221 lines
36 KiB
XML
221 lines
36 KiB
XML
<document id="63785C58E0E346D1B94D577160487221" ID-DOI="10.1080/00222933.2018.1481235" ID-ISSN="1464-5262" ID-Zenodo-Dep="4747141" ID-ZooBank="63A31AD2-F049-42CB-A45B-557014DC286E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620670765052" checkinUser="carolina" docAuthor="Taylor, Paul D., Martha, Silviu O. & Gordon, Dennis P." docDate="2018" docId="03EB8789FFF2446CA1B0762EFBBBF915" docLanguage="en" docName="JNATHIST.52.25-26.1657-1721.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222933.2018.1481235" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Hoplitaechmella Voigt 1949" docType="treatment" docVersion="4" lastPageNumber="1683" masterDocId="FFD2FFF1FFE84477A00C7447FF92FFBE" masterDocTitle="Synopsis of ‘ onychocellid’ cheilostome bryozoan genera" masterLastPageNumber="1721" masterPageNumber="1657" pageNumber="1682" updateTime="1698914560039" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="60B62125B18CB8D5D468AEAD91EB93BB">
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<mods:title id="F6082A9F445AEBB805C9B3B5B126BF89">Synopsis of ‘ onychocellid’ cheilostome bryozoan genera</mods:title>
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<mods:namePart id="6BF8DC135DC6D4AA4C30D5D0514402CB">Taylor, Paul D.</mods:namePart>
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<mods:namePart id="6B5A98CA964EA222E0438504E6676F3F">Martha, Silviu O.</mods:namePart>
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<mods:namePart id="03E02F01A737ABA176D2DE680D8F7C00">Gordon, Dennis P.</mods:namePart>
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<mods:typeOfResource id="5F6FC3BD1EA2A1088EB675458804321B">text</mods:typeOfResource>
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<mods:title id="EE986FF4E7A51C8BA1F045572A4CA0BB">Journal of Natural History</mods:title>
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<mods:part id="F9C7EE132A7ADC2E6AA9112C297AA579">
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<mods:date id="3C02C53E96E5C48129A0E901A1FE2E45">2018</mods:date>
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<mods:title id="EDB874E57A7155FA2356B695583AC29B">J. Nat. Hist.</mods:title>
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<mods:number id="4DAAFBFE694FFD73B6E3DA41A74597EA">2018-07-03</mods:number>
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<mods:number id="D9688D1D7104A18EBBD7546BA17C0241">52</mods:number>
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<mods:number id="76FE42011F0F1015BDCF9E3203C8DE52">25 - 26</mods:number>
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<mods:start id="D98954EB6335B541F7C1ABCFFA346AB2">1657</mods:start>
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<mods:url id="65846C9422702EA503DA6BD5B651C60C">http://dx.doi.org/10.1080/00222933.2018.1481235</mods:url>
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<mods:identifier id="CB105733E04819425A49C996F557ACA0" type="Zenodo-Dep">4747141</mods:identifier>
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<treatment id="03EB8789FFF2446CA1B0762EFBBBF915" ID-DOI="http://doi.org/10.5281/zenodo.4747854" ID-Zenodo-Dep="4747854" LSID="urn:lsid:plazi:treatment:03EB8789FFF2446CA1B0762EFBBBF915" httpUri="http://treatment.plazi.org/id/03EB8789FFF2446CA1B0762EFBBBF915" lastPageId="27" lastPageNumber="1683" pageId="26" pageNumber="1682">
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<subSubSection id="C3586514FFF2446DA1B0762EFCF5FD3C" box="[444,871,617,642]" pageId="26" pageNumber="1682" type="nomenclature">
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<paragraph id="8BFD369FFFF2446DA1B0762EFCF5FD3C" blockId="26.[444,871,617,676]" box="[444,871,617,642]" pageId="26" pageNumber="1682">
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<heading id="D0B581F3FFF2446DA1B0762EFCF5FD3C" box="[444,871,617,642]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="1682" reason="2">
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Genus
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<taxonomicName id="4C424D1CFFF2446DA203762EFCF5FD3C" ID-CoL="85FG7" authority="Voigt, 1949" authorityName="Voigt" authorityYear="1949" box="[527,871,617,642]" class="Gymnolaemata" family="Onychocellidae" genus="Hoplitaechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="26" pageNumber="1682" phylum="Bryozoa" rank="genus">
