumber="17">Ptychogastria polaris</emphasis>
Allman
</taxonomicName>
[…]”. However, no further details were given relative to the position of these groups with respect to the radial canals. Their presence, if conFrmed again, would not be surprising, given that more or less distinct groups of tentacles can be found in both
<taxonomicName id="4C424D0EFF83FFB1FC1505F8FB0EE716" box="[989,1154,1013,1039]" class="Hydrozoa" family="Ptychogastriidae" genus="Ptychogastria" kingdom="Animalia" order="Trachymedusae" pageId="16" pageNumber="17" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF83FFB1FC1505F8FB0EE716" box="[989,1154,1013,1039]" italics="true" pageId="16" pageNumber="17">Ptychogastria</emphasis>
</taxonomicName>
and
<taxonomicName id="4C424D0EFF83FFB1FB7705F8FAE3E716" box="[1215,1391,1013,1039]" class="Hydrozoa" family="Ptychogastriidae" genus="Glaciambulata" kingdom="Animalia" order="Trachymedusae" pageId="16" pageNumber="16" phylum="Cnidaria" rank="genus" status="gen. nov.">
<emphasis id="B936EA9FFF83FFB1FB7705F8FAE3E716" box="[1215,1391,1013,1039]" italics="true" pageId="16" pageNumber="17">Glaciambulata</emphasis>
</taxonomicName>
, the sole marked difference in this respect being the total absence of tentacles with adhesive pads in
<taxonomicName id="4C424D0EFF83FFB1FF750231FEE4E74C" box="[189,360,1084,1109]" class="Hydrozoa" family="Ptychogastriidae" genus="Tesserogastria" kingdom="Animalia" order="Trachymedusae" pageId="16" pageNumber="17" phylum="Cnidaria" rank="genus">
<emphasis id="B936EA9FFF83FFB1FF750231FEE4E74C" box="[189,360,1084,1109]" italics="true" pageId="16" pageNumber="17">Tesserogastria</emphasis>
</taxonomicName>
. Though no such structures have been observed, “The amount of debris so frequently found to be sticking to the tentacle ends indicate that they are adhesive” (
<bibRefCitation id="EFD34B7CFF83FFB1FBF80252FB4CE760" author="Beyer F." box="[1072,1216,1118,1145]" journalOrPublisher="Nytt Magasin for Zoologi" pageId="16" pageNumber="17" pagination="121 - 143" part="6" refString="Beyer F. 1959. A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi 6: 121 - 143." title="A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps" type="journal article" year="1959">Beyer 1959</bibRefCitation>
). The so called adhesion is, most probably, due to the crowded condition of the nematocysts in the tentacle tips (
<bibRefCitation id="EFD34B7CFF83FFB1FAE7028FFF77E7D9" author="Beyer F." journalOrPublisher="Nytt Magasin for Zoologi" pageId="16" pageNumber="17" pagination="121 - 143" part="6" refString="Beyer F. 1959. A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi 6: 121 - 143." title="A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps" type="journal article" year="1959">Beyer 1959</bibRefCitation>
), and the possible glutinant nature of some of them (
<bibRefCitation id="EFD34B7CFF83FFB1FCAB02A8FBABE7A6" author="Hesthagen I. H." box="[867,1063,1189,1215]" journalOrPublisher="Norwegian Journal of Zoology" pageId="16" pageNumber="17" pagination="1 - 19" part="19" refString="Hesthagen I. H. 1971. On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer. Norwegian Journal of Zoology 19 (1): 1 - 19." title="On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer" type="journal article" year="1971">Hesthagen 1971</bibRefCitation>
).
