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<document id="1CD699BC5193172B0071C2241748EB46" ID-DOI="10.1080/00222933.2018.1481235" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747141" ID-ZooBank="63A31AD2-F049-42CB-A45B-557014DC286E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670765052" checkinUser="carolina" docAuthor="Taylor, Paul D., Martha, Silviu O. &amp; Gordon, Dennis P." docDate="2018" docId="03EB8789FFFC4462A1FC76C3FD7FF901" docLanguage="en" docName="JNATHIST.52.25-26.1657-1721.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222933.2018.1481235" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Euritina Canu 1900" docType="treatment" docVersion="3" lastPageNumber="1677" masterDocId="FFD2FFF1FFE84477A00C7447FF92FFBE" masterDocTitle="Synopsis of onychocellid cheilostome bryozoan genera" masterLastPageNumber="1721" masterPageNumber="1657" pageNumber="1676" updateTime="1698914560039" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B51B912DC536B2604AF3B30E95C3ED0A">Synopsis of onychocellid cheilostome bryozoan genera</mods:title>
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<mods:namePart id="FA1F9B3D3D2AFD3BF9CB37C42A8EE8FB">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="B956A2DFBCA1A0D0F2C878F0165B592C">Martha, Silviu O.</mods:namePart>
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<mods:title id="4D1B4B8B041D9EFF3E9C1AF11134F658">Journal of Natural History</mods:title>
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<mods:date id="BEAD172D99B81D0FA1F88DA0CE148A57">2018</mods:date>
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<mods:title id="028813113D093F85901F843CAB234C71">J. Nat. Hist.</mods:title>
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<subSubSection id="C3586514FFFC4463A1FC76C3FCA0FD23" box="[496,818,644,669]" pageId="20" pageNumber="1676" type="nomenclature">
<paragraph id="8BFD369FFFFC4463A1FC76C3FCA0FD23" blockId="20.[496,818,644,704]" box="[496,818,644,669]" pageId="20" pageNumber="1676">
<heading id="D0B581F3FFFC4463A1FC76C3FCA0FD23" box="[496,818,644,669]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="1676" reason="2">
Genus
<taxonomicName id="4C424D1CFFFC4463A24876C3FCA0FD23" authority="Canu, 1900" authorityName="Canu" authorityYear="1900" box="[580,818,644,669]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="1676" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFC4463A24876C3FD36FD23" bold="true" box="[580,676,644,669]" italics="true" pageId="20" pageNumber="1676">Euritina</emphasis>
<bibRefCitation id="EFD34B6EFFFC4463A2A176C3FCA0FD23" author="Canu F" box="[685,818,644,669]" pageId="20" pageNumber="1676" pagination="334 - 463" refId="ref24273" refString="Canu F. 1900. Revision des bryozoaires du Cretace figures par d' Orbigny. II. Cheilostomata. Bull Soc Geol Fr, Ser. 3 (28): 334 - 463." type="journal article" year="1900">Canu, 1900</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFFC4463A25D76E0FD40FD7E" box="[593,722,679,704]" pageId="20" pageNumber="1676" type="description">
<paragraph id="8BFD369FFFFC4463A25D76E0FD40FD7E" blockId="20.[496,818,644,704]" box="[593,722,679,704]" pageId="20" pageNumber="1676">
(
<figureCitation id="13792A1AFFFC4463A25576E0FD58FD7E" box="[601,714,679,704]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="20" pageNumber="1676">Figure 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFFC4463A0AC76ABFDEFFCF4" pageId="20" pageNumber="1676" type="type_taxon">
<paragraph id="8BFD369FFFFC4463A0AC76ABFEA6FCBB" blockId="20.[160,1155,748,842]" box="[160,308,748,773]" pageId="20" pageNumber="1676">
<emphasis id="B936EA8DFFFC4463A0AC76ABFEA6FCBB" bold="true" box="[160,308,748,773]" italics="true" pageId="20" pageNumber="1676">
<typeStatus id="54F9883DFFFC4463A0AC76ABFF4AFCBB" box="[160,216,748,773]" pageId="20" pageNumber="1676">Type</typeStatus>
species
</emphasis>
</paragraph>
<paragraph id="8BFD369FFFFC4463A0AC7757FDEFFCF4" blockId="20.[160,1155,748,842]" pageId="20" pageNumber="1676">
<taxonomicName id="4C424D1CFFFC4463A0AC7757FD97FC96" authority="d' Orbigny, 1851" authorityName="d' Orbigny" authorityYear="1851" box="[160,517,783,808]" class="Gymnolaemata" family="Coscinopleuridae" genus="Eschara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="1676" phylum="Bryozoa" rank="species" species="eurita">
<emphasis id="B936EA8DFFFC4463A0AC7757FEADFC96" box="[160,319,784,808]" italics="true" pageId="20" pageNumber="1676">Eschara eurita</emphasis>
d Orbigny, 1851
</taxonomicName>
, by original designation.