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<emphasis id="B936EA8DFFF2446DA203762EFD45FD3C" bold="true" box="[527,727,617,642]" italics="true" pageId="26" pageNumber="1682">Hoplitaechmella</emphasis>
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<bibRefCitation id="EFD34B6EFFF2446DA2EC762EFCF5FD3C" author="Voigt E" box="[736,871,617,642]" pageId="26" pageNumber="1682" pagination="1 - 49" refId="ref28686" refString="Voigt E. 1949. Cheilostome Bryozoen aus der Quadratenkreide Nordwestdeutschlands. Mitt Geol Staatsinst Hamburg. 19: 1 - 49." type="journal article" year="1949">Voigt, 1949</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3586514FFF2446DA25D76CCFD40FD1A" box="[593,722,651,676]" pageId="26" pageNumber="1682" type="description">
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<paragraph id="8BFD369FFFF2446DA25D76CCFD40FD1A" blockId="26.[444,871,617,676]" box="[593,722,651,676]" pageId="26" pageNumber="1682">
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||
(
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<figureCitation id="13792A1AFFF2446DA25576CCFD58FD1A" box="[601,714,651,676]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3586514FFF2446DA0AC7693FB13FC03" pageId="26" pageNumber="1682" type="type_taxon">
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<paragraph id="8BFD369FFFF2446DA0AC7693FEA6FD53" blockId="26.[160,1155,724,957]" box="[160,308,724,749]" pageId="26" pageNumber="1682">
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<emphasis id="B936EA8DFFF2446DA0AC7693FEA6FD53" bold="true" box="[160,308,724,749]" italics="true" pageId="26" pageNumber="1682">
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<typeStatus id="54F9883DFFF2446DA0AC7693FF4AFD53" box="[160,216,724,749]" pageId="26" pageNumber="1682">Type</typeStatus>
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species
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</emphasis>
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</paragraph>
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<paragraph id="8BFD369FFFF2446DA0AC76BFFB13FC03" blockId="26.[160,1155,724,957]" pageId="26" pageNumber="1682">
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<taxonomicName id="4C424D1CFFF2446DA0AC76BFFD12FCAE" authority="von Hagenow, 1839" authorityName="von Hagenow" authorityYear="1839" box="[160,640,760,784]" class="Gymnolaemata" family="Celleporidae" genus="Cellepora" kingdom="Animalia" order="Cheilostomatida" pageId="26" pageNumber="1682" phylum="Bryozoa" rank="species" species="vespertilio">
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<emphasis id="B936EA8DFFF2446DA0AC76BFFEEDFCAE" box="[160,383,760,784]" italics="true" pageId="26" pageNumber="1682">Cellepora vespertilio</emphasis>
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von Hagenow, 1839
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</taxonomicName>
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, by original designation.
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<materialsCitation id="3B2A3CC2FFF2446DA3B276BFFCA1FC8C" collectorName="Cretaceous & Maastrichtian" country="Germany" location="Rugen" pageId="26" pageNumber="1682" specimenCount="1" stateProvince="Mecklenburg-Vorpommern" typeStatus="holotype">
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<collectorName id="26B75349FFF2446DA3B276BFFBA9FCAE" box="[958,1083,760,784]" pageId="26" pageNumber="1682">Cretaceous</collectorName>
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, early
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<collectorName id="26B75349FFF2446DA0AC775DFEA5FC8C" box="[160,311,794,818]" pageId="26" pageNumber="1682">Maastrichtian</collectorName>
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,
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<location id="8E9D6044FFF2446DA14D775DFE19FC8C" LSID="urn:lsid:plazi:treatment:03EB8789FFF2446CA1B0762EFBBBF915:8E9D6044FFF2446DA14D775DFE19FC8C" box="[321,395,794,818]" country="Germany" name="Rugen" pageId="26" pageNumber="1682" stateProvince="Mecklenburg-Vorpommern">Rügen</location>
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,