</paragraph>
<paragraph id="8BFD368DFF83FFB1FF7502EBFE1AE63D" blockId="16.[189,1398,1254,1316]" pageId="16" pageNumber="17">
The development from the egg, and the behavior of the medusa were thoroughly documented by
<bibRefCitation id="EFD34B7CFF83FFB1FF750304FE1EE63D" author="Hesthagen I. H." box="[189,402,1289,1316]" journalOrPublisher="Norwegian Journal of Zoology" pageId="16" pageNumber="17" pagination="1 - 19" part="19" refString="Hesthagen I. H. 1971. On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer. Norwegian Journal of Zoology 19 (1): 1 - 19." title="On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer" type="journal article" year="1971">Hesthagen (1971)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3586506FF83FFB1FF750346FDF0E6E4" pageId="16" pageNumber="17" type="biology_ecology">
<paragraph id="8BFD368DFF83FFB1FF750346FEACE67C" blockId="16.[189,1399,1355,1534]" box="[189,288,1355,1381]" pageId="16" pageNumber="17">
<heading id="D0B581E1FF83FFB1FF750346FEACE67C" bold="true" box="[189,288,1355,1381]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="6">
<emphasis id="B936EA9FFF83FFB1FF750346FEACE67C" bold="true" box="[189,288,1355,1381]" pageId="16" pageNumber="17">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFD368DFF83FFB1FF750377FDF0E6E4" blockId="16.[189,1399,1355,1534]" pageId="16" pageNumber="17">
A soft bottom species, never captured in plankton nets or vertical hauls (
<bibRefCitation id="EFD34B7CFF83FFB1FBF80377FB4DE68D" author="Beyer F." box="[1072,1217,1401,1428]" journalOrPublisher="Nytt Magasin for Zoologi" pageId="16" pageNumber="17" pagination="121 - 143" part="6" refString="Beyer F. 1959. A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps. Nytt Magasin for Zoologi 6: 121 - 143." title="A new, bottom-living trachymedusa from the Oslofjord. Description of the species, and a general discussion of the life conditions and fauna of the fjord deeps" type="journal article" year="1959">Beyer 1959</bibRefCitation>
). The medusae move extremely slowly, and have never been observed to swim in captivity, unless they were forced to do so and, in this case, their swimming was jerky. Specimens of this species have been collected between 60350 m (
<bibRefCitation id="EFD34B7CFF83FFB1FE6303EEFDE3E6E4" author="Hesthagen I. H." box="[427,623,1507,1533]" journalOrPublisher="Norwegian Journal of Zoology" pageId="16" pageNumber="17" pagination="1 - 19" part="19" refString="Hesthagen I. H. 1971. On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer. Norwegian Journal of Zoology 19 (1): 1 - 19." title="On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer" type="journal article" year="1971">Hesthagen 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3586506FF83FFB1FF750029FAE0E574" box="[189,1388,1572,1646]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BFD368DFF83FFB1FF750029FEDAE527" blockId="16.[189,342,1572,1598]" box="[189,342,1572,1598]" pageId="16" pageNumber="17">
<heading id="D0B581E1FF83FFB1FF750029FEDAE527" bold="true" box="[189,342,1572,1598]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<emphasis id="B936EA9FFF83FFB1FF750029FEDAE527" bold="true" box="[189,342,1572,1598]" pageId="16" pageNumber="17">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BFD368DFF83FFB1FF75005EFAE0E574" blockId="16.[189,1388,1619,1646]" box="[189,1388,1619,1646]" pageId="16" pageNumber="17">
Only known from the Norwegian coast of the North Sea, between Bergen and Oslo (
<bibRefCitation id="EFD34B7CFF83FFB1FB53005EFAECE574" author="Hesthagen I. H." box="[1179,1376,1619,1645]" journalOrPublisher="Norwegian Journal of Zoology" pageId="16" pageNumber="17" pagination="1 - 19" part="19" refString="Hesthagen I. H. 1971. On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer. Norwegian Journal of Zoology 19 (1): 1 - 19." title="On the biology of the bottom-dwelling Trachymedusa Tesserogastria musculosa Beyer" type="journal article" year="1971">Hesthagen 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>