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Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et
<collectingRegion id="4986F87DFFFC4463A1EA7775FDB2FCF4" box="[486,544,818,842]" country="France" name="Pays de la Loire" pageId="20" pageNumber="1676">Loire</collectingRegion>
,
<collectingCountry id="F355760FFFFC4463A2207775FDEBFCF4" box="[556,633,818,842]" name="France" pageId="20" pageNumber="1676">France</collectingCountry>
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.
</paragraph>
</subSubSection>
<caption id="DF3D6617FFFC4463A0AC7252FE2BF901" ID-DOI="http://doi.org/10.5281/zenodo.4747165" ID-Zenodo-Dep="4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="20" pageNumber="1676" startId="20.[160,225,1557,1580]" targetBox="[163,1153,890,1527]" targetPageId="20">
<paragraph id="8BFD369FFFFC4463A0AC7252FE2BF901" blockId="20.[160,1157,1557,1727]" pageId="20" pageNumber="1676">
<emphasis id="B936EA8DFFFC4463A0AC7252FE9AF992" bold="true" box="[160,264,1557,1580]" pageId="20" pageNumber="1676">Figure 10.</emphasis>
(a
<emphasis id="B936EA8DFFFC4463A12F7252FEA2F992" box="[291,304,1557,1580]" italics="true" pageId="20" pageNumber="1676"></emphasis>
d)
<taxonomicName id="4C424D1CFFFC4463A1407252FE47F992" baseAuthorityName="d'Orbigny" baseAuthorityYear="1851" box="[332,469,1557,1580]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="1676" phylum="Bryozoa" rank="species" species="eurita">
<emphasis id="B936EA8DFFFC4463A1407252FE47F992" box="[332,469,1557,1580]" italics="true" pageId="20" pageNumber="1676">Euritina eurita</emphasis>
</taxonomicName>
(d
<emphasis id="B936EA8DFFFC4463A1FD7252FE64F992" box="[497,502,1557,1580]" italics="true" pageId="20" pageNumber="1676"></emphasis>
Orbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f)
<taxonomicName id="4C424D1CFFFC4463A40C7217FEC6F93A" authority="Canu and Bassler" authorityName="Canu and Bassler" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="20" pageNumber="1676" phylum="Bryozoa" rank="species" species="tecta">
<emphasis id="B936EA8DFFFC4463A40C7217FB11F9D9" box="[1024,1155,1616,1639]" italics="true" pageId="20" pageNumber="1676">Euritina tecta</emphasis>
Canu and Bassler
</taxonomicName>
; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm.
</paragraph>
</caption>
<subSubSection id="C3586514FFFD4462A0AC74C9FBB6FE24" pageId="21" pageNumber="1677" type="diagnosis">
<paragraph id="8BFD369FFFFD4462A0AC74C9FE86FF19" blockId="21.[160,1157,142,410]" box="[160,276,142,167]" pageId="21" pageNumber="1677">
<emphasis id="B936EA8DFFFD4462A0AC74C9FE86FF19" bold="true" box="[160,276,142,167]" italics="true" pageId="21" pageNumber="1677">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BFD369FFFFD4462A0AC74F5FBB6FE24" blockId="21.[160,1157,142,410]" pageId="21" pageNumber="1677">
Colony erect, bifoliate (
<figureCitation id="13792A1AFFFD4462A1A674F5FD84FF74" box="[426,534,178,202]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="21" pageNumber="1677">Figure 10</figureCitation>
(a)). Autozooids elongate (
<figureCitation id="13792A1AFFFD4462A34F74F5FC23FF74" box="[835,945,178,202]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="21" pageNumber="1677">Figure 10</figureCitation>
(b)), subhexagonal, rounded distally; zooidal boundaries grooved. Cryptocyst extensive, granular, sloping distally, with proximal and two proximolateral facets (
<figureCitation id="13792A1AFFFD4462A2C674B0FCA3FEB1" box="[714,817,247,271]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="21" pageNumber="1677">Figure 10</figureCitation>
(c)). Gymnocyst lacking. Opesia terminal, occupying about one-third of frontal surface; inverted pear shaped, longer than wide. Ovicells hyperstomial (
<figureCitation id="13792A1AFFFD4462A1D5757BFDD0FEEA" box="[473,578,316,340]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="21" pageNumber="1677">Figure 10</figureCitation>
(d)). Avicularia vicarious, symmetrical (
<figureCitation id="13792A1AFFFD4462A3ED757BFBD8FEEA" box="[993,1098,316,340]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="21" pageNumber="1677">Figure 10</figureCitation>
(b,c)), about half the width of an autozooid and somewhat shorter, typically budded at row bifurcations; opesia longitudinally elliptical; rostrum short, rounded, floor smooth.