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<collectingRegion id="4986F87DFFF2446DA198775DFD50FC8C" box="[404,706,794,818]" country="Germany" name="Mecklenburg-Vorpommern" pageId="26" pageNumber="1682">Mecklenburg-Vorpommern</collectingRegion>
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,
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<collectingCountry id="F355760FFFF2446DA2C0775DFCA1FC8C" box="[716,819,794,818]" name="Germany" pageId="26" pageNumber="1682">Germany</collectingCountry>
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</materialsCitation>
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. The type material of this species has been lost (see
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<bibRefCitation id="EFD34B6EFFF2446DA1A4777AFDB0FCEB" author="Voigt E" box="[424,546,829,853]" pageId="26" pageNumber="1682" pagination="1 - 80" refId="ref28800" refString="Voigt E. 1959. Revision der von F. v. Hagenow 1838 - 1850 aus der Schreibkreide von Rugen veroffentlichten Bryozoen. Geol. 8: 1 - 80." type="journal article" year="1959">Voigt 1959</bibRefCitation>
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) and no
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<typeStatus id="54F9883DFFF2446DA280777AFD75FCEB" box="[652,743,829,853]" pageId="26" pageNumber="1682" type="neotype">neotype</typeStatus>
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has been designated. Although von Hagenow (1839) did not illustrate his species,
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<specimenCount id="9D44FD16FFF2446DA2A87727FCD9FCC6" box="[676,843,864,888]" pageId="26" pageNumber="1682" type="generic">two specimens</specimenCount>
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from the type locality were figured in
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<bibRefCitation id="EFD34B6EFFF2446DA10377C5FE3FFC24" author="Marsson T" box="[271,429,898,922]" pageId="26" pageNumber="1682" pagination="3 - 112" refId="ref27283" refString="Marsson T. 1887. Die Bryozoen der weissen Schreibkreide der Insel Rugen. Palaontol Abh. 4: 3 - 112." type="journal article" year="1887">Marsson (1887</bibRefCitation>
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, p. 9, fig. 15).
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<materialsCitation id="3B2A3CC2FFF2446DA23277C5FBEFFC03" collectorName="No" country="Germany" location="Rugen" municipality="Voigt Collection" pageId="26" pageNumber="1682" specimenCount="1" stateProvince="Mecklenburg-Vorpommern" typeStatus="holotype">
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<collectorName id="26B75349FFF2446DA23277C5FDCCFC24" box="[574,606,898,922]" pageId="26" pageNumber="1682">No</collectorName>
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further material of this species has been described from Rügen but we here figure a
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<location id="8E9D6044FFF2446DA21E77E2FDCBFC03" LSID="urn:lsid:plazi:treatment:03EB8789FFF2446CA1B0762EFBBBF915:8E9D6044FFF2446DA21E77E2FDCBFC03" box="[530,601,933,957]" country="Germany" municipality="Voigt Collection" name="Rugen" pageId="26" pageNumber="1682" stateProvince="Mecklenburg-Vorpommern">Rügen</location>
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specimen in the
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<collectingMunicipality id="6B99ACE5FFF2446DA31477E2FC55FC03" box="[792,967,933,957]" pageId="26" pageNumber="1682">Voigt Collection</collectingMunicipality>
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(
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<figureCitation id="13792A1AFFF2446DA3D977E2FBAFFC03" box="[981,1085,933,957]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
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(g–i))
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3586514FFF2446DA0AC77AAFC7CFA18" pageId="26" pageNumber="1682" type="diagnosis">
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<paragraph id="8BFD369FFFF2446DA0AC77AAFE86FBB8" blockId="26.[160,1157,1005,1446]" box="[160,276,1005,1030]" pageId="26" pageNumber="1682">
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<emphasis id="B936EA8DFFF2446DA0AC77AAFE86FBB8" bold="true" box="[160,276,1005,1030]" italics="true" pageId="26" pageNumber="1682">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BFD369FFFF2446DA0AC7056FC7CFA18" blockId="26.[160,1157,1005,1446]" pageId="26" pageNumber="1682">
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Colony encrusting, zooidal boundaries grooved (
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<figureCitation id="13792A1AFFF2446DA2DA7056FCD6FB97" box="[726,836,1041,1065]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
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(a)). Pore chambers present (
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<figureCitation id="13792A1AFFF2446DA0AB7073FEB6FBF2" box="[167,292,1076,1100]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
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(e)). Autozooids rounded hexagonal. Cryptocyst extensive, granular. Gymnocyst variably developed proximally and laterally. Opesia D shaped to bell shaped (
|
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<figureCitation id="13792A1AFFF2446DA0AB703EFE80FB2F" box="[167,274,1145,1169]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
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(b)), small, about one-quarter of zooid length, terminal, lacking opesiular indentations and constrictions; oral spines present, usually numbering six; cryptocyst granular. Ovicells hyperstomial (
|
||
<figureCitation id="13792A1AFFF2446DA19170F9FD9BFB68" box="[413,521,1214,1238]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
|
||
(h), the ooecium not granular and apparently of exterior wall (
|
||
<figureCitation id="13792A1AFFF2446DA0D770A6FED5FB47" box="[219,327,1249,1273]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
|
||
(i)). Intramural buds common (
|
||
<figureCitation id="13792A1AFFF2446DA29A70A6FD6DFB47" box="[662,767,1249,1273]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
|
||
(c,h)). Avicularia of two main
|
||
<typeStatus id="54F9883DFFF2446DA44E70A6FB12FB47" box="[1090,1152,1249,1273]" pageId="26" pageNumber="1682">types</typeStatus>
|
||
: small interzooidal avicularia often located distolaterally of autozooids, symmetrical, opesia long, rostrum short (
|
||
<figureCitation id="13792A1AFFF2446DA1997161FD97FA80" box="[405,517,1318,1342]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
|
||
(b)); large vicarious avicularia, rare, symmetrical, opesia either the same size as those of the interzooidal avicularia (
|
||
<figureCitation id="13792A1AFFF2446DA354710EFC54FADF" box="[856,966,1353,1377]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
|
||
(g)), or long and distally tapering, rostrum extending over cryptocyst of distal autozooid, condyles developed (
|
||
<figureCitation id="13792A1AFFF2446DA0E571C9FEC6FA18" box="[233,340,1422,1446]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
|
||
(d)). Kenozooids with central foramen present (
|
||
<figureCitation id="13792A1AFFF2446DA36E71C9FC5FFA18" box="[866,973,1422,1446]" captionStart="Figure 14" captionStartId="27.[160,225,1131,1154]" captionTargetBox="[163,1153,142,1100]" captionTargetId="figure-76@27.[163,1153,142,1100]" captionTargetPageId="27" captionText="Figure 14. Hoplitaechmella vespertilio (von Hagenow).(a–f) NHMUK D15590;Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g–i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c–h = 200 µm; b, i = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="26" pageNumber="1682">Figure 14</figureCitation>
|
||
(g).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3586514FFF2446CA0AC7191FBBBF915" lastPageId="27" lastPageNumber="1683" pageId="26" pageNumber="1682" type="discussion">
|
||
<paragraph id="8BFD369FFFF2446DA0AC7191FE94FA51" blockId="26.[160,1156,1494,1727]" box="[160,262,1494,1519]" pageId="26" pageNumber="1682">
|
||
<emphasis id="B936EA8DFFF2446DA0AC7191FE94FA51" bold="true" box="[160,262,1494,1519]" italics="true" pageId="26" pageNumber="1682">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFD369FFFF2446DA0AC71BDFDCEF901" blockId="26.[160,1156,1494,1727]" pageId="26" pageNumber="1682">
|
||
Aside from the
|
||
<typeStatus id="54F9883DFFF2446DA15771BDFE1FF9AC" box="[347,397,1530,1554]" pageId="26" pageNumber="1682">type</typeStatus>
|
||
species,
|
||
<bibRefCitation id="EFD34B6EFFF2446DA1F071BDFD1BF9AC" author="Voigt E" box="[508,649,1530,1554]" pageId="26" pageNumber="1682" pagination="1 - 49" refId="ref28686" refString="Voigt E. 1949. Cheilostome Bryozoen aus der Quadratenkreide Nordwestdeutschlands. Mitt Geol Staatsinst Hamburg. 19: 1 - 49." type="journal article" year="1949">Voigt (1949)</bibRefCitation>