</paragraph>
</subSubSection>
<subSubSection id="C3586514FFFD4462A0AC759FFD7FF901" pageId="21" pageNumber="1677" type="discussion">
<paragraph id="8BFD369FFFFD4462A0AC759FFE94FE4F" blockId="21.[160,1156,472,1606]" box="[160,262,472,497]" pageId="21" pageNumber="1677">
<emphasis id="B936EA8DFFFD4462A0AC759FFE94FE4F" bold="true" box="[160,262,472,497]" italics="true" pageId="21" pageNumber="1677">Remarks</emphasis>
</paragraph>
<paragraph id="8BFD369FFFFD4462A0AC75BCFE23FD7E" blockId="21.[160,1156,472,1606]" pageId="21" pageNumber="1677">
The family-level classification of
<taxonomicName id="4C424D1CFFFD4462A22F75BCFDE5FDAD" authorityName="Canu" authorityYear="1900" box="[547,631,507,531]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A22F75BCFDE5FDAD" box="[547,631,507,531]" italics="true" pageId="21" pageNumber="1677">Euritina</emphasis>
</taxonomicName>
has been debated, the genus having been placed in
<taxonomicName id="4C424D1CFFFD4462A1187659FE4DFD88" authorityName="Jullien" authorityYear="1888" box="[276,479,542,566]" class="Gymnolaemata" family="Aspidostomatidae" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="family">Aspidostomatidae</taxonomicName>
(e.g.
<bibRefCitation id="EFD34B6EFFFD4462A22E7659FCBAFD88" author="Canu F &amp; Bassler RS" box="[546,808,542,566]" pageId="21" pageNumber="1677" pagination="1 - 879" refId="ref24397" refString="Canu F, Bassler RS. 1920. North American Early Tertiary Bryozoa. Bull U S Natl Mus. 106: 1 - 879." type="journal article" year="1920">Canu and Bassler 1920</bibRefCitation>
),
<taxonomicName id="4C424D1CFFFD4462A3337659FC7EFD88" authorityName="Jullien" authorityYear="1882" box="[831,1004,542,566]" class="Gymnolaemata" family="Onychocellidae" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="family">Onychocellidae</taxonomicName>
(e.g.
<bibRefCitation id="EFD34B6EFFFD4462A4227659FEF3FDE6" author="Gordon DP &amp; Taylor PD" pageId="21" pageNumber="1677" pagination="83 - 97" refId="ref26071" refString="Gordon DP, Taylor PD 2005. The cheilostomatous genera of Alcide d' Orbigny - nomenclatural and taxonomic status. In: Moyano HI, Wyse Jackson PN, editors. Bryozoan Studies 2004. Proceedings of the 13 th International Bryozoology Association conference, Concepcion / Chile; January 11 - 16 2004; p. 83 - 97. London: CRC Press." type="journal article" year="2005">Gordon and Taylor 2005</bibRefCitation>
, table 2;
<bibRefCitation id="EFD34B6EFFFD4462A1D57607FD65FDE6" author="Taylor PD &amp; Martha SO" box="[473,759,576,600]" pageId="21" pageNumber="1677" pagination="19 - 31" refId="ref28446" refString="Taylor PD, Martha SO. 2017. Cenomanian cheilostome bryozoans from Devon, England. Ann Paleontol. 103: 19 - 31." type="journal article" year="2017">Taylor and Martha 2017</bibRefCitation>
) or
<taxonomicName id="4C424D1CFFFD4462A33C7607FC2CFDE6" box="[816,958,576,600]" class="Gymnolaemata" family="Calloporidae" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="family">Calloporidae</taxonomicName>
(e.g.
<bibRefCitation id="EFD34B6EFFFD4462A40B7607FEC0FDC5" author="Taylor PD &amp; McKinney FK" pageId="21" pageNumber="1677" pagination="1 - 346" refId="ref28471" refString="Taylor PD, McKinney FK. 2006. Cretaceous Bryozoa from the Campanian and Maastrichtian of the Atlantic and Gulf Coastal Plains, United States. Scripta Geol. 132: 1 - 346." type="journal article" year="2006">Taylor and McKinney 2006</bibRefCitation>
). Only broken ovicells are known in the
<typeStatus id="54F9883DFFFD4462A3487624FCE4FDC5" box="[836,886,611,635]" pageId="21" pageNumber="1677">type</typeStatus>
species (
<figureCitation id="13792A1AFFFD4462A3E57624FBCBFDC5" box="[1001,1113,611,635]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="21" pageNumber="1677">Figure 10</figureCitation>
(d)). Although these are clearly hyperstomial, it is unclear whether the ooecium is gymnocystal or cryptocyst-like.
</paragraph>
<paragraph id="8BFD369FFFFD4462A0CC768CFCBCFB18" blockId="21.[160,1156,472,1606]" pageId="21" pageNumber="1677">
Apart from the
<typeStatus id="54F9883DFFFD4462A162768CFE0DFD5D" box="[366,415,715,739]" pageId="21" pageNumber="1677">type</typeStatus>
species,
<bibRefCitation id="EFD34B6EFFFD4462A209768CFD19FD5D" author="Canu F" box="[517,651,715,739]" pageId="21" pageNumber="1677" pagination="334 - 463" refId="ref24273" refString="Canu F. 1900. Revision des bryozoaires du Cretace figures par d' Orbigny. II. Cheilostomata. Bull Soc Geol Fr, Ser. 3 (28): 334 - 463." type="journal article" year="1900">Canu (1900)</bibRefCitation>
assigned to the genus two other Cretaceous species (
<emphasis id="B936EA8DFFFD4462A10E76A9FE01FCB8" box="[258,403,750,774]" italics="true" pageId="21" pageNumber="1677">Eschara delia</emphasis>
d Orbigny, 1851 and
<taxonomicName id="4C424D1CFFFD4462A29E76A9FC29FCB8" authority="Canu, 1900" authorityName="Canu" authorityYear="1900" box="[658,955,750,774]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="species" species="welshi">
<emphasis id="B936EA8DFFFD4462A29E76A9FCA3FCB8" box="[658,817,750,774]" italics="true" pageId="21" pageNumber="1677">Euritina welshi</emphasis>
<bibRefCitation id="EFD34B6EFFFD4462A33076A9FC29FCB8" author="Canu F" box="[828,955,750,774]" pageId="21" pageNumber="1677" pagination="334 - 463" refId="ref24273" refString="Canu F. 1900. Revision des bryozoaires du Cretace figures par d' Orbigny. II. Cheilostomata. Bull Soc Geol Fr, Ser. 3 (28): 334 - 463." type="journal article" year="1900">Canu, 1900</bibRefCitation>
</taxonomicName>
) and one Recent species (
<taxonomicName id="4C424D1CFFFD4462A0F37757FD09FC96" authority="Busk, 1884" authorityName="Busk" authorityYear="1884" box="[255,667,784,808]" class="Gymnolaemata" family="Calloporidae" genus="Amphiblestrum" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="species" species="papillatum">
<emphasis id="B936EA8DFFFD4462A0F37757FD8FFC96" box="[255,541,784,808]" italics="true" pageId="21" pageNumber="1677">Amphiblestrum papillatum</emphasis>
<bibRefCitation id="EFD34B6EFFFD4462A2287757FD09FC96" author="Busk G" box="[548,667,784,808]" pageId="21" pageNumber="1677" pagination="1 - 216" refId="ref24224" refString="Busk G. 1884. Report on the Polyzoa collected by H. M. S. Challenger during the years 1873 - 1876. Part I. - the Cheilostomata. Rep Sci Res Voyage HMS &quot; Challenger &quot;, Zool. 10: 1 - 216." type="journal article" year="1884">Busk, 1884</bibRefCitation>
</taxonomicName>
). The Recent species, which has tiny adventitious avicularia (
<bibRefCitation id="EFD34B6EFFFD4462A1537774FE43FCF5" author="Busk G" box="[351,465,819,843]" pageId="21" pageNumber="1677" pagination="1 - 216" refId="ref24224" refString="Busk G. 1884. Report on the Polyzoa collected by H. M. S. Challenger during the years 1873 - 1876. Part I. - the Cheilostomata. Rep Sci Res Voyage HMS &quot; Challenger &quot;, Zool. 10: 1 - 216." type="journal article" year="1884">Busk 1884</bibRefCitation>
, p. 33, fig. 1), was not listed by
<bibRefCitation id="EFD34B6EFFFD4462A3207774FBBEFCF5" author="Canu F &amp; Bassler RS" box="[812,1068,819,843]" pageId="21" pageNumber="1677" pagination="1 - 879" refId="ref24397" refString="Canu F, Bassler RS. 1920. North American Early Tertiary Bryozoa. Bull U S Natl Mus. 106: 1 - 879." type="journal article" year="1920">Canu and Bassler (1920</bibRefCitation>
, p. 257) when they redescribed
<taxonomicName id="4C424D1CFFFD4462A1A47711FE68FCD0" authorityName="Canu" authorityYear="1900" box="[424,506,854,878]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A1A47711FE68FCD0" box="[424,506,854,878]" italics="true" pageId="21" pageNumber="1677">Euritina</emphasis>
</taxonomicName>
and seems not to belong to this genus.
<bibRefCitation id="EFD34B6EFFFD4462A3C97711FF77FC2E" author="Canu F &amp; Bassler RS" pageId="21" pageNumber="1677" pagination="1 - 879" refId="ref24397" refString="Canu F, Bassler RS. 1920. North American Early Tertiary Bryozoa. Bull U S Natl Mus. 106: 1 - 879." type="journal article" year="1920">Canu and Bassler (1920)</bibRefCitation>
listed several additional Paleocene species of
<taxonomicName id="4C424D1CFFFD4462A2ED773FFCA1FC2E" authorityName="Canu" authorityYear="1900" box="[737,819,888,912]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A2ED773FFCA1FC2E" box="[737,819,888,912]" italics="true" pageId="21" pageNumber="1677">Euritina</emphasis>
</taxonomicName>
, among which is
<taxonomicName id="4C424D1CFFFD4462A3FA773FFEFFFC0D" authority="Canu and Bassler, 1920" authorityName="Canu and Bassler" authorityYear="1920" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="species" species="tecta">
<emphasis id="B936EA8DFFFD4462A3FA773FFBD1FC2E" box="[1014,1091,888,912]" italics="true" pageId="21" pageNumber="1677">E. tecta</emphasis>
<bibRefCitation id="EFD34B6EFFFD4462A446773FFEFFFC0D" author="Canu F &amp; Bassler RS" pageId="21" pageNumber="1677" pagination="1 - 879" refId="ref24397" refString="Canu F, Bassler RS. 1920. North American Early Tertiary Bryozoa. Bull U S Natl Mus. 106: 1 - 879." type="journal article" year="1920">Canu and Bassler, 1920</bibRefCitation>
</taxonomicName>
from the Clayton Formation of
<collectingRegion id="4986F87DFFFD4462A2F977DCFCC4FC0D" box="[757,854,923,947]" country="United States of America" name="Arkansas" pageId="21" pageNumber="1677">Arkansas</collectingRegion>
(
<figureCitation id="13792A1AFFFD4462A36077DCFC4EFC0D" box="[876,988,923,947]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="21" pageNumber="1677">Figure 10</figureCitation>
(e,f)). Scanning electron microscope images of the
<typeStatus id="54F9883DFFFD4462A24477F9FD14FC68" box="[584,646,958,982]" pageId="21" pageNumber="1677">types</typeStatus>
, kindly supplied by JoAnn Sanner (USNM), show clearly the gymnocystal ooecium (
<figureCitation id="13792A1AFFFD4462A26677A7FD47FC46" box="[618,725,992,1016]" captionStart="Figure 10" captionStartId="20.[160,225,1557,1580]" captionTargetBox="[163,1153,890,1527]" captionTargetId="figure-168@20.[163,1153,890,1527]" captionTargetPageId="20" captionText="Figure 10. (ad) Euritina eurita (dOrbigny); lectotype, MNHN R61484; Cretaceous, Turonian, Sainte- Maure-de-Touraine, Indre-et Loire, France; (a) bifoliate frond; (b) autozooids and slightly smaller avicularia; (c) autozooids and avicularium (right of centre); (d) broken ovicells. (e,f) Euritina tecta Canu and Bassler; syntypes, USNM 63807; Paleocene, Clayton Formation, Mabelvale, Little Rock, Arkansas; (e) overgrowth; (f) autozooids, some ovicellate (e.g. arrow). Scale bars: a = 1 mm; b, e, f = 500 µm; c, d = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4747165" httpUri="https://zenodo.org/record/4747165/files/figure.png" pageId="21" pageNumber="1677">Figure 10</figureCitation>
(f)) of the hyperstomial ovicells in this Danian species. Two new species of multilamellar encrusters were assigned to
<taxonomicName id="4C424D1CFFFD4462A4027044FBF2FBA5" authorityName="Canu" authorityYear="1900" box="[1038,1120,1027,1051]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A4027044FBF2FBA5" box="[1038,1120,1027,1051]" italics="true" pageId="21" pageNumber="1677">Euritina</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFD34B6EFFFD4462A0AC7061FE4BFB80" author="Taylor PD &amp; McKinney FK" box="[160,473,1062,1086]" pageId="21" pageNumber="1677" pagination="1 - 346" refId="ref28471" refString="Taylor PD, McKinney FK. 2006. Cretaceous Bryozoa from the Campanian and Maastrichtian of the Atlantic and Gulf Coastal Plains, United States. Scripta Geol. 132: 1 - 346." type="journal article" year="2006">Taylor and McKinney (2006)</bibRefCitation>
from the Maastrichtian of the south-eastern
<collectingCountry id="F355760FFFFD4462A3D97061FB94FB80" box="[981,1030,1062,1086]" name="United States of America" pageId="21" pageNumber="1677">USA</collectingCountry>
. Both have facetted cryptocysts but
<taxonomicName id="4C424D1CFFFD4462A1BF700FFC25FBDE" authority="Taylor and McKinney, 2006" authorityName="Taylor and McKinney" authorityYear="2006" box="[435,951,1096,1120]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="species" species="metapolymorpha">
<emphasis id="B936EA8DFFFD4462A1BF700FFD16FBDE" box="[435,644,1096,1120]" italics="true" pageId="21" pageNumber="1677">E. metapolymorpha</emphasis>
<bibRefCitation id="EFD34B6EFFFD4462A280700FFC25FBDE" author="Taylor PD &amp; McKinney FK" box="[652,951,1096,1120]" pageId="21" pageNumber="1677" pagination="1 - 346" refId="ref28471" refString="Taylor PD, McKinney FK. 2006. Cretaceous Bryozoa from the Campanian and Maastrichtian of the Atlantic and Gulf Coastal Plains, United States. Scripta Geol. 132: 1 - 346." type="journal article" year="2006">Taylor and McKinney, 2006</bibRefCitation>
</taxonomicName>
is unusual for this genus (and for
<taxonomicName id="4C424D1CFFFD4462A143702CFE68FB3D" authorityName="Jullien" authorityYear="1882" box="[335,506,1131,1155]" class="Gymnolaemata" family="Onychocellidae" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="family">Onychocellidae</taxonomicName>
) in that the autozooids and avicularia have a narrow but distinct gymnocyst along their proximal and lateral edges.
</paragraph>
<paragraph id="8BFD369FFFFD4462A0CC70F7FEE1F9F8" blockId="21.[160,1156,472,1606]" pageId="21" pageNumber="1677">
<bibRefCitation id="EFD34B6EFFFD4462A0CC70F7FEFEFB76" author="Osburn RC" box="[192,364,1200,1224]" pageId="21" pageNumber="1677" pagination="1 - 269" refId="ref27630" refString="Osburn RC. 1950. Bryozoa of the Pacific coast of America. Part 1, Cheilostomata - Anasca. Rep Allan Hancock Pac Exped. 14: 1 - 269." type="journal article" year="1950">Osburn (1950)</bibRefCitation>
referred the extant species
<taxonomicName id="4C424D1CFFFD4462A2D070F7FBC9FB76" authority="(Osburn, 1950)" baseAuthorityName="Osburn" baseAuthorityYear="1950" box="[732,1115,1200,1224]" class="Gymnolaemata" family="Microporidae" genus="Reussina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="species" species="arctica">
<emphasis id="B936EA8DFFFD4462A2D070F7FC0AFB76" box="[732,920,1200,1224]" italics="true" pageId="21" pageNumber="1677">Reussina arctica</emphasis>
(
<bibRefCitation id="EFD34B6EFFFD4462A3A370F7FBC0FB76" author="Osburn RC" box="[943,1106,1200,1224]" pageId="21" pageNumber="1677" pagination="1 - 269" refId="ref27630" refString="Osburn RC. 1950. Bryozoa of the Pacific coast of America. Part 1, Cheilostomata - Anasca. Rep Allan Hancock Pac Exped. 14: 1 - 269." type="journal article" year="1950">Osburn, 1950</bibRefCitation>
)
</taxonomicName>
to
<taxonomicName id="4C424D1CFFFD4462A0AC7094FF64FB55" authorityName="Canu" authorityYear="1900" box="[160,246,1235,1259]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A0AC7094FF64FB55" box="[160,246,1235,1259]" italics="true" pageId="21" pageNumber="1677">Euritina</emphasis>
</taxonomicName>
but this species has oral spine bases and the cryptocyst is ringed by large pores (?opesiules) (Gordon 2009, fig. 1AD), outweighing similarities with the
<typeStatus id="54F9883DFFFD4462A45C70B1FB11FAB0" box="[1104,1155,1270,1294]" pageId="21" pageNumber="1677">type</typeStatus>
species of the genus in the facetted structure of the cryptocyst. It is now assigned to
<taxonomicName id="4C424D1CFFFD4462A0AC717CFE8BFAED" authorityName="Gordon" authorityYear="2009" box="[160,281,1339,1363]" class="Gymnolaemata" family="Microporidae" genus="Reussinella" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A0AC717CFE8BFAED" box="[160,281,1339,1363]" italics="true" pageId="21" pageNumber="1677">Reussinella</emphasis>
</taxonomicName>
, a replacement name introduced by Gordon (2009) for
<taxonomicName id="4C424D1CFFFD4462A3C3717CFF4AFAC8" authority="Kluge, 1962" authorityName="Kluge" authorityYear="1962" class="Gymnolaemata" family="Microporidae" genus="Reussina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A3C3717CFBBDFAED" box="[975,1071,1339,1363]" italics="true" pageId="21" pageNumber="1677">Reussina</emphasis>
<bibRefCitation id="EFD34B6EFFFD4462A431717CFF4AFAC8" author="Kluge GA" pageId="21" pageNumber="1677" pagination="1 - 584" refId="ref26713" refString="Kluge GA. 1962. Мsaнki сiviрныk мoрСССР [Bryozoans of the northern seas of the USSR]. Opred Faune SSSR. 76: 1 - 584." type="journal article" year="1962">Kluge, 1962</bibRefCitation>
</taxonomicName>
, a junior homonym of
<taxonomicName id="4C424D1CFFFD4462A1F77119FC9AFAC8" authority="Neviani, 1896" authorityName="Neviani" authorityYear="1896" box="[507,776,1374,1398]" class="Gymnolaemata" family="Microporidae" genus="Reussina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A1F77119FDC9FAC8" box="[507,603,1374,1398]" italics="true" pageId="21" pageNumber="1677">Reussina</emphasis>
<bibRefCitation id="EFD34B6EFFFD4462A2647119FC9AFAC8" author="Neviani A" box="[616,776,1374,1398]" pageId="21" pageNumber="1677" pagination="79 - 100" refId="ref27492" refString="Neviani A. 1896. Briozoi fossili della Farnesina e Monte Mario presso Roma. Paleontogr Ital. 1: 79 - 100." type="journal article" year="1896">Neviani, 1896</bibRefCitation>
</taxonomicName>
, who placed it provisionally in
<taxonomicName id="4C424D1CFFFD4462A0AC71C7FEA9FA26" authorityName="Gray" authorityYear="1848" box="[160,315,1408,1432]" class="Gymnolaemata" family="Microporidae" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="family">Microporidae</taxonomicName>
. With the exclusion of this species from
<taxonomicName id="4C424D1CFFFD4462A31871C7FCF8FA26" authorityName="Canu" authorityYear="1900" box="[788,874,1408,1432]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A31871C7FCF8FA26" box="[788,874,1408,1432]" italics="true" pageId="21" pageNumber="1677">Euritina</emphasis>
</taxonomicName>
, the upper range of the genus becomes Thanetian, the youngest recorded species being
<taxonomicName id="4C424D1CFFFD4462A3A571E4FEBBFA60" authority="(Gabb and Horn, 1862)" baseAuthorityName="Gabb and Horn" baseAuthorityYear="1862" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="species" species="torta">
<emphasis id="B936EA8DFFFD4462A3A571E4FC6EFA05" box="[937,1020,1443,1467]" italics="true" pageId="21" pageNumber="1677">E. torta</emphasis>
(
<bibRefCitation id="EFD34B6EFFFD4462A40171E4FEB3FA60" author="Gabb WM &amp; Horn GH" pageId="21" pageNumber="1677" pagination="111 - 179" refId="ref25825" refString="Gabb WM, Horn GH. 1862. Monograph of the fossil Polyzoa of the Secondary and Tertiary formations of North America. J Acad Nat Sci Phila. 5: 111 - 179." type="journal article" year="1862">Gabb and Horn, 1862</bibRefCitation>
)
</taxonomicName>
from the Vincentown Limesand of
<collectingRegion id="4986F87DFFFD4462A2D97181FCCFFA60" box="[725,861,1478,1502]" country="United States of America" name="New Jersey" pageId="21" pageNumber="1677">New Jersey</collectingRegion>
. Provisionally, the oldest known species is the early Cenomanian
<taxonomicName id="4C424D1CFFFD4462A29271AFFBD4F9BE" authority="(Roemer, 1840)" baseAuthorityName="Roemer" baseAuthorityYear="1840" box="[670,1094,1512,1536]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="species" species="denticulata">
<emphasis id="B936EA8DFFFD4462A29271AFFC12F9BE" box="[670,896,1512,1536]" italics="true" pageId="21" pageNumber="1677">Euritina denticulata</emphasis>
(
<bibRefCitation id="EFD34B6EFFFD4462A39B71AFFBACF9BE" author="Roemer FA" box="[919,1086,1512,1536]" pageId="21" pageNumber="1677" refId="ref27937" refString="Roemer FA. 1840. Die Versteinerungen des norddeutschen Kreidegebirges. Hannover: Hahn' sche Hofbuchhandlung." type="book" year="1840">Roemer, 1840</bibRefCitation>
)
</taxonomicName>
(see
<bibRefCitation id="EFD34B6EFFFD4462A0AC724CFE51F99D" author="Taylor PD &amp; Martha SO" box="[160,451,1547,1571]" pageId="21" pageNumber="1677" pagination="19 - 31" refId="ref28446" refString="Taylor PD, Martha SO. 2017. Cenomanian cheilostome bryozoans from Devon, England. Ann Paleontol. 103: 19 - 31." type="journal article" year="2017">Taylor and Martha 2017</bibRefCitation>
), a species which, however, lacks the facetted cryptocyst typical of
<taxonomicName id="4C424D1CFFFD4462A1147269FEFCF9F8" authorityName="Canu" authorityYear="1900" box="[280,366,1582,1606]" class="Gymnolaemata" family="Aspidostomatidae" genus="Euritina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1677" phylum="Bryozoa" rank="genus">
<emphasis id="B936EA8DFFFD4462A1147269FEFCF9F8" box="[280,366,1582,1606]" italics="true" pageId="21" pageNumber="1677">Euritina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFD369FFFFD4462A0AC72C3FF79F923" blockId="21.[160,749,1668,1727]" box="[160,235,1668,1693]" pageId="21" pageNumber="1677">
<emphasis id="B936EA8DFFFD4462A0AC72C3FF79F923" bold="true" box="[160,235,1668,1693]" italics="true" pageId="21" pageNumber="1677">Range</emphasis>
</paragraph>
<paragraph id="8BFD369FFFFD4462A0AC72E0FD7FF901" blockId="21.[160,749,1668,1727]" box="[160,749,1703,1727]" pageId="21" pageNumber="1677">Cretaceous (Cenomanian) to Paleocene (Thanetian).</paragraph>
</subSubSection>
</treatment>
</document>