|
||
assigned four other species from the Late Cretaceous to
|
||
<taxonomicName id="4C424D1CFFF2446DA14E725AFE60F98B" authorityName="Voigt" authorityYear="1949" box="[322,498,1565,1589]" class="Gymnolaemata" family="Onychocellidae" genus="Hoplitaechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="26" pageNumber="1682" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B936EA8DFFF2446DA14E725AFE60F98B" box="[322,498,1565,1589]" italics="true" pageId="26" pageNumber="1682">Hoplitaechmella</emphasis>
|
||
</taxonomicName>
|
||
when he erected the genus. The dimorphic avicularia and oral spine bases are characteristic, as is the presence of pore chambers as well as autozooidal frontal walls that may have a narrow but appreciable marginal gymnocyst. The latter two characters in particular raise some doubts about the classification of
|
||
<taxonomicName id="4C424D1CFFF2446DA0AC72E0FEC2F901" authorityName="Voigt" authorityYear="1949" box="[160,336,1703,1727]" class="Gymnolaemata" family="Onychocellidae" genus="Hoplitaechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="26" pageNumber="1682" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B936EA8DFFF2446DA0AC72E0FEC2F901" box="[160,336,1703,1727]" italics="true" pageId="26" pageNumber="1682">Hoplitaechmella</emphasis>
|
||
</taxonomicName>
|
||
within
|
||
<taxonomicName id="4C424D1CFFF2446DA1A572E0FDCAF901" authorityName="Jullien" authorityYear="1882" box="[425,600,1703,1727]" class="Gymnolaemata" family="Onychocellidae" kingdom="Animalia" order="Cheilostomatida" pageId="26" pageNumber="1682" phylum="Bryozoa" rank="family">Onychocellidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF3D6617FFF3446CA0AC702CFC54FAF1" ID-DOI="http://doi.org/10.5281/zenodo.4747173" ID-Zenodo-Dep="4747173" httpUri="https://zenodo.org/record/4747173/files/figure.png" pageId="27" pageNumber="1683" startId="27.[160,225,1131,1154]" targetBox="[163,1153,142,1100]" targetPageId="27">
|
||
<paragraph id="8BFD369FFFF3446CA0AC702CFC54FAF1" blockId="27.[160,1156,1131,1359]" pageId="27" pageNumber="1683">
|
||
<emphasis id="B936EA8DFFF3446CA0AC702CFE95FB3C" bold="true" box="[160,263,1131,1154]" pageId="27" pageNumber="1683">Figure 14.</emphasis>
|
||
<taxonomicName id="4C424D1CFFF3446CA103702CFD30FB3C" authority="(von Hagenow)" baseAuthorityName="von Hagenow" baseAuthorityYear="1839" box="[271,674,1131,1154]" class="Gymnolaemata" family="Onychocellidae" genus="Hoplitaechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="27" pageNumber="1683" phylum="Bryozoa" rank="species" species="vespertilio">
|
||
<emphasis id="B936EA8DFFF3446CA103702CFD9BFB3C" box="[271,521,1131,1154]" italics="true" pageId="27" pageNumber="1683">Hoplitaechmella vespertilio</emphasis>
|
||
(von Hagenow)
|
||
</taxonomicName>
|
||
. (a
|
||
<emphasis id="B936EA8DFFF3446CA2B0702CFD5AFB3C" box="[700,712,1131,1154]" italics="true" pageId="27" pageNumber="1683">–</emphasis>
|
||
f) NHMUK D15590; Cretaceous,probably lower Maastrichtian, Trimingham, Norfolk, UK; (a) autozooids and small avicularia; (b) autozooid opesia with oral spine bases and a pair of small avicularia; (c) autozooid with a sheet-like lamina of additional cryptocystal calcification; (d) large vicarious avicularium; (e) damaged growing edge showing pore chambers; (f) early astogeny with apparent ancestrula arrowed. (g
|
||
<emphasis id="B936EA8DFFF3446CA33A70A7FCD0FB49" box="[822,834,1248,1271]" italics="true" pageId="27" pageNumber="1683">–</emphasis>
|
||
i) SMF 29623; Cretaceous, lower Maastrichtian, Rügen, Mecklenburg-Vorpommern, Germany; (g) kenozooid (left) and vicarious avicularium with small opesiae; (h) ovicellate zooid containing apparent intermural bud; (i) detail of ovicell flanked by small interzooidal avicularia. Scale bars: a, c
|
||
<emphasis id="B936EA8DFFF3446CA2BC717FFD2EFAF1" box="[688,700,1336,1359]" italics="true" pageId="27" pageNumber="1683">–</emphasis>
|
||
h = 200 µm; b, i = 100 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFD369FFFF3446CA0CC71CCFEF9F990" blockId="27.[160,1155,1419,1582]" pageId="27" pageNumber="1683">
|
||
A poorly preserved colony from the Late Paleocene to Early Eocene Red Bluff Tuff of Chatham Island,
|
||
<collectingCountry id="F355760FFFF3446CA15171E9FE66FA78" box="[349,500,1454,1478]" name="New Zealand" pageId="27" pageNumber="1683">New Zealand</collectingCountry>
|
||
, was questionably assigned to
|
||
<taxonomicName id="4C424D1CFFF3446CA35F71E9FB91FA78" authorityName="Voigt" authorityYear="1949" box="[851,1027,1454,1478]" class="Gymnolaemata" family="Onychocellidae" genus="Hoplitaechmella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="27" pageNumber="1683" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B936EA8DFFF3446CA35F71E9FB91FA78" box="[851,1027,1454,1478]" italics="true" pageId="27" pageNumber="1683">Hoplitaechmella</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFD34B6EFFF3446CA42271E9FEFBFA57" author="Gordon DP & Taylor PD" pageId="27" pageNumber="1683" pagination="1 - 45" refId="ref26034" refString="Gordon DP, Taylor PD. 1999. Latest Paleocene to earliest Eocene bryozoans from Chatham Island, New Zealand. Bull Brit Mus (Nat Hist), Geol. 55: 1 - 45." type="journal article" year="1999">Gordon and Taylor (1999)</bibRefCitation>
|
||
. Otherwise, the genus is known only from the European and Central Asian Late Cretaceous (
|
||
<bibRefCitation id="EFD34B6EFFF3446CA1A771B4FDBAF9B5" author="Voigt E" box="[427,552,1523,1547]" pageId="27" pageNumber="1683" pagination="1 - 49" refId="ref28686" refString="Voigt E. 1949. Cheilostome Bryozoen aus der Quadratenkreide Nordwestdeutschlands. Mitt Geol Staatsinst Hamburg. 19: 1 - 49." type="journal article" year="1949">Voigt 1949</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD34B6EFFF3446CA23B71B4FDFCF9B5" author="Voigt E" box="[567,622,1523,1547]" pageId="27" pageNumber="1683" pagination="15 - 109" refId="ref28709" refString="Voigt E. 1951. Das Maastricht-Vorkommen von Ilten bei Hannover und seine Fauna. Mitt Geol Staatsinst Hamburg. 20: 15 - 109." type="journal article" year="1951">1951</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD34B6EFFF3446CA27171B4FD26F9B5" author="Voigt E" box="[637,692,1523,1547]" pageId="27" pageNumber="1683" refId="ref28832" refString="Voigt E. 1962. Viрkнiмiлovыi мsaнki ivрopiйсkoй caсti СССР i нikotoрыk сopрidiльныk obлaсtiй [Upper Cretaceous bryozoans of the European part of the USSR and neighbouring territories]. Мoсkva [Moskva]: Izdatiльсtvo Мoсkovсkogo uнiviрсitita [Moscow University Press]." type="book" year="1962">1962</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD34B6EFFF3446CA2CE71B4FD6BF9B5" author="Voigt E" box="[706,761,1523,1547]" pageId="27" pageNumber="1683" pagination="5 - 95" refId="ref28876" refString="Voigt E. 1967. Oberkreide-Bryozoen aus den asiatischen Gebieten der UdSSR. Mitt Geol Staatsinst Hamburg. 36: 5 - 95." type="journal article" year="1967">1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD34B6EFFF3446CA30471B4FCADF9B5" author="Voigt E" box="[776,831,1523,1547]" pageId="27" pageNumber="1683" pagination="33 - 88" refId="ref29011" refString="Voigt E. 1979. Bryozoen der Kunrader Schichten in Sud-Limburg (Oberkreide, Ob. Maastrichtium). I. Cheilostomata. Grondb Hamer. 2: 33 - 88." type="journal article" year="1979">1979</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD34B6EFFF3446CA34171B4FC7DF9B5" author="Veenstra HJ" box="[845,1007,1523,1547]" pageId="27" pageNumber="1683" refId="ref28506" refString="Veenstra HJ. 1963. Microscopic studies of boulder clays. Groningen: Rijksuniversiteit Groningen." type="book" year="1963">Veenstra 1963</bibRefCitation>
|
||
) and Danian (
|
||
<bibRefCitation id="EFD34B6EFFF3446CA0AA7251FECCF990" author="Berthelsen O" box="[166,350,1558,1582]" pageId="27" pageNumber="1683" refId="ref23703" refString="Berthelsen O. 1962. Cheilostome Bryozoa in the Danian deposits of East Denmark. KObenhavn: C. A. Reitzel." type="book" year="1962">Berthelsen 1962</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFD369FFFF3446CA0AC7237FF79F937" blockId="27.[160,1065,1648,1707]" box="[160,235,1648,1673]" pageId="27" pageNumber="1683">
|
||
<emphasis id="B936EA8DFFF3446CA0AC7237FF79F937" bold="true" box="[160,235,1648,1673]" italics="true" pageId="27" pageNumber="1683">Range</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFD369FFFF3446CA0AC72D4FBBBF915" blockId="27.[160,1065,1648,1707]" box="[160,1065,1683,1707]" pageId="27" pageNumber="1683">Cretaceous (Campanian) to Paleocene (Danian,?Thanetian),?Eocene (Ypresian).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |