875 lines
132 KiB
XML
875 lines
132 KiB
XML
<document id="3C066748697C6C64DFBE9A03788B807C" ID-CLB-Dataset="294285" ID-DOI="10.4202/app.00115.2014" ID-GBIF-Dataset="fe4529e6-7d02-4afc-b3bf-4783c1df2e65" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626269" IM.illustrations_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1707248722526" checkinUser="felipe" docAuthor="Colpaert, Wouter, Bosselaers, Mark & Lambert, Olivier" docDate="2015" docId="A6740C2FFFFAFFB0FCF9FB72E842FAC9" docLanguage="en" docName="ActaPalaeontolPol.60.1.1-10.pdf" docOrigin="Acta Palaeontologica Polonica 60 (1)" docSource="http://dx.doi.org/10.4202/app.00115.2014" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Brabocetus gigaseorum Colpaert, Bosselaers & Lambert 2015, sp. nov." docType="treatment" docUuid="3A577BF5-B6A6-4F9C-8524-2F40912A8F11" docUuidSource="ZooBank" docVersion="7" lastPageNumber="7" masterDocId="5A4D7457FFFBFFB6FFD1FF99ED72FFDA" masterDocTitle="Out of the Pacific: A second fossil porpoise from the Pliocene of the North Sea Basin" masterLastPageNumber="10" masterPageNumber="1" pageNumber="2" updateTime="1715888006825" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DBCD3A31767512B346D36C7E89DDE879">Out of the Pacific: A second fossil porpoise from the Pliocene of the North Sea Basin</mods:title>
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<mods:namePart id="BF44290D410C2040F076BE984E8B5D23">Colpaert, Wouter</mods:namePart>
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<mods:namePart id="50E4D4BD1EDDCA7C25E63AC83D7B518B">Bosselaers, Mark</mods:namePart>
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<mods:namePart id="362A5891CAAC9C31850471344EA14EF6">Lambert, Olivier</mods:namePart>
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<mods:typeOfResource id="C7B706F953191EC9362170006709239C">text</mods:typeOfResource>
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<mods:title id="D4D0869DB5DE34525AA423742D69FDF1">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="CF8F1270D60733F44585649B64ACD353">
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<mods:date id="0E237F61CEB52C6287E0B9BAE08E316E">2015</mods:date>
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<mods:number id="30A5398B8AE609164405A18037C2B997">2014-12-14</mods:number>
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<treatment id="A6740C2FFFFAFFB0FCF9FB72E842FAC9" ID-DOI="http://doi.org/10.5281/zenodo.10989429" ID-GBIF-Taxon="225696677" ID-Zenodo-Dep="10989429" LSID="urn:lsid:zoobank.org:act:3A577BF5-B6A6-4F9C-8524-2F40912A8F11" httpUri="http://treatment.plazi.org/id/A6740C2FFFFAFFB0FCF9FB72E842FAC9" lastPageId="6" lastPageNumber="7" pageId="1" pageNumber="2">
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<subSubSection id="66C7EEB2FFFAFFB7FCF9FB72E9B1FAD2" box="[808,1219,1259,1288]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="2E62BD39FFFAFFB7FCF9FB72E9B1FAD2" blockId="1.[808,1219,1259,1288]" box="[808,1219,1259,1288]" pageId="1" pageNumber="2">
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<heading id="752A0A55FFFAFFB7FCF9FB72E9B1FAD2" box="[808,1219,1259,1288]" fontSize="12" level="2" pageId="1" pageNumber="2" reason="2">
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<taxonomicName id="E9DDC6BAFFFAFFB7FCF9FB72E92BFAD2" authority="Colpaert, Bosselaers & Lambert, 2015" authorityName="Colpaert, Bosselaers & Lambert" authorityYear="2015" box="[808,1113,1259,1288]" class="Mammalia" family="Phocoenidae" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="1" pageNumber="2" phylum="Chordata" rank="species" species="gigaseorum" status="sp. nov.">
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<emphasis id="1CA9612BFFFAFFB7FCF9FB72E92BFAD2" box="[808,1113,1259,1288]" italics="true" pageId="1" pageNumber="2">Brabocetus gigaseorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="079ADC50FFFAFFB7FBB0FB75E9B1FAD2" box="[1121,1219,1260,1288]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="66C7EEB2FFFAFFB7FCF9FA80EECAFAF4" box="[808,952,1305,1326]" pageId="1" pageNumber="2" type="description">
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<paragraph id="2E62BD39FFFAFFB7FCF9FA80EECAFAF4" blockId="1.[808,952,1305,1326]" box="[808,952,1305,1326]" pageId="1" pageNumber="2">
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<figureCitation id="B6E6A1BCFFFAFFB7FCF9FA80EE19FAF4" box="[808,875,1305,1326]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="1" pageNumber="2">Figs. 2</figureCitation>
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,
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<figureCitation id="B6E6A1BCFFFAFFB7FCA4FA80EEF0FAF4" box="[885,898,1305,1326]" captionStart="Fig" captionStartId="4.[862,894,973,993]" captionTargetBox="[868,1529,164,947]" captionTargetId="figure-663@4.[867,1528,166,911]" captionTargetPageId="4" captionText="Fig. 3. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Anterior view of the facial region. B. Posterior view. C. Ventral view of the right squamosal." figureDoi="http://doi.org/10.5281/zenodo.10626273" httpUri="https://zenodo.org/record/10626273/files/figure.png" pageId="1" pageNumber="2">3</figureCitation>
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,
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<figureCitation id="B6E6A1BCFFFAFFB7FC5CFA80EEC6FAF4" box="[909,948,1305,1326]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="5.[98,131,929,949]" captionStartId-1="6.[151,184,505,525]" captionStartId-2="6.[862,895,1824,1844]" captionTargetBox-0="[98,1477,158,906]" captionTargetBox-1="[162,1521,162,482]" captionTargetBox-2="[920,1476,1335,1799]" captionTargetId-0="figure-619@5.[97,1481,157,580]" captionTargetId-2="figure-810@6.[1016,1310,1335,1765]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="6" captionText-0="Fig. 5. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Right lateral view, photo (A1) and corresponding line drawing (A2). Dotted lines indicate the dorsal margin of the temporal fossa (A1), the reconstructed right postorbital process, and poorly visible sutures (A2). B. Left lateral view." captionText-1="Fig. 6. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Right ventrolateral view of the palate and orbital regions. Photo (A) and corresponding line drawing (B). Dotted lines indicate poorly visible sutures. Dark grey surface indicates fracture surfaces." captionText-2="Fig. 7. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Fragmentary basicranium in ventral view. Dotted lines indicate poorly visible sutures, the right tympanosquamosal recess, the groove for the mandibular nerve V3, and the presumed area of origin for scalenus medius." figureDoi-0="http://doi.org/10.5281/zenodo.10626277" figureDoi-1="http://doi.org/10.5281/zenodo.10626279" figureDoi-2="http://doi.org/10.5281/zenodo.10626283" httpUri-0="https://zenodo.org/record/10626277/files/figure.png" httpUri-1="https://zenodo.org/record/10626279/files/figure.png" httpUri-2="https://zenodo.org/record/10626283/files/figure.png" pageId="1" pageNumber="2">5–7</figureCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="66C7EEB2FFFAFFB7FCF9FADCE97BFA6A" pageId="1" pageNumber="2" type="etymology">
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<paragraph id="2E62BD39FFFAFFB7FCF9FADCE97BFA6A" blockId="1.[808,1481,1349,1456]" pageId="1" pageNumber="2">
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<emphasis id="1CA9612BFFFAFFB7FCF9FADCEEFDFA80" box="[808,911,1349,1370]" italics="true" pageId="1" pageNumber="2">Etymology</emphasis>
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: Honouring Paul and Pierre Gigase (father and son) for their long-term effort in collecting fossil marine mammal remains in the area of Antwerp and for generously donating the specimen IRSNB M. 2171 to the IRSNB.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="66C7EEB2FFFAFFB7FCF9FA25EEB2F87C" pageId="1" pageNumber="2" type="materials_examined">
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||
<paragraph id="2E62BD39FFFAFFB7FCF9FA25EEBFF9F2" blockId="1.[808,1481,1468,1576]" pageId="1" pageNumber="2">
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||
<materialsCitation id="9EB5B764FFFAFFB7FCF9FA25EEBFF9F2" ID-GBIF-Occurrence="4845992301" collectionCode="IRSNB" pageId="1" pageNumber="2" specimenCount="1" typeStatus="holotype">
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<emphasis id="1CA9612BFFFAFFB7FCF9FA25EEF2FA0B" box="[808,896,1468,1489]" italics="true" pageId="1" pageNumber="2">
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<typeStatus id="F166039BFFFAFFB7FCF9FA25EEF2FA0B" box="[808,896,1468,1489]" pageId="1" pageNumber="2">Holotype</typeStatus>
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||
</emphasis>
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||
:
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<collectionCode id="48CC25FCFFFAFFB7FC5FFA24EEA7FA08" box="[910,981,1469,1490]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="1" pageNumber="2">IRSNB</collectionCode>
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M. 2171, a partial skull in three fragments, lacking the anterior part of the rostrum, part of the left supraorbital process, the nasals, the pterygoids, and some elements of the cerebral cavity and basicranium.
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</materialsCitation>
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</paragraph>
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<paragraph id="2E62BD39FFFAFFB7FCF9F9ACE976F97A" blockId="1.[808,1481,1588,1696]" pageId="1" pageNumber="2">
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<emphasis id="1CA9612BFFFAFFB7FCF9F9ACEED6F993" box="[808,932,1588,1610]" italics="true" pageId="1" pageNumber="2">Type locality</emphasis>
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:
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<materialsCitation id="9EB5B764FFFAFFB7FC60F9ACEE8CF97A" ID-GBIF-Occurrence="4845992302" country="Belgium" county="The" latitude="51.26389" location="Beverentunnel" longLatPrecision="17" longitude="4.2719445" municipality="Antwerp Harbour" pageId="1" pageNumber="2" specimenCount="1">
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<location id="2B02EBE2FFFAFFB7FC60F9ACE94DF990" LSID="urn:lsid:plazi:treatment:A6740C2FFFFAFFB0FCF9FB72E842FAC9:2B02EBE2FFFAFFB7FC60F9ACE94DF990" box="[945,1087,1589,1610]" country="Belgium" county="The" latitude="51.26389" longLatPrecision="17" longitude="4.2719445" municipality="Antwerp Harbour" name="Beverentunnel" pageId="1" pageNumber="2">Beverentunnel</location>
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, left bank of the
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<location id="2B02EBE2FFFAFFB7FB34F9ACE819F990" LSID="urn:lsid:plazi:treatment:A6740C2FFFFAFFB0FCF9FB72E842FAC9:2B02EBE2FFFAFFB7FB34F9ACE819F990" box="[1253,1387,1589,1610]" country="Belgium" county="The" latitude="51.26389" longLatPrecision="17" longitude="4.2719445" municipality="Antwerp Harbour" name="River Scheldt" pageId="1" pageNumber="2">River Scheldt</location>
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,
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<collectingMunicipality id="CE062743FFFAFFB7FAA4F9ACEE08F9BC" pageId="1" pageNumber="2">Antwerp Harbour</collectingMunicipality>
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, northern
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<collectingCountry id="56CAFDA9FFFAFFB7FC31F9C8E941F9BC" box="[992,1075,1617,1638]" name="Belgium" pageId="1" pageNumber="2">Belgium</collectingCountry>
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(
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<figureCitation id="B6E6A1BCFFFAFFB7FB92F9C8E90CF9BC" box="[1091,1150,1617,1638]" captionStart="Fig" captionStartId="1.[98,131,582,602]" captionTargetBox="[100,767,161,557]" captionTargetId="figure-792@1.[416,758,161,557]" captionTargetPageId="1" captionText="Fig. 1. Maps indicating the type localities of fossil phocoenids in the port of Antwerp, north-west of Antwerp, Belgium, southern margin of the North Sea Basin. A. Map of northern Belgium. B. Southern margin of the North Sea Basin.C. Port of Antwerp. The small asterisk indicates the type locality of Septemtriocetus bosselaersi, in the Verrebroek Dock, and the large asterisk indicates the type locality of Brabocetus gigaseorum gen. et sp. nov., at the entrance of the Beveren Tunnel. Modified from Lambert (2008)." figureDoi="http://doi.org/10.5281/zenodo.10989425" httpUri="https://zenodo.org/record/10989425/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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).
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<collectingCounty id="C703C5B5FFFAFFB7FB44F9C8E9C8F9BC" box="[1173,1210,1617,1638]" pageId="1" pageNumber="2">The</collectingCounty>
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locality is situated two kilometres north of the village of Kallo. Geographic coordinates:
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<geoCoordinate id="4BE9DBFEFFFAFFB7FA69F9F7EEFBF97A" degrees="51" direction="north" minutes="15" orientation="latitude" pageId="1" pageNumber="2" precision="15" seconds="50" value="51.26389">
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N 51°15
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<emphasis id="1CA9612BFFFAFFB7FCB3F912EE15F97A" bold="true" box="[866,871,1675,1696]" italics="true" pageId="1" pageNumber="2">′</emphasis>
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50
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<emphasis id="1CA9612BFFFAFFB7FCAEF912EEFBF97A" bold="true" box="[895,905,1675,1696]" italics="true" pageId="1" pageNumber="2">″</emphasis>
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</geoCoordinate>
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,
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<geoCoordinate id="4BE9DBFEFFFAFFB7FC44F912EE8CF97A" box="[917,1022,1675,1696]" degrees="4" direction="east" minutes="16" orientation="longitude" pageId="1" pageNumber="2" precision="15" seconds="19" value="4.2719445">
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E 4°16
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<emphasis id="1CA9612BFFFAFFB7FC06F912EEAEF97A" bold="true" box="[983,988,1675,1696]" italics="true" pageId="1" pageNumber="2">′</emphasis>
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19
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<emphasis id="1CA9612BFFFAFFB7FC25F912EE8CF97A" bold="true" box="[1012,1022,1675,1696]" italics="true" pageId="1" pageNumber="2">″</emphasis>
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</geoCoordinate>
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||
</materialsCitation>
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||
.
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||
</paragraph>
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<paragraph id="2E62BD39FFFAFFB7FCF9F934EEB2F87C" blockId="1.[808,1482,1708,1958]" pageId="1" pageNumber="2">
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<emphasis id="1CA9612BFFFAFFB7FCF9F934EED4F91B" box="[808,934,1708,1730]" italics="true" pageId="1" pageNumber="2">Type horizon</emphasis>
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: The skull was found in the Kattendijk Formation,
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<quantity id="E92510DCFFFAFFB7FA73F934E8BBF918" box="[1442,1481,1709,1730]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="2" unit="m" value="2.0">2 m</quantity>
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below the Oorderen Sands Member, Lillo Formation, in a layer of grey glauconitic sand without mollusc shells. The Kattendijk Formation is dated from the early Pliocene (Zanclean) based on dinoflagellates, foraminifers, and molluscs (
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<bibRefCitation id="4A4CC0C8FFFAFFB7FBEBF886E845F8EE" author="Vandenberghe, N. & Laga, P. & Steurbaut, E. & Hardenbol, J. & Vail, P. R." box="[1082,1335,1823,1844]" pageId="1" pageNumber="2" pagination="119 - 154" refId="ref9754" refString="Vandenberghe, N., Laga, P., Steurbaut, E., Hardenbol, J., and Vail, P. R. 1998. Tertiary sequence stratigraphy at the southern border of the North Sea Basin in Belgium. Special Publication SEPM 60: 119 - 154." type="journal article" year="1998">Vandenberghe et al. 1998</bibRefCitation>
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||
;
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<bibRefCitation id="4A4CC0C8FFFAFFB7FA93F886EE2EF88B" author="Louwye, S. & Head, M. J. & De Schepper, S." pageId="1" pageNumber="2" pagination="353 - 378" refId="ref9063" refString="Louwye, S., Head, M. J., and De Schepper, S. 2004. Dinoflagellate cyst stratigraphy and palaeoecology of the Pliocene in northern Belgium, southern North Sea Basin. Geological Magazine 141: 353 - 378." type="journal article" year="2004">Louwye et al. 2004</bibRefCitation>
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), more precisely between 5 and 4.4 Ma (
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<bibRefCitation id="4A4CC0C8FFFAFFB7FB09F8A5E8CDF88B" author="De Schepper, S. & Head, M. J. & Louwye, S." box="[1240,1471,1852,1873]" pageId="1" pageNumber="2" pagination="92 - 112" refId="ref8501" refString="De Schepper, S., Head, M. J., and Louwye, S. 2009. Pliocene dinoflagellate cyst stratigraphy, palaeoecology and sequence stratigraphy of the Tunnel-Canal Dock, Belgium. Geological Magazine 146: 92 - 112." type="journal article" year="2009">De Schepper et al. 2009</bibRefCitation>
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||
). This unit represents a neritic deposition (
|
||
<quantity id="E92510DCFFFAFFB7FB6DF8C1E866F8B7" box="[1212,1300,1880,1901]" metricMagnitude="1" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="2" unit="m" value="40.0" valueMax="50.0" valueMin="30.0">30–50 m</quantity>
|
||
water depth) with open-marine influence, under a warm to temperate climate (
|
||
<bibRefCitation id="4A4CC0C8FFFAFFB7FAB2F8ECEEC7F87C" author="De Schepper, S. & Head, M. J. & Louwye, S." pageId="1" pageNumber="2" pagination="92 - 112" refId="ref8501" refString="De Schepper, S., Head, M. J., and Louwye, S. 2009. Pliocene dinoflagellate cyst stratigraphy, palaeoecology and sequence stratigraphy of the Tunnel-Canal Dock, Belgium. Geological Magazine 146: 92 - 112." type="journal article" year="2009">De Schepper et al. 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7AA2EDB1FFF9FFB4FF48FD31E979FD2E" ID-DOI="http://doi.org/10.5281/zenodo.10626271" ID-Zenodo-Dep="10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="2" pageNumber="3" startId="2.[153,186,680,700]" targetBox="[154,1530,165,658]" targetPageId="2" targetType="figure">
|
||
<paragraph id="2E62BD39FFF9FFB4FF48FD31E979FD2E" blockId="2.[151,1535,678,756]" pageId="2" pageNumber="3">
|
||
Fig. 2. Skull of the phocoenid cetacean
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FDD9FD3FEF93FD66" authority="Colpaert & Bosselaers & Lambert, 2015" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[520,737,678,700]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="gigaseorum" status="gen. et sp. nov.">
|
||
<emphasis id="1CA9612BFFF9FFB4FDD9FD3FEF93FD66" box="[520,737,678,700]" italics="true" pageId="2" pageNumber="3">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="079ADC50FFF9FFB4FD39FD31EE07FD66" box="[744,885,680,700]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (
|
||
<emphasis id="1CA9612BFFF9FFB4FE81FD5BEC12FD0D" bold="true" box="[336,352,706,727]" pageId="2" pageNumber="3">A</emphasis>
|
||
) and corresponding line drawing (
|
||
<emphasis id="1CA9612BFFF9FFB4FD73FD5BEFC3FD0D" bold="true" box="[674,689,706,727]" pageId="2" pageNumber="3">B</emphasis>
|
||
). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="66C7EEB2FFF9FFB4FF46FC8BEF83F937" pageId="2" pageNumber="3" type="diagnosis">
|
||
<paragraph id="2E62BD39FFF9FFB4FF46FC8BEF83F937" blockId="2.[151,824,785,1773]" pageId="2" pageNumber="3">
|
||
<emphasis id="1CA9612BFFF9FFB4FF46FC8BEC7BFCF1" box="[151,265,786,811]" italics="true" pageId="2" pageNumber="3">Diagnosis</emphasis>
|
||
.—
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FEFDFC88EF45FCF1" authority="Colpaert & Bosselaers & Lambert, 2015" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[300,567,785,811]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="gigaseorum" status="sp. nov.">
|
||
<emphasis id="1CA9612BFFF9FFB4FEFDFC88EF45FCF1" box="[300,567,785,811]" italics="true" pageId="2" pageNumber="3">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="079ADC50FFF9FFB4FDEFFC8BEFE7FCF1" box="[574,661,786,811]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has a cranium intermediate in size between that of the extant species
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FCD3FCABECDBFCB1" baseAuthorityName="G. Cuvier" baseAuthorityYear="1829" class="Mammalia" family="Phocoenidae" genus="Neophocaena" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="phocaenoides">
|
||
<emphasis id="1CA9612BFFF9FFB4FCD3FCABECDBFCB1" italics="true" pageId="2" pageNumber="3">Neophocaena phocaenoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FE36FCC8EFB3FCB1" baseAuthorityName="True" baseAuthorityYear="1885" box="[487,705,849,875]" class="Mammalia" family="Phocoenidae" genus="Phocoenoides" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="dalli">
|
||
<emphasis id="1CA9612BFFF9FFB4FE36FCC8EFB3FCB1" box="[487,705,849,875]" italics="true" pageId="2" pageNumber="3">Phocoenoides dalli</emphasis>
|
||
</taxonomicName>
|
||
. It differs from all other phocoenids except
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FDC2FCE8EFB4FC51" authorityName="Lambert" authorityYear="2008" box="[531,710,881,907]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FDC2FCE8EFB4FC51" box="[531,710,881,907]" italics="true" pageId="2" pageNumber="3">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
in having a frontal boss that is longer than wide. It differs from all other phocoenids except
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FEBAFC28EF5CFC11" authorityName="Ichishima & Kimura" authorityYear="2005" box="[363,558,945,971]" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FEBAFC28EF5CFC11" box="[363,558,945,971]" italics="true" pageId="2" pageNumber="3">Haborophocoena</emphasis>
|
||
</taxonomicName>
|
||
in the anteromedial sulcus being shorter than the posteromedial sulcus. It possibly differs from all extinct phocoenids except
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FDAEFC68EE6BFBD1" box="[639,793,1009,1035]" class="Mammalia" family="Phocoenidae" genus="Miophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FDAEFC68EE6BFBD1" box="[639,793,1009,1035]" italics="true" pageId="2" pageNumber="3">Miophocaena</emphasis>
|
||
</taxonomicName>
|
||
in the presence of an additional longitudinal sulcus across the premaxillary eminences. It differs from all phocoenids except
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FF01FBC8ECE1FBB1" authorityName="Ichishima & Kimura" authorityYear="2005" box="[208,403,1105,1131]" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FF01FBC8ECE1FBB1" box="[208,403,1105,1131]" italics="true" pageId="2" pageNumber="3">Haborophocoena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FE74FBC8EF4DFBB1" box="[421,575,1105,1131]" class="Mammalia" family="Phocoenidae" genus="Miophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FE74FBC8EF4DFBB1" box="[421,575,1105,1131]" italics="true" pageId="2" pageNumber="3">Miophocaena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FD80FBC8EE43FBB1" box="[593,817,1105,1131]" class="Mammalia" family="Phocoenidae" genus="Piscolithax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="tedfordi">
|
||
<emphasis id="1CA9612BFFF9FFB4FD80FBC8EE43FBB1" box="[593,817,1105,1131]" italics="true" pageId="2" pageNumber="3">Piscolithax tedfordi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FF16FBEBEC08FB56" authorityName="Lambert" authorityYear="2008" box="[199,378,1138,1164]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FF16FBEBEC08FB56" box="[199,378,1138,1164]" italics="true" pageId="2" pageNumber="3">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
, in that the level of the anterior margin of the bony nares is in line with the postorbital process of the frontal; from all other phocoenids except
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FD45FB2BEDCDFB31" authorityName="Ichishima & Kimura" authorityYear="2005" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FD45FB2BEDCDFB31" italics="true" pageId="2" pageNumber="3">Haborophocoena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FF1EFB4BEC1BFB36" box="[207,361,1234,1260]" class="Mammalia" family="Phocoenidae" genus="Miophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FF1EFB4BEC1BFB36" box="[207,361,1234,1260]" italics="true" pageId="2" pageNumber="3">Miophocaena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FEA8FB4BEF61FB36" authorityName="Palmer" authorityYear="1899" box="[377,531,1234,1260]" class="Mammalia" family="Phocoenidae" genus="Neophocaena" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FEA8FB4BEF61FB36" box="[377,531,1234,1260]" italics="true" pageId="2" pageNumber="3">Neophocaena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FDF3FB4BEE72FB36" box="[546,768,1234,1260]" class="Mammalia" family="Phocoenidae" genus="Piscolithax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="tedfordi">
|
||
<emphasis id="1CA9612BFFF9FFB4FDF3FB4BEE72FB36" box="[546,768,1234,1260]" italics="true" pageId="2" pageNumber="3">Piscolithax tedfordi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FF46FB6BEC38FAD6" authorityName="Lambert" authorityYear="2008" box="[151,330,1266,1292]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FF46FB6BEC38FAD6" box="[151,330,1266,1292]" italics="true" pageId="2" pageNumber="3">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
in that the first anterior dorsal infraorbital foramen is distinctly posterior to the antorbital notch; from
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FF46FAABEC28FA96" authorityName="Ichishima & Kimura" authorityYear="2005" box="[151,346,1330,1356]" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FF46FAABEC28FA96" box="[151,346,1330,1356]" italics="true" pageId="2" pageNumber="3">Haborophocoena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FEB6FAABEF73FA96" box="[359,513,1330,1356]" class="Mammalia" family="Phocoenidae" genus="Miophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FEB6FAABEF73FA96" box="[359,513,1330,1356]" italics="true" pageId="2" pageNumber="3">Miophocaena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FDDFFAABEFA6FA96" authorityName="Ichishima & Kimura" authorityYear="2000" box="[526,724,1330,1356]" class="Mammalia" family="Phocoenidae" genus="Numataphocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FDDFFAABEFA6FA96" box="[526,724,1330,1356]" italics="true" pageId="2" pageNumber="3">Numataphocoena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FD30FAABED8FFAB6" class="Mammalia" family="Phocoenidae" genus="Piscolithax" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="1CA9612BFFF9FFB4FD30FAABEDB5FAB6" italics="true" pageId="2" pageNumber="3">Piscolithax</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FED6FACBECC3FAB6" authorityName="Murakami, Shimada, Hikida & Hirano" authorityYear="2012" box="[263,433,1362,1388]" class="Mammalia" family="Phocoenidae" genus="Pterophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FED6FACBECC3FAB6" box="[263,433,1362,1388]" italics="true" pageId="2" pageNumber="3">Pterophocaena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FE6CFACBEF39FAB6" authorityName="Racicot, Deméré, Beatty & Boessenecker" authorityYear="2014" box="[445,587,1362,1388]" class="Mammalia" family="Phocoenidae" genus="Semirostrum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FE6CFACBEF39FAB6" box="[445,587,1362,1388]" italics="true" pageId="2" pageNumber="3">Semirostrum</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FD54FACBEE4AFAB6" authorityName="Lambert" authorityYear="2008" box="[645,824,1362,1388]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FD54FACBEE4AFAB6" box="[645,824,1362,1388]" italics="true" pageId="2" pageNumber="3">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
in the anterolaterally widely-open antorbital notch, not laterally limited by an extension of the antorbital process; and from extant phocoenids in a more pronounced asymmetry of the vertex, with medial sutures more distinctly shifted to the left side. It further differs from the other North Sea fossil phocoenid
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FEC2F98AECB4F9F7" authorityName="Lambert" authorityYear="2008" box="[275,454,1555,1581]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FEC2F98AECB4F9F7" box="[275,454,1555,1581]" italics="true" pageId="2" pageNumber="3">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
in that it possesses: an anteriorly longer pterygoid sinus fossa, reaching the level of the antorbital notch; a higher premaxillary eminence, overhanging the posterolateral sulcus; a significantly transversely narrower frontal boss; a lower, not dorsally pointed temporal fossa; a deeper and longer squamosal fossa; and a postglenoid process of the squamosal being thickened in lateral view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="66C7EEB2FFF9FFB0FF46F89FE842FAC9" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="2E62BD39FFF9FFB4FF46F89FEC50F8C5" blockId="2.[151,290,1798,1823]" box="[151,290,1798,1823]" pageId="2" pageNumber="3">
|
||
<emphasis id="1CA9612BFFF9FFB4FF46F89FEC50F8C5" box="[151,290,1798,1823]" italics="true" pageId="2" pageNumber="3">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFF9FFB4FF46F8B6E9E9FA4F" blockId="2.[151,824,1839,1960]" lastBlockId="2.[862,1535,786,1429]" pageId="2" pageNumber="3">
|
||
Based on the anteriorly converging and rectilinear lateral margins of the maxillae at the rostrum base, a tentative reconstruction of the outline of the pointed rostrum of
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FD26F8F4EC19F87D" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="gigaseorum">
|
||
<emphasis id="1CA9612BFFF9FFB4FD26F8F4EC19F87D" italics="true" pageId="2" pageNumber="3">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
is provided (
|
||
<figureCitation id="B6E6A1BCFFF9FFB4FDD3F816EF3BF872" box="[514,585,1935,1960]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
). This reconstruction suggests a rostrum length shorter than the cranium length, a feature shared with extant phocoenids and possibly
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FA1CFCABEE9AFCB6" authorityName="Lambert" authorityYear="2008" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FA1CFCABEE9AFCB6" italics="true" pageId="2" pageNumber="3">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
, but unlike the longer-snouted extinct phocoenids
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FC45FCEAE97FFC56" authorityName="de Muizon" authorityYear="1986" box="[916,1037,883,908]" class="Mammalia" family="Phocoenidae" genus="Lomacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FC45FCEAE97FFC56" box="[916,1037,883,908]" italics="true" pageId="2" pageNumber="3">Lomacetus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FBCDFCEBE9E8FC56" authorityName="de Muizon" authorityYear="1983" box="[1052,1178,882,908]" class="Mammalia" family="Phocoenidae" genus="Piscolithax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FBCDFCEBE9E8FC56" box="[1052,1178,882,908]" italics="true" pageId="2" pageNumber="3">Piscolithax</emphasis>
|
||
</taxonomicName>
|
||
, and probably
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FA95FCEBE88DFC56" authorityName="Barnes" authorityYear="1985" box="[1348,1535,882,908]" class="Mammalia" family="Phocoenidae" genus="Salumiphocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF9FFB4FA95FCEBE88DFC56" box="[1348,1535,882,908]" italics="true" pageId="2" pageNumber="3">Salumiphocaena</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="4A4CC0C8FFF9FFB4FCB4FC0DEE88FC77" author="Wilson, L. E." box="[869,1018,916,941]" pageId="2" pageNumber="3" pagination="1 - 34" refId="ref9934" refString="Wilson, L. E. 1973. A delphinid (Mammalia, Cetacea) from the Miocene of Palos Verdes Hills, California. University of California Publications in Geological Sciences 103: 1 - 34." type="journal article" year="1973">Wilson 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFF9FFB4FBD6FC0DE9E5FC77" author="Barnes, L. G." box="[1031,1175,916,941]" pageId="2" pageNumber="3" pagination="1 - 46" refId="ref8032" refString="Barnes, L. G. 1984. Fossil odontocetes (Mammalia: Cetacea) from the Almejas Formation, Isla Cedros, Mexico. PaleoBios 42: 1 - 46." type="journal article" year="1984">Barnes 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFF9FFB4FB75FC0DE84CFC77" author="Muizon, C. de" box="[1188,1342,916,941]" pageId="2" pageNumber="3" pagination="1 - 188" refId="ref9203" refString="Muizon, C. de 1984. Les Vertebres de la Formation Pisco (Perou). Deuxieme partie: Les Odontocetes (Cetacea, Mammalia) du Pliocene inferieur du Sud-Sacaco. Travaux de l'Institut Francais d'Etudes Andines 27: 1 - 188." type="journal article" year="1984">Muizon 1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4A4CC0C8FFF9FFB4FA9BFC0DE8E1FC77" author="Muizon, C. de" box="[1354,1427,916,941]" pageId="2" pageNumber="3" pagination="1 - 244" refId="ref9271" refString="Muizon, C. de 1988 b. Les Vertebres fossiles de la Formation Pisco (Perou). Troisieme partie: Les Odontocetes (Cetacea, Mammalia) du Miocene. Travaux de l'Institut Francais d'Etudes Andines 42: 1 - 244." type="journal article" year="1988">1988b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFF9FFB4FA71FC0DEEE9FC14" author="Lambert, O." pageId="2" pageNumber="3" pagination="863 - 872" refId="ref9029" refString="Lambert, O. 2008. A new porpoise (Cetacea, Odontoceti, Phocoenidae) from the Pliocene of the North Sea. Journal of Vertebrate Paleontology 28: 863 - 872." type="journal article" year="2008">Lambert 2008</bibRefCitation>
|
||
). However, a more complete specimen is needed to confirm this statement. The dimensions of the cranium (
|
||
<tableCitation id="635F8882FFF9FFB4FA09FC4CEEE4FBD5" captionStart="Table 1" captionStartId="3.[98,151,160,181]" captionTargetBox="[105,688,236,1060]" captionTargetId="graphics-815@3.[98,769,227,1063]" captionTargetPageId="3" captionText="Table 1. Measurements (in mm) on the skull of Brabocetus gigaseorum gen. et sp. nov. IRSNB M. 2171 (holotype). e, estimate; + incomplete" httpUri="http://table.plazi.org/id/7AA2EDB1FFF8FFB5FFB3FF39EE72FF08" pageId="2" pageNumber="3" tableUuid="7AA2EDB1FFF8FFB5FFB3FF39EE72FF08">Table 1</tableCitation>
|
||
; pre- and postorbital widths, length from the antorbital notch to the nuchal crest) are roughly intermediate between those of the extant species
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FB40FBACE8BCFB95" baseAuthorityName="G. Cuvier" baseAuthorityYear="1829" box="[1169,1486,1077,1103]" class="Mammalia" family="Phocoenidae" genus="Neophocaena" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="phocaenoides">
|
||
<emphasis id="1CA9612BFFF9FFB4FB40FBACE8BCFB95" box="[1169,1486,1077,1103]" italics="true" pageId="2" pageNumber="3">Neophocaena phocaenoides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FC8FFBCFE946FBAA" baseAuthorityName="True" baseAuthorityYear="1885" box="[862,1076,1110,1136]" class="Mammalia" family="Phocoenidae" genus="Phocoenoides" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="dalli">
|
||
<emphasis id="1CA9612BFFF9FFB4FC8FFBCFE946FBAA" box="[862,1076,1110,1136]" italics="true" pageId="2" pageNumber="3">Phocoenoides dalli</emphasis>
|
||
</taxonomicName>
|
||
, and slightly smaller than in the
|
||
<typeStatus id="F166039BFFF9FFB4FA4CFBCEE88DFBAA" box="[1437,1535,1111,1136]" pageId="2" pageNumber="3">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="E9DDC6BAFFF9FFB4FCACFBEEE9C5FB4A" authorityName="Lambert" authorityYear="2008" box="[893,1207,1142,1169]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="bosselaersi">
|
||
<emphasis id="1CA9612BFFF9FFB4FCACFBEEE9C5FB4A" box="[893,1207,1142,1169]" italics="true" pageId="2" pageNumber="3">Septemtriocetus bosselaersi</emphasis>
|
||
</taxonomicName>
|
||
. The small temporal fossa is still anteroposteriorly longer than the orbit and the roof of the former is slightly higher. The cranium displays a high degree of asymmetry that is especially developed in the region of the bony nares and vertex. This asymmetry is expressed as a shift to the left (ca. 15° in comparison to the sagittal plane of the skull), which is particularly conspicuous in anterior view (
|
||
<figureCitation id="B6E6A1BCFFF9FFB4FCB7FAC5EEB0FAAF" box="[870,962,1372,1397]" captionStart="Fig" captionStartId="4.[862,894,973,993]" captionTargetBox="[868,1529,164,947]" captionTargetId="figure-663@4.[867,1528,166,911]" captionTargetPageId="4" captionText="Fig. 3. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Anterior view of the facial region. B. Posterior view. C. Ventral view of the right squamosal." figureDoi="http://doi.org/10.5281/zenodo.10626273" httpUri="https://zenodo.org/record/10626273/files/figure.png" pageId="2" pageNumber="3">Fig. 3A</figureCitation>
|
||
). Differences in width and height of left and right elements are detailed below.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFF9FFB4FC8FFA3FE883F93F" blockId="2.[862,1536,1446,1960]" pageId="2" pageNumber="3">
|
||
<emphasis id="1CA9612BFFF9FFB4FC8FFA3FEEABFA1A" box="[862,985,1446,1472]" italics="true" pageId="2" pageNumber="3">Premaxilla</emphasis>
|
||
.—In dorsal view, from the preserved anterior end of the rostrum, the premaxilla widens until the level of the antorbital notch, where its width equals the width of the maxilla (
|
||
<figureCitation id="B6E6A1BCFFF9FFB4FC10F990E974F9F8" box="[961,1030,1545,1570]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
). From that level onwards, the lateral margin of the bone becomes slightly concave over
|
||
<quantity id="E92510DCFFF9FFB4FA9BF9B0E8BAF998" box="[1354,1480,1577,1602]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">30–35 mm</quantity>
|
||
; it is followed backwards by the convex lateral margin of the premaxillary eminence. The maximum width of the mesorostral groove occurs at the level of the antorbital notches, from where the opening narrows backwards to terminate where the medial margins of the premaxillary eminences nearly abut.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFF9FFB5FC55F975E947FE43" blockId="2.[862,1536,1446,1960]" lastBlockId="3.[808,1482,160,409]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
Best preserved on the right side, the tiny elliptical premaxillary foramen (transverse diameter =
|
||
<quantity id="E92510DCFFF9FFB4FAECF894E8F0F8FC" box="[1341,1410,1805,1830]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="3" unit="mm" value="2.0">2 mm</quantity>
|
||
) is located posterior to the level of the antorbital notch. A short and shallow anteromedial sulcus originates from each premaxillary foramen; by contrast the posteromedial sulcus is longer. Consequently, the portion of the prenarial triangle posterior to the premaxillary foramen is longer than the portion anterior to the foramen. A roughly similar condition is seen only in
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FFB3FB08EC51FB71" authorityName="Ichishima & Kimura" authorityYear="2005" box="[98,291,1169,1195]" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FFB3FB08EC51FB71" box="[98,291,1169,1195]" italics="true" pageId="3" pageNumber="4">Haborophocoena</emphasis>
|
||
</taxonomicName>
|
||
, whereas other phocoenids, including the extant species, display a longer anteromedial sulcus. Together with the long posterolateral sulcus, the posteromedial sulcus demarcates the anteriorly pointed and moderately elevated premaxillary eminence (
|
||
<figureCitation id="B6E6A1BCFFF8FFB5FEA6FA8EECBBFAEA" box="[375,457,1303,1328]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="3" pageNumber="4">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="B6E6A1BCFFF8FFB5FE08FA8EEC8CFAEA" box="[473,510,1303,1328]" captionStart="Fig" captionStartId="4.[862,894,973,993]" captionTargetBox="[868,1529,164,947]" captionTargetId="figure-663@4.[867,1528,166,911]" captionTargetPageId="4" captionText="Fig. 3. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Anterior view of the facial region. B. Posterior view. C. Ventral view of the right squamosal." figureDoi="http://doi.org/10.5281/zenodo.10626273" httpUri="https://zenodo.org/record/10626273/files/figure.png" pageId="3" pageNumber="4">3A</figureCitation>
|
||
). The left eminence is narrower (maximum width
|
||
<quantity id="E92510DCFFF8FFB5FE4BFAA1EC9BFA8B" box="[410,489,1336,1361]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="4" unit="mm" value="22.0">22 mm</quantity>
|
||
for
|
||
<quantity id="E92510DCFFF8FFB5FDCDFAA1EF1EFA8B" box="[540,620,1336,1361]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="4" unit="mm" value="25.0">25 mm</quantity>
|
||
on the right) and higher (maximum height relative to the lateral maxilla
|
||
<quantity id="E92510DCFFF8FFB5FFB3FAE2EDC4FA4E" box="[98,182,1403,1428]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="4" unit="mm" value="7.5">7.5 mm</quantity>
|
||
for
|
||
<quantity id="E92510DCFFF8FFB5FF35FAE2EC51FA4E" box="[228,291,1403,1428]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="4" unit="mm" value="6.0">6 mm</quantity>
|
||
on the right) than the right eminence. Each eminence is transversely and longitudinally convex, with the lateral margin distinctly overhanging the posterolateral sulcus and part of the maxilla on the left side). The right eminence is crossed by a thin longitudinal groove, tentatively interpreted as the additional sulcus observed in extant phocoenids and
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FFB3F9D8EF53F981" authority="(Murakami et al. 2012 b)" baseAuthorityName="Murakami" baseAuthorityYear="2012" box="[98,545,1601,1627]" class="Mammalia" family="Phocoenidae" genus="Miophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FFB3F9D8ED89F981" box="[98,251,1601,1627]" italics="true" pageId="3" pageNumber="4">Miophocaena</emphasis>
|
||
(
|
||
<bibRefCitation id="4A4CC0C8FFF8FFB5FEDDF9DBEF6BF981" author="Murakami, M. & Shimada, C. & Hikida, Y. & Hirano, H." box="[268,537,1602,1627]" pageId="3" pageNumber="4" pagination="1172 - 1185" refId="ref9426" refString="Murakami, M., Shimada, C., Hikida, Y., and Hirano, H. 2012 b. Two new extinct basal phocoenids (Cetacea, Odontoceti, Delphinoidea), from the upper Miocene Koetoi Formation of Japan and their phylogenetic significance. Journal of Vertebrate Paleontology 32: 1172 - 1185." type="journal article" year="2012">Murakami et al. 2012b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. An alternative, but less likely interpretation would be that this groove is due to a longitudinal fracture of the eminence following the path of an underlying canal inside the premaxilla. A wider groove is exposed only on the posterior part of the left eminence, the dorsal exposure of which was possibly increased due to surface wear. The posteromedial corner of each eminence is truncated (slightly damaged on the left side), as it is in most extinct phocoenids and the fossil delphinoid
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FD95F8D2EFE4F8BF" authorityName="Barnes" authorityYear="1984" box="[580,662,1867,1893]" class="Mammalia" family="Albireonidae" genus="Albireo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FD95F8D2EFE4F8BF" box="[580,662,1867,1893]" italics="true" pageId="3" pageNumber="4">Albireo</emphasis>
|
||
</taxonomicName>
|
||
. This feature most likely indicates the presence of maxillary ossicles on the anterior margin of the bony nares, as observed in extant phocoenids (
|
||
<bibRefCitation id="4A4CC0C8FFF8FFB5FC69FF39E93EFF63" author="Barnes, L. G." box="[952,1100,160,185]" pageId="3" pageNumber="4" pagination="1 - 46" refId="ref8032" refString="Barnes, L. G. 1984. Fossil odontocetes (Mammalia: Cetacea) from the Almejas Formation, Isla Cedros, Mexico. PaleoBios 42: 1 - 46." type="journal article" year="1984">Barnes 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFF8FFB5FB8BFF39E871FF63" author="Muizon, C. de" box="[1114,1283,160,185]" pageId="3" pageNumber="4" pagination="159 - 227" refId="ref9248" refString="Muizon, C. de 1988 a. Les relations phylogenetiques des Delphinida. Annales de Paleontologie 74: 159 - 227." type="journal article" year="1988">Muizon 1988a</bibRefCitation>
|
||
). Although wider on the right side of the skull, the portion of the premaxilla along the bony nares is narrow and ends before the posterolateral corner of the naris. A similar condition is observed in all phocoenids, except
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FBFFFE87E985FEE2" authorityName="Murakami, Shimada, Hikida & Hirano" authorityYear="2012" box="[1070,1271,286,312]" class="Mammalia" family="Phocoenidae" genus="Archaeophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FBFFFE87E985FEE2" box="[1070,1271,286,312]" italics="true" pageId="3" pageNumber="4">Archaeophocaena</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FAD9FE87EEF2FE82" baseAuthorityName="Ichishima and Kimura" baseAuthorityYear="2009" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="minutus">
|
||
<emphasis id="1CA9612BFFF8FFB5FAD9FE87EEF2FE82" italics="true" pageId="3" pageNumber="4">Haborophocoena minutus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FC6AFEA7E921FE82" box="[955,1107,318,344]" class="Mammalia" family="Phocoenidae" genus="Miophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FC6AFEA7E921FE82" box="[955,1107,318,344]" italics="true" pageId="3" pageNumber="4">Miophocaena</emphasis>
|
||
</taxonomicName>
|
||
, which display premaxillae wider and longer towards the vertex (
|
||
<bibRefCitation id="4A4CC0C8FFF8FFB5FB57FEF9E8B7FEA3" author="Ichishima, H. & Kimura, M." box="[1158,1477,352,377]" pageId="3" pageNumber="4" pagination="855 - 874" refId="ref8993" refString="Ichishima, H. and Kimura, M. 2009. A new species of Haborophocoena, an Early Pliocene phocoenid cetacean from Hokkaido, Japan. Marine Mammal Science 25: 855 - 874." type="journal article" year="2009">Ichishima and Kimura 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFF8FFB5FCF9FE19E95BFE43" author="Murakami, M. & Shimada, C. & Hikida, Y. & Hirano, H." box="[808,1065,384,409]" pageId="3" pageNumber="4" pagination="1172 - 1185" refId="ref9426" refString="Murakami, M., Shimada, C., Hikida, Y., and Hirano, H. 2012 b. Two new extinct basal phocoenids (Cetacea, Odontoceti, Delphinoidea), from the upper Miocene Koetoi Formation of Japan and their phylogenetic significance. Journal of Vertebrate Paleontology 32: 1172 - 1185." type="journal article" year="2012">Murakami et al. 2012b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="7AA2EDB1FFF8FFB5FFB3FF39EE72FF08" ID-Table-UUID="7AA2EDB1FFF8FFB5FFB3FF39EE72FF08" httpUri="http://table.plazi.org/id/7AA2EDB1FFF8FFB5FFB3FF39EE72FF08" pageId="3" pageNumber="4" startId="3.[98,151,160,181]" targetBox="[105,688,236,1060]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="2E62BD39FFF8FFB5FFB3FF39EE72FF08" blockId="3.[98,770,159,210]" pageId="3" pageNumber="4">
|
||
Table 1. Measurements (in mm) on the skull of
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FDCEFF06EE70FF6F" authority="Colpaert & Bosselaers & Lambert, 2015" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[543,770,159,181]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="gigaseorum" status="gen. et sp. nov.">
|
||
<emphasis id="1CA9612BFFF8FFB5FDCEFF06EE70FF6F" box="[543,770,159,181]" italics="true" pageId="3" pageNumber="4">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="079ADC50FFF8FFB5FFB3FF24ED80FF08" box="[98,242,189,210]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
IRSNB M. 2171 (holotype). e, estimate; + incomplete
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7AA2EDB1FFF8FFB5FCD1FF24EE76FF08" box="[768,772,189,210]" pageId="3" pageNumber="4" startId="3.[768,772,189,210]" targetType="table">
|
||
<paragraph id="2E62BD39FFF8FFB5FCD1FF24EE76FF08" blockId="3.[768,772,189,210]" box="[768,772,189,210]" pageId="3" pageNumber="4">.</paragraph>
|
||
</caption>
|
||
<paragraph id="2E62BD39FFF8FFB5FFB8FF75EFD6FBFE" pageId="3" pageNumber="4">
|
||
<table id="5CDD4F99FFF80049FFB8FF75EFC2FBFE" box="[105,688,236,1060]" gridcols="2" gridrows="21" pageId="3" pageNumber="4">
|
||
<tr id="90EDBF7BFFF80049FFB8FF75EFC2FEDB" box="[105,688,236,257]" gridrow="0" pageId="3" pageNumber="4">
|
||
<th id="D33CD607FFF80049FFB8FF75EF59FEDB" box="[105,555,236,257]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">Condylobasal length</th>
|
||
<th id="D33CD607FFF80049FD50FF75EFC2FEDB" box="[641,688,236,257]" gridcol="1" gridrow="0" pageId="3" pageNumber="4">e300</th>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FE95EFC2FEFB" box="[105,688,268,289]" gridrow="1" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FE95EF59FEFB" box="[105,555,268,289]" gridcol="0" gridrow="1" pageId="3" pageNumber="4">Rostrum length as preserved</td>
|
||
<td id="D33CD607FFF80049FD50FE95EFC2FEFB" box="[641,688,268,289]" gridcol="1" gridrow="1" pageId="3" pageNumber="4">+66</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FEB5EFC2FE87" box="[105,688,300,349]" gridrow="2" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FEB5EF59FE87" box="[105,555,300,349]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">Distance between antorbital notch and posterior end of alveolar groove</td>
|
||
<td id="D33CD607FFF80049FD50FEB5EFC2FE87" box="[641,688,300,349]" gridcol="1" gridrow="2" pageId="3" pageNumber="4">33</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FEFEEFC2FEA6" box="[105,688,359,380]" gridrow="3" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FEFEEF59FEA6" box="[105,555,359,380]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">Width of rostrum at base</td>
|
||
<td id="D33CD607FFF80049FD50FEFEEFC2FEA6" box="[641,688,359,380]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">e73</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FE1EEFC2FE46" box="[105,688,391,412]" gridrow="4" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FE1EEF59FE46" box="[105,555,391,412]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">Width of premaxillae at rostrum base</td>
|
||
<td id="D33CD607FFF80049FD50FE1EEFC2FE46" box="[641,688,391,412]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">40</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FE3EEFC2FE02" box="[105,688,423,472]" gridrow="5" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FE3EEF59FE02" box="[105,555,423,472]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">Longitudinal distance between nuchal crest and antorbital notches</td>
|
||
<td id="D33CD607FFF80049FD50FE3EEFC2FE02" box="[641,688,423,472]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">122</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FE7AEFC2FDCE" box="[105,688,483,532]" gridrow="6" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FE7AEF59FDCE" box="[105,555,483,532]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">Longitudinal distance between anterior margin of bony nares and antorbital notches</td>
|
||
<td id="D33CD607FFF80049FD50FE7AEFC2FDCE" box="[641,688,483,532]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">57</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FD87EFC2FD95" box="[105,688,542,591]" gridrow="7" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FD87EF59FD95" box="[105,555,542,591]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">Distance between premaxillary foramen and anterior margin of bony nares</td>
|
||
<td id="D33CD607FFF80049FD50FD87EFC2FD95" box="[641,688,542,591]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">44</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FDC3EFC2FDB5" box="[105,688,602,623]" gridrow="8" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FDC3EF59FDB5" box="[105,555,602,623]" gridcol="0" gridrow="8" pageId="3" pageNumber="4">Preorbital width of skull</td>
|
||
<td id="D33CD607FFF80049FD50FDC3EFC2FDB5" box="[641,688,602,623]" gridcol="1" gridrow="8" pageId="3" pageNumber="4">e115</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FDE3EFC2FD55" box="[105,688,634,655]" gridrow="9" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FDE3EF59FD55" box="[105,555,634,655]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">Postorbital width of skull</td>
|
||
<td id="D33CD607FFF80049FD50FDE3EFC2FD55" box="[641,688,634,655]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">e148</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FD03EFC2FD75" box="[105,688,666,687]" gridrow="10" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FD03EF59FD75" box="[105,555,666,687]" gridcol="0" gridrow="10" pageId="3" pageNumber="4">Maximum width of premaxillae on cranium</td>
|
||
<td id="D33CD607FFF80049FD50FD03EFC2FD75" box="[641,688,666,687]" gridcol="1" gridrow="10" pageId="3" pageNumber="4">52</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FD23EFC2FD15" box="[105,688,698,719]" gridrow="11" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FD23EF59FD15" box="[105,555,698,719]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">Width of bony nares</td>
|
||
<td id="D33CD607FFF80049FD50FD23EFC2FD15" box="[641,688,698,719]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">40</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FD40EFC2FD34" box="[105,688,729,750]" gridrow="12" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FD40EF59FD34" box="[105,555,729,750]" gridcol="0" gridrow="12" pageId="3" pageNumber="4">Maximum width of fossae for nasals</td>
|
||
<td id="D33CD607FFF80049FD50FD40EFC2FD34" box="[641,688,729,750]" gridcol="1" gridrow="12" pageId="3" pageNumber="4">37</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FD60EFC2FCD4" box="[105,688,761,782]" gridrow="13" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FD60EF59FCD4" box="[105,555,761,782]" gridcol="0" gridrow="13" pageId="3" pageNumber="4">Length of orbit</td>
|
||
<td id="D33CD607FFF80049FD50FD60EFC2FCD4" box="[641,688,761,782]" gridcol="1" gridrow="13" pageId="3" pageNumber="4">e50</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FC80EFC2FCF4" box="[105,688,793,814]" gridrow="14" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FC80EF59FCF4" box="[105,555,793,814]" gridcol="0" gridrow="14" pageId="3" pageNumber="4">Length of temporal fossa</td>
|
||
<td id="D33CD607FFF80049FD50FC80EFC2FCF4" box="[641,688,793,814]" gridcol="1" gridrow="14" pageId="3" pageNumber="4">e75</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FCA0EFC2FC5C" box="[105,688,825,902]" gridrow="15" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FCA0EF59FC5C" box="[105,555,825,902]" gridcol="0" gridrow="15" pageId="3" pageNumber="4">Distance between anterior tip of zygomatic process of squamosal and ventral tip of postglenoid process</td>
|
||
<td id="D33CD607FFF80049FD50FCA0EFC2FC5C" box="[641,688,825,902]" gridcol="1" gridrow="15" pageId="3" pageNumber="4">42</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FC09EFC2FC7F" box="[105,688,912,933]" gridrow="16" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FC09EF59FC7F" box="[105,555,912,933]" gridcol="0" gridrow="16" pageId="3" pageNumber="4">Distance between jugular notches</td>
|
||
<td id="D33CD607FFF80049FD50FC09EFC2FC7F" box="[641,688,912,933]" gridcol="1" gridrow="16" pageId="3" pageNumber="4">e85</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FC29EFC2FC1F" box="[105,688,944,965]" gridrow="17" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FC29EF59FC1F" box="[105,555,944,965]" gridcol="0" gridrow="17" pageId="3" pageNumber="4">Width of occipital condyles</td>
|
||
<td id="D33CD607FFF80049FD50FC29EFC2FC1F" box="[641,688,944,965]" gridcol="1" gridrow="17" pageId="3" pageNumber="4">74</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FC49EFC2FC3F" box="[105,688,976,997]" gridrow="18" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FC49EF59FC3F" box="[105,555,976,997]" gridcol="0" gridrow="18" pageId="3" pageNumber="4">Maximum width of right occipital condyle</td>
|
||
<td id="D33CD607FFF80049FD50FC49EFC2FC3F" box="[641,688,976,997]" gridcol="1" gridrow="18" pageId="3" pageNumber="4">29</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FC69EFC2FBDF" box="[105,688,1008,1029]" gridrow="19" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FC69EF59FBDF" box="[105,555,1008,1029]" gridcol="0" gridrow="19" pageId="3" pageNumber="4">Width of foramen magnum</td>
|
||
<td id="D33CD607FFF80049FD50FC69EFC2FBDF" box="[641,688,1008,1029]" gridcol="1" gridrow="19" pageId="3" pageNumber="4">37</td>
|
||
</tr>
|
||
<tr id="90EDBF7BFFF80049FFB8FB96EFC2FBFE" box="[105,688,1039,1060]" gridrow="20" pageId="3" pageNumber="4">
|
||
<td id="D33CD607FFF80049FFB8FB96EF59FBFE" box="[105,555,1039,1060]" gridcol="0" gridrow="20" pageId="3" pageNumber="4">Height of foramen magnum</td>
|
||
<td id="D33CD607FFF80049FD50FB96EFC2FBFE" box="[641,688,1039,1060]" gridcol="1" gridrow="20" pageId="3" pageNumber="4">36</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFF8FFB5FCF9FE30E9CDFA9C" blockId="3.[808,1482,425,1960]" pageId="3" pageNumber="4">
|
||
<emphasis id="1CA9612BFFF8FFB5FCF9FE30EE0DFE19" box="[808,895,425,451]" italics="true" pageId="3" pageNumber="4">Maxilla</emphasis>
|
||
.—Only the right antorbital notch is preserved. In dorsal view, the notch is anterolaterally widely open (
|
||
<figureCitation id="B6E6A1BCFFF8FFB5FA4DFE52EE4BFDDE" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
). Consisting of the maxilla and lacrimal, the posterior wall is directed transversely, lacking an anterolateral projection. This major difference with
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FBB9FDB3E869FD9E" authorityName="Lambert" authorityYear="2008" box="[1128,1307,554,580]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FBB9FDB3E869FD9E" box="[1128,1307,554,580]" italics="true" pageId="3" pageNumber="4">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
is observed in most extant phocoenids,
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FB95FDD3E9BCFDBE" authorityName="de Muizon" authorityYear="1988" box="[1092,1230,586,612]" class="Mammalia" family="Phocoenidae" genus="Australithax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FB95FDD3E9BCFDBE" box="[1092,1230,586,612]" italics="true" pageId="3" pageNumber="4">Australithax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FB0FFDD2E825FDBE" authorityName="de Muizon" authorityYear="1986" box="[1246,1367,587,612]" class="Mammalia" family="Phocoenidae" genus="Lomacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FB0FFDD2E825FDBE" box="[1246,1367,587,612]" italics="true" pageId="3" pageNumber="4">Lomacetus</emphasis>
|
||
</taxonomicName>
|
||
, and possibly
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FCB4FDF3E952FD5E" authorityName="Barnes" authorityYear="1985" box="[869,1056,618,644]" class="Mammalia" family="Phocoenidae" genus="Salumiphocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFF8FFB5FCB4FDF3E952FD5E" box="[869,1056,618,644]" italics="true" pageId="3" pageNumber="4">Salumiphocaena</emphasis>
|
||
</taxonomicName>
|
||
. Although incomplete in this area, the left maxilla bears three dorsal infraorbital foramina posteromedial to the corresponding notch: two small anterior foramina with a diameter not exceeding
|
||
<quantity id="E92510DCFFF8FFB5FB4CFD55E9AEFD3F" box="[1181,1244,716,741]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="4" unit="mm" value="2.0">2 mm</quantity>
|
||
and a larger posterior foramen with a transverse diameter of
|
||
<quantity id="E92510DCFFF8FFB5FB06FD75E869FCDF" box="[1239,1307,748,773]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="4" unit="mm" value="6.0">6 mm</quantity>
|
||
. The anteromedial-most foramen is conjoined with a shallow groove anterolaterally. On the right side, a large opening posteromedial to the antorbital notch is probably due to damage of an area pierced by several dorsal infraorbital foramina. In the orbital region, the dorsal surface of the maxilla is smooth and moderately convex. Its lateral margin is thin. A large posterior dorsal infraorbital foramen is in line transversely with the level of the lost postorbital process of the frontal, and
|
||
<quantity id="E92510DCFFF8FFB5FAAAFC74E8BBFBDC" box="[1403,1481,1005,1030]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="4" unit="mm" value="12.0">12 mm</quantity>
|
||
lateral to the premaxilla. The left maxilla is exposed dorsally as a narrow strip medial to the corresponding premaxilla on the anterolateral corner of the bony naris, but this exposure may have been accentuated by wear of the overlying premaxilla. The dorsal surface of the maxilla is distinctly convex between the temporal fossa and the vertex. The maxilla reaches the nuchal crest over most of its posterior border, except in its posteromedial corner, where the concave dorsal surface of the frontal is exposed. The maxilla only contacts the frontal boss in the anterior part of the latter.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFF8FFB5FC9FFAD7E9A7F932" blockId="3.[808,1482,425,1960]" pageId="3" pageNumber="4">
|
||
In lateral view, the medial part of the maxilla ascends gradually towards the vertex, although less abruptly than in
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FCF9FA15EEA0FA7D" box="[808,978,1420,1447]" class="Mammalia" family="Phocoenidae" genus="Phocoena" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="1CA9612BFFF8FFB5FCF9FA15EEEAFA7C" box="[808,920,1420,1446]" italics="true" pageId="3" pageNumber="4">Phocoena</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E9DDC6BAFFF8FFB5FBC1FA15E999FA7C" baseAuthorityName="True" baseAuthorityYear="1885" box="[1040,1259,1420,1446]" class="Mammalia" family="Phocoenidae" genus="Phocoenoides" kingdom="Animalia" order="Cetacea" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="dalli">
|
||
<emphasis id="1CA9612BFFF8FFB5FBC1FA15E999FA7C" box="[1040,1259,1420,1446]" italics="true" pageId="3" pageNumber="4">Phocoenoides dalli</emphasis>
|
||
</taxonomicName>
|
||
, whereas the bone slopes down posterolaterally from the orbit region to the postorbital process. The posterior end of the alveolar groove is preserved on the left side, beginning
|
||
<quantity id="E92510DCFFF8FFB5FB0BFA77E85AF9DD" box="[1242,1320,1518,1543]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="4" unit="mm" value="33.0">33 mm</quantity>
|
||
anterior to the antorbital notch. However, the spongy bone of this area is partially worn and no detail of the alveoli (if originally present) is preserved. The palatal surface of the maxillae between the anterior tips of the palatines is wide and flat, as in other phocoenids and monodontids. On each maxilla, a longitudinal sulcus indicates the presence of a major palatine foramen anterior to the maxilla-palatine suture.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFF8FFB2FC9FF976EFBEFEE0" blockId="3.[808,1482,425,1960]" lastBlockId="4.[151,825,160,314]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
An anteroposteriorly directed thin groove on the lateral side of the maxilla originates medial to the jugal and extends for a short distance (
|
||
<quantity id="E92510DCFFF8FFB5FC1CF8B6E96FF892" box="[973,1053,1839,1864]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="4" unit="mm" value="25.0">25 mm</quantity>
|
||
) on the rostral base (
|
||
<figureCitation id="B6E6A1BCFFF8FFB5FAD1F8B6E830F892" box="[1280,1346,1839,1864]" captionStart="Fig" captionStartId="6.[151,184,505,525]" captionTargetBox="[162,1521,162,482]" captionTargetPageId="6" captionText="Fig. 6. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Right ventrolateral view of the palate and orbital regions. Photo (A) and corresponding line drawing (B). Dotted lines indicate poorly visible sutures. Dark grey surface indicates fracture surfaces." figureDoi="http://doi.org/10.5281/zenodo.10626279" httpUri="https://zenodo.org/record/10626279/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
|
||
). According to
|
||
<bibRefCitation id="4A4CC0C8FFF8FFB5FC94F8D6E918F8B2" author="Mead, J. G. & Fordyce, R. E." box="[837,1130,1871,1896]" pageId="3" pageNumber="4" pagination="1 - 248" refId="ref9166" refString="Mead, J. G. and Fordyce, R. E. 2009. The therian skull: a lexicon with emphasis on the odontocetes. Smithsonian Contributions to Zoology 627: 1 - 248." type="journal article" year="2009">Mead and Fordyce (2009)</bibRefCitation>
|
||
, this groove may correspond to the anterior extent of the zygomatic recess, the latter housing the maxillary process of the jugal. Ventromedial to this groove, a broader and anteroposteriorly elongated depression is interpreted as a fossa for an anterior extent of the pterygoid sinus, possibly homologous to the anterior sinus fossa present in extant delphinids but presumably not in extant phocoenids (
|
||
<bibRefCitation id="4A4CC0C8FFFFFFB2FF4FFEB8ECD7FEE0" author="Fraser, F. C. & Purves, P. E." box="[158,421,289,314]" pageId="4" pageNumber="5" pagination="1 - 140" refId="ref8718" refString="Fraser, F. C. and Purves, P. E. 1960. Hearing in cetaceans: Evolution of the accessory air sacs and the structure of the outer and middle ear in recent cetaceans. Bulletin of the British Museum (Natural History), Zoology 7: 1 - 140." type="journal article" year="1960">Fraser and Purves 1960</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFFFFFB2FE61FEB8EFB2FEE0" author="Mead, J. G. & Fordyce, R. E." box="[432,704,289,314]" pageId="4" pageNumber="5" pagination="1 - 248" refId="ref9166" refString="Mead, J. G. and Fordyce, R. E. 2009. The therian skull: a lexicon with emphasis on the odontocetes. Smithsonian Contributions to Zoology 627: 1 - 248." type="journal article" year="2009">Mead and Fordyce 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFFFFFB2FF46FED3EFD0FDDC" blockId="4.[151,824,330,518]" pageId="4" pageNumber="5">
|
||
<emphasis id="1CA9612BFFFFFFB2FF46FED3EC52FEBE" box="[151,288,330,356]" italics="true" pageId="4" pageNumber="5">Mesethmoid</emphasis>
|
||
.—Although the mesethmoid is not visible in dorsal view between the premaxillary eminences, it forms a thin and low nasal septum between the bony nares. Below the frontal boss, the transverse plate-like mesethmoid forms the posterior wall of the bony nares. This plate is proportionally wider and less erect than in extant phocoenids.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFFFFFB2FF46FD8EEFFCFC0F" blockId="4.[151,825,535,981]" pageId="4" pageNumber="5">
|
||
<emphasis id="1CA9612BFFFFFFB2FF46FD8EEDAAFDEB" box="[151,216,535,561]" italics="true" pageId="4" pageNumber="5">Nasal</emphasis>
|
||
.—As in the
|
||
<typeStatus id="F166039BFFFFFFB2FEA1FD81ECA0FDEB" box="[368,466,536,561]" pageId="4" pageNumber="5">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FE2CFD8EEE4AFDEB" authorityName="Lambert" authorityYear="2008" box="[509,824,535,561]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="bosselaersi">
|
||
<emphasis id="1CA9612BFFFFFFB2FE2CFD8EEE4AFDEB" box="[509,824,535,561]" italics="true" pageId="4" pageNumber="5">Septemtriocetus bosselaersi</emphasis>
|
||
</taxonomicName>
|
||
and several other fossil phocoenid specimens, both nasals are lost. Anteroventrolateral to the frontal boss, a shallow fossa is observed above the mesethmoid on each side of the specimen (
|
||
<figureCitation id="B6E6A1BCFFFFFFB2FEC2FD00EC28FD68" box="[275,346,665,690]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
). This fossa is crossed by an oblique low ridge descending from the frontal boss ventrolaterally and the deepest portion of each fossa is located dorsolaterally. Similar features on a juvenile skull of
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FDB3FD60EDC1FCE9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Mammalia" family="Phocoenidae" genus="Phocoena" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="phocoena">
|
||
<emphasis id="1CA9612BFFFFFFB2FDB3FD60EDC1FCE9" italics="true" pageId="4" pageNumber="5">Phocoena phocoena</emphasis>
|
||
</taxonomicName>
|
||
, of which one nasal could be removed (
|
||
<figureCitation id="B6E6A1BCFFFFFFB2FD53FC83EFBAFCE9" box="[642,712,794,819]" captionStart="Fig" captionStartId="4.[862,895,1532,1552]" captionTargetBox="[945,1450,1109,1506]" captionTargetId="figure-801@4.[946,1448,1106,1506]" captionTargetPageId="4" captionText="Fig. 4. Anterior view of the facial region of the skull of the extant porpoise Phocoena phocoena (Linnaeus, 1758) with both nasals present (A) and right nasal removed (B), to show the corresponding fossae on the corresponding frontal and maxilla. Dotted lines indicate the outline of the right nasal, both when present and when removed." figureDoi="http://doi.org/10.5281/zenodo.10626275" httpUri="https://zenodo.org/record/10626275/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
|
||
), indicate that these fossae in the frontals held the nasals in life. Consequently, originally each wide nasal extended much more laterally than the frontal boss, with a lateral limit in line with the lateral margin of the bony nares. The mesethmoid formed a dorsomedial projection between the nasals.
|
||
</paragraph>
|
||
<caption id="7AA2EDB1FFFFFFB2FC8FFC54E960FBEF" ID-DOI="http://doi.org/10.5281/zenodo.10626273" ID-Zenodo-Dep="10626273" httpUri="https://zenodo.org/record/10626273/files/figure.png" pageId="4" pageNumber="5" startId="4.[862,894,973,993]" targetBox="[868,1529,164,947]" targetPageId="4" targetType="figure">
|
||
<paragraph id="2E62BD39FFFFFFB2FC8FFC54E960FBEF" blockId="4.[862,1536,972,1077]" pageId="4" pageNumber="5">
|
||
Fig. 3. Skull of the phocoenid cetacean
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FB17FC55E8E9FC38" authority="Colpaert & Bosselaers & Lambert, 2015" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[1222,1435,972,994]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="gigaseorum" status="gen. et sp. nov.">
|
||
<emphasis id="1CA9612BFFFFFFB2FB17FC55E8E9FC38" box="[1222,1435,972,994]" italics="true" pageId="4" pageNumber="5">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="079ADC50FFFFFFB2FA73FC54EEF7FC27" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
from the Pliocene of northern Belgium IRSNB M. 2171 (holotype).
|
||
<emphasis id="1CA9612BFFFFFFB2FC8FFB9DEE1CFBC3" bold="true" box="[862,878,1028,1049]" pageId="4" pageNumber="5">A</emphasis>
|
||
. Anterior view of the facial region.
|
||
<emphasis id="1CA9612BFFFFFFB2FB63FB9DE9B3FBC3" bold="true" box="[1202,1217,1028,1049]" pageId="4" pageNumber="5">B</emphasis>
|
||
. Posterior view.
|
||
<emphasis id="1CA9612BFFFFFFB2FA89FB9DE81AFBC0" bold="true" box="[1368,1384,1028,1050]" pageId="4" pageNumber="5">C</emphasis>
|
||
. Ventral view of the right squamosal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2E62BD39FFFFFFB2FF46FC7CEE5EF9DE" blockId="4.[151,824,997,1960]" pageId="4" pageNumber="5">
|
||
<emphasis id="1CA9612BFFFFFFB2FF46FC7CED9EFC25" box="[151,236,997,1023]" italics="true" pageId="4" pageNumber="5">Frontal</emphasis>
|
||
.—Posterior to the fossae for the nasals, the frontal boss is longer than wide (
|
||
<figureCitation id="B6E6A1BCFFFFFFB2FE69FB9EEC8EFBFA" box="[440,508,1031,1056]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
), as in
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FD9FFB9FEE73FBFA" authorityName="Lambert" authorityYear="2008" box="[590,769,1030,1056]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FD9FFB9FEE73FBFA" box="[590,769,1030,1056]" italics="true" pageId="4" pageNumber="5">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
, and transversely narrower than in any other phocoenid, somewhat more similar to the monodontid
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FE2AFBDFEFDBFBBA" authorityName="Lacepede" authorityYear="1804" box="[507,681,1094,1120]" class="Mammalia" family="Monodontidae" genus="Delphinapterus" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FE2AFBDFEFDBFBBA" box="[507,681,1094,1120]" italics="true" pageId="4" pageNumber="5">Delphinapterus</emphasis>
|
||
</taxonomicName>
|
||
. It should be noted that this area is unknown in
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FDC0FBFFEFC9FB5A" authorityName="Murakami, Shimada, Hikida & Hirano" authorityYear="2012" box="[529,699,1126,1152]" class="Mammalia" family="Phocoenidae" genus="Pterophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FDC0FBFFEFC9FB5A" box="[529,699,1126,1152]" italics="true" pageId="4" pageNumber="5">Pterophocaena</emphasis>
|
||
</taxonomicName>
|
||
and poorly known in
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FED8FB1EECA6FB7B" authorityName="Murakami, Shimada, Hikida & Hirano" authorityYear="2012" box="[265,468,1159,1185]" class="Mammalia" family="Phocoenidae" genus="Archaeophocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FED8FB1EECA6FB7B" box="[265,468,1159,1185]" italics="true" pageId="4" pageNumber="5">Archaeophocaena</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FDDDFB1EEFA0FB7B" authorityName="Ichishima & Kimura" authorityYear="2000" box="[524,722,1159,1185]" class="Mammalia" family="Phocoenidae" genus="Numataphocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FDDDFB1EEFA0FB7B" box="[524,722,1159,1185]" italics="true" pageId="4" pageNumber="5">Numataphocoena</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="4A4CC0C8FFFFFFB2FD30FB11ECF6FB1B" author="Ichishima, H. & Kimura, M." pageId="4" pageNumber="5" pagination="561 - 576" refId="ref8911" refString="Ichishima, H. and Kimura, M. 2000. A new fossil porpoise (Cetacea; Delphinoidea; Phocoenidae) from the early Pliocene Horokaoshirarika Formation, Hokkaido, Japan. Journal of Vertebrate Paleontology 20: 561 - 576." type="journal article" year="2000">Ichishima and Kimura 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFFFFFB2FE5CFB31EFFEFB1B" author="Murakami, M. & Shimada, C. & Hikida, Y. & Hirano, H." box="[397,652,1192,1217]" pageId="4" pageNumber="5" pagination="1172 - 1185" refId="ref9426" refString="Murakami, M., Shimada, C., Hikida, Y., and Hirano, H. 2012 b. Two new extinct basal phocoenids (Cetacea, Odontoceti, Delphinoidea), from the upper Miocene Koetoi Formation of Japan and their phylogenetic significance. Journal of Vertebrate Paleontology 32: 1172 - 1185." type="journal article" year="2012">Murakami et al. 2012b</bibRefCitation>
|
||
). The right wall of this transversely pinched frontal boss is relatively high, with a dorsal margin forming the top of the skull and slightly overhanging the rest of the wall. This right wall is located far beyond the sagittal plane of the skull on the left side, as is also seen in
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FE99FAD1EF79FAB8" authorityName="Ichishima & Kimura" authorityYear="2005" box="[328,523,1352,1378]" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FE99FAD1EF79FAB8" box="[328,523,1352,1378]" italics="true" pageId="4" pageNumber="5">Haborophocoena</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FD98FAD0EF8EFAB9" authorityName="Lambert" authorityYear="2008" box="[585,764,1353,1379]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FD98FAD0EF8EFAB9" box="[585,764,1353,1379]" italics="true" pageId="4" pageNumber="5">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
. The left wall slopes more gradually towards the corresponding maxilla. Although the interfrontal suture is visible on the anterior surface of the frontal boss, the state of preservation of the posterior part of the boss precludes a detailed description, including the potential presence of an interparietal (
|
||
<figureCitation id="B6E6A1BCFFFFFFB2FD08FA72EE6DF9DE" box="[729,799,1515,1540]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="7AA2EDB1FFFFFFB2FC8FFA65E985F95A" ID-DOI="http://doi.org/10.5281/zenodo.10626275" ID-Zenodo-Dep="10626275" httpUri="https://zenodo.org/record/10626275/files/figure.png" pageId="4" pageNumber="5" startId="4.[862,895,1532,1552]" targetBox="[945,1450,1109,1506]" targetPageId="4" targetType="figure">
|
||
<paragraph id="2E62BD39FFFFFFB2FC8FFA65E985F95A" blockId="4.[862,1535,1532,1664]" pageId="4" pageNumber="5">
|
||
Fig. 4. Anterior view of the facial region of the skull of the extant porpoise
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FC8FF98FE9B2F9F6" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[862,1216,1558,1580]" class="Mammalia" family="Phocoenidae" genus="Phocoena" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="phocoena">
|
||
<emphasis id="1CA9612BFFFFFFB2FC8FF98FE96BF9F1" box="[862,1049,1558,1579]" italics="true" pageId="4" pageNumber="5">Phocoena phocoena</emphasis>
|
||
(Linnaeus, 1758)
|
||
</taxonomicName>
|
||
with both nasals present (
|
||
<emphasis id="1CA9612BFFFFFFB2FA6CF98FE8BFF9F1" bold="true" box="[1469,1485,1558,1579]" pageId="4" pageNumber="5">A</emphasis>
|
||
) and right nasal removed (
|
||
<emphasis id="1CA9612BFFFFFFB2FBF4F9ABE946F99D" bold="true" box="[1061,1076,1586,1607]" pageId="4" pageNumber="5">B</emphasis>
|
||
), to show the corresponding fossae on the corresponding frontal and maxilla. Dotted lines indicate the outline of the right nasal, both when present and when removed.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2E62BD39FFFFFFB2FF6CF995EF8EF97C" blockId="4.[151,824,997,1960]" pageId="4" pageNumber="5">
|
||
In lateral view, the preorbital process of the frontal is barely thickened, roughly continuous with the supraorbital portion (
|
||
<figureCitation id="B6E6A1BCFFFFFFB2FF24F9D5EC45F9BF" box="[245,311,1612,1637]" captionStart="Fig" captionStartId="5.[98,131,929,949]" captionTargetBox="[98,1477,158,906]" captionTargetId="figure-619@5.[97,1481,157,580]" captionTargetPageId="5" captionText="Fig. 5. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Right lateral view, photo (A1) and corresponding line drawing (A2). Dotted lines indicate the dorsal margin of the temporal fossa (A1), the reconstructed right postorbital process, and poorly visible sutures (A2). B. Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.10626277" httpUri="https://zenodo.org/record/10626277/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
|
||
). Although no postorbital process is preserved, the broken surface at the base of the right process indicates a process in line with the anterior half of the bony nares.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFFFFFB3FF6CF934ECC2FA74" blockId="4.[151,824,997,1960]" lastBlockId="5.[98,771,1040,1454]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
In ventral view of the orbit region, a deep fossa for the postorbital lobe of the pterygoid sinus excavates the frontal medial to the infratemporal crest (
|
||
<figureCitation id="B6E6A1BCFFFFFFB2FDDAF977EF3FF8DD" box="[523,589,1774,1799]" captionStart="Fig" captionStartId="6.[151,184,505,525]" captionTargetBox="[162,1521,162,482]" captionTargetPageId="6" captionText="Fig. 6. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Right ventrolateral view of the palate and orbital regions. Photo (A) and corresponding line drawing (B). Dotted lines indicate poorly visible sutures. Dark grey surface indicates fracture surfaces." figureDoi="http://doi.org/10.5281/zenodo.10626279" httpUri="https://zenodo.org/record/10626279/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
|
||
). This fossa is deeper than in the
|
||
<typeStatus id="F166039BFFFFFFB2FECAF897EC0FF8FD" box="[283,381,1806,1831]" pageId="4" pageNumber="5">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FE77F894EF92F8FC" authorityName="Lambert" authorityYear="2008" box="[422,736,1804,1831]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="bosselaersi">
|
||
<emphasis id="1CA9612BFFFFFFB2FE77F894EF92F8FC" box="[422,736,1804,1831]" italics="true" pageId="4" pageNumber="5">Septemtriocetus bosselaersi</emphasis>
|
||
</taxonomicName>
|
||
, but its extent is variable in extant phocoenids (OL personal observations). The fossa for the preorbital lobe of the pterygoid sinus extends posterodorsally for a short distance above the frontal groove, close to the configuration of the same in
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FD0FF817EEB2F961" authorityName="Lambert" authorityYear="2008" box="[734,960,1698,1960]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FD0FF817EEB2F961" box="[734,960,1698,1960]" italics="true" pageId="4" pageNumber="5">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
. This feature is also present in some extinct phocoenids, as well as some extant delphinids, whereas it is more developed in extant phocoenids (
|
||
<bibRefCitation id="4A4CC0C8FFFFFFB2FB0AF97FE880F925" author="Fraser, F. C. & Purves, P. E." box="[1243,1522,1766,1791]" pageId="4" pageNumber="5" pagination="1 - 140" refId="ref8718" refString="Fraser, F. C. and Purves, P. E. 1960. Hearing in cetaceans: Evolution of the accessory air sacs and the structure of the outer and middle ear in recent cetaceans. Bulletin of the British Museum (Natural History), Zoology 7: 1 - 140." type="journal article" year="1960">Fraser and Purves 1960</bibRefCitation>
|
||
). The roof of the temporal fossa extends for two thirds of the length of the fossa. This roof is proportionally longer anteroposteriorly than in extant phocoenids (most likely linked to the proportionally less transversely inflated cerebral hemispheres), but transversely narrower than in
|
||
<taxonomicName id="E9DDC6BAFFFFFFB2FA93F817E887F872" authorityName="Lambert" authorityYear="2008" box="[1346,1525,1934,1960]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="4" pageNumber="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFFFFB2FA93F817E887F872" box="[1346,1525,1934,1960]" italics="true" pageId="4" pageNumber="5">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="1CA9612BFFFEFFB3FFB3FB89EC75FBF0" box="[98,263,1040,1066]" italics="true" pageId="5" pageNumber="6">Supraoccipital</emphasis>
|
||
.—The nuchal crest is only poorly elevated against the frontal boss, unlike that in
|
||
<taxonomicName id="E9DDC6BAFFFEFFB3FDCDFBA8EFA3FB91" box="[540,721,1073,1099]" class="Mammalia" family="Phocoenidae" genus="Piscolithax" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="1CA9612BFFFEFFB3FDCDFBA8EFE8FB91" box="[540,666,1073,1099]" italics="true" pageId="5" pageNumber="6">Piscolithax</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
and several extant phocoenids (
|
||
<bibRefCitation id="4A4CC0C8FFFEFFB3FE4EFBCBEF4DFBB1" author="Muizon, C. de" box="[415,575,1106,1131]" pageId="5" pageNumber="6" pagination="1 - 188" refId="ref9203" refString="Muizon, C. de 1984. Les Vertebres de la Formation Pisco (Perou). Deuxieme partie: Les Odontocetes (Cetacea, Mammalia) du Pliocene inferieur du Sud-Sacaco. Travaux de l'Institut Francais d'Etudes Andines 27: 1 - 188." type="journal article" year="1984">Muizon 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFFEFFB3FD9EFBCBEF85FBB1" author="Lambert, O." box="[591,759,1106,1131]" pageId="5" pageNumber="6" pagination="863 - 872" refId="ref9029" refString="Lambert, O. 2008. A new porpoise (Cetacea, Odontoceti, Phocoenidae) from the Pliocene of the North Sea. Journal of Vertebrate Paleontology 28: 863 - 872." type="journal article" year="2008">Lambert 2008</bibRefCitation>
|
||
). Lateral to the vertex, the nuchal crest is higher and more acute on the right side (
|
||
<figureCitation id="B6E6A1BCFFFEFFB3FEB7FB0AECADFB76" box="[358,479,1171,1196]" captionStart="Fig" captionStartId="4.[862,894,973,993]" captionTargetBox="[868,1529,164,947]" captionTargetId="figure-663@4.[867,1528,166,911]" captionTargetPageId="4" captionText="Fig. 3. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Anterior view of the facial region. B. Posterior view. C. Ventral view of the right squamosal." figureDoi="http://doi.org/10.5281/zenodo.10626273" httpUri="https://zenodo.org/record/10626273/files/figure.png" pageId="5" pageNumber="6">Fig. 3A, B</figureCitation>
|
||
), overhanging the maxilla and frontal.At the top of the right nuchal crest lies the weakly developed external occipital protuberance. A shallow groove extends from the protuberance towards the foramen magnum. The supraoccipital shield slopes more steeply posteroventrally (mean angle of ca. 50° to the horizontal plane over the first
|
||
<quantity id="E92510DCFFFEFFB3FF14FACCEC6BFAB4" box="[197,281,1365,1390]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="6" unit="mm" value="45.0">45 mm</quantity>
|
||
, taken mediodorsally in lateral view) than in extant phocoenids, due to its less posteriorly developed cerebral hemispheres (
|
||
<figureCitation id="B6E6A1BCFFFEFFB3FE8FFA0CECD0FA74" box="[350,418,1429,1454]" captionStart="Fig" captionStartId="5.[98,131,929,949]" captionTargetBox="[98,1477,158,906]" captionTargetId="figure-619@5.[97,1481,157,580]" captionTargetPageId="5" captionText="Fig. 5. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Right lateral view, photo (A1) and corresponding line drawing (A2). Dotted lines indicate the dorsal margin of the temporal fossa (A1), the reconstructed right postorbital process, and poorly visible sutures (A2). B. Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.10626277" httpUri="https://zenodo.org/record/10626277/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="7AA2EDB1FFFEFFB3FFB3FC38EE6CFC37" ID-DOI="http://doi.org/10.5281/zenodo.10626277" ID-Zenodo-Dep="10626277" httpUri="https://zenodo.org/record/10626277/files/figure.png" pageId="5" pageNumber="6" startId="5.[98,131,929,949]" targetBox="[98,1477,158,906]" targetPageId="5" targetType="figure">
|
||
<paragraph id="2E62BD39FFFEFFB3FFB3FC38EE6CFC37" blockId="5.[98,1482,927,1010]" pageId="5" pageNumber="6">
|
||
Fig. 5. Skull of the phocoenid cetacean
|
||
<taxonomicName id="E9DDC6BAFFFEFFB3FE09FC06EFC0FC6F" authority="Colpaert & Bosselaers & Lambert, 2015" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[472,690,927,949]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="gigaseorum" status="gen. et sp. nov.">
|
||
<emphasis id="1CA9612BFFFEFFB3FE09FC06EFC0FC6F" box="[472,690,927,949]" italics="true" pageId="5" pageNumber="6">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="079ADC50FFFEFFB3FD6BFC38EE38FC6F" box="[698,842,929,949]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
from the Pliocene of northern Belgium IRSNB M. 2171 (holotype).
|
||
<emphasis id="1CA9612BFFFEFFB3FFB3FC22ED00FC0A" bold="true" box="[98,114,955,976]" pageId="5" pageNumber="6">A</emphasis>
|
||
. Right lateral view, photo (A
|
||
<subScript id="B259BF7CFFFEFFB3FEA7FC53EC0FFC0C" attach="left" box="[374,381,970,982]" fontSize="5" pageId="5" pageNumber="6">1</subScript>
|
||
) and corresponding line drawing (A
|
||
<subScript id="B259BF7CFFFEFFB3FD15FC53EFB9FC0C" attach="right" box="[708,715,970,982]" fontSize="5" pageId="5" pageNumber="6">2</subScript>
|
||
). Dotted lines indicate the dorsal margin of the temporal fossa (A
|
||
<subScript id="B259BF7CFFFEFFB3FAC4FC53E86EFC0C" attach="left" box="[1301,1308,970,982]" fontSize="5" pageId="5" pageNumber="6">1</subScript>
|
||
), the reconstructed right postorbital process, and poorly visible sutures (A
|
||
<subScript id="B259BF7CFFFEFFB3FD9EFC7FEF24FC28" attach="left" box="[591,598,998,1010]" fontSize="5" pageId="5" pageNumber="6">2</subScript>
|
||
).
|
||
<emphasis id="1CA9612BFFFEFFB3FDB8FC41EF0AFC37" bold="true" box="[617,632,984,1005]" pageId="5" pageNumber="6">B</emphasis>
|
||
. Left lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2E62BD39FFFEFFB3FFB3FA26E860F9AA" blockId="5.[98,771,1471,1723]" lastBlockId="5.[808,1481,1042,1648]" pageId="5" pageNumber="6">
|
||
<emphasis id="1CA9612BFFFEFFB3FFB3FA26EC4BFA03" box="[98,313,1471,1497]" italics="true" pageId="5" pageNumber="6">Lacrimal and jugal</emphasis>
|
||
.—Only the right lacrimal and part of the corresponding jugal are preserved. In dorsal and lateral views
|
||
<figureCitation id="B6E6A1BCFFFEFFB3FFBBF998EDC8F9C0" box="[106,186,1537,1562]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="B6E6A1BCFFFEFFB3FF16F998EDA4F9C0" box="[199,214,1537,1562]" captionStart="Fig" captionStartId="5.[98,131,929,949]" captionTargetBox="[98,1477,158,906]" captionTargetId="figure-619@5.[97,1481,157,580]" captionTargetPageId="5" captionText="Fig. 5. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Right lateral view, photo (A1) and corresponding line drawing (A2). Dotted lines indicate the dorsal margin of the temporal fossa (A1), the reconstructed right postorbital process, and poorly visible sutures (A2). B. Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.10626277" httpUri="https://zenodo.org/record/10626277/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
|
||
), the lacrimal is visible on the anterior margin of the antorbital process. In ventral view, the triangular lacrimal occupies about 50% of the surface of the orbit region (
|
||
<figureCitation id="B6E6A1BCFFFEFFB3FD60F9D8EF86F980" box="[689,756,1601,1626]" captionStart="Fig" captionStartId="6.[151,184,505,525]" captionTargetBox="[162,1521,162,482]" captionTargetPageId="6" captionText="Fig. 6. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Right ventrolateral view of the palate and orbital regions. Photo (A) and corresponding line drawing (B). Dotted lines indicate poorly visible sutures. Dark grey surface indicates fracture surfaces." figureDoi="http://doi.org/10.5281/zenodo.10626279" httpUri="https://zenodo.org/record/10626279/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
|
||
). The suture with the jugal is not visible. Only the base of the styliform portion of the jugal is preserved, located medial to the antorbital notch and displaying a concave lateral surface.
|
||
<emphasis id="1CA9612BFFFEFFB3FFB3F955EC11F93C" box="[98,355,1740,1766]" italics="true" pageId="5" pageNumber="6">Palatine and pterygoid</emphasis>
|
||
.—The maxilla-palatine suture is difficult to follow, only discernable in its medial portion where the suture is directed anterolaterally (
|
||
<figureCitation id="B6E6A1BCFFFEFFB3FDC4F897EF2FF8FD" box="[533,605,1806,1831]" captionStart="Fig" captionStartId="6.[151,184,505,525]" captionTargetBox="[162,1521,162,482]" captionTargetPageId="6" captionText="Fig. 6. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Right ventrolateral view of the palate and orbital regions. Photo (A) and corresponding line drawing (B). Dotted lines indicate poorly visible sutures. Dark grey surface indicates fracture surfaces." figureDoi="http://doi.org/10.5281/zenodo.10626279" httpUri="https://zenodo.org/record/10626279/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
|
||
). The anterior tips of right and left palatines are widely separated (
|
||
<quantity id="E92510DCFFFEFFB3FD78F8B7EF88F89D" box="[681,762,1838,1863]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="6" unit="mm" value="54.0">54 mm</quantity>
|
||
) as in most other extant delphinoids. The robust ridge of the palatine directed posteromedioventrally and marking the limit between the palate and the orbit/pterygoid sinus region is keeled and more prominent than in extant phocoenids, although it is not as sharp as in
|
||
<taxonomicName id="E9DDC6BAFFFEFFB3FB52FBA8E844FB91" authorityName="Lambert" authorityYear="2008" box="[1155,1334,1073,1099]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFEFFB3FB52FBA8E844FB91" box="[1155,1334,1073,1099]" italics="true" pageId="5" pageNumber="6">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
. In the latter, this ridge has been proposed to correspond to the anterior limit of a lobe of the pterygoid
|
||
<taxonomicName id="E9DDC6BAFFFEFFB3FB5EFBEAE8F7FB56" authority="(Lambert 2008)" baseAuthorityName="Lambert" baseAuthorityYear="2008" box="[1167,1413,1139,1164]" class="Mammalia" family="Phocoenidae" genus="Phocoena" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="sinus">
|
||
sinus (
|
||
<bibRefCitation id="4A4CC0C8FFFEFFB3FB09FBEAE80FFB56" author="Lambert, O." box="[1240,1405,1139,1164]" pageId="5" pageNumber="6" pagination="863 - 872" refId="ref9029" refString="Lambert, O. 2008. A new porpoise (Cetacea, Odontoceti, Phocoenidae) from the Pliocene of the North Sea. Journal of Vertebrate Paleontology 28: 863 - 872." type="journal article" year="2008">Lambert 2008</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Considering the presence of this oblique ridge in extant monodontids, which are known to lack an anterior
|
||
<taxonomicName id="E9DDC6BAFFFEFFB3FAE7FB2AEE99FB37" authority="(Fraser and Purves 1960)" baseAuthorityName="Fraser and Purves" baseAuthorityYear="1960" class="Mammalia" family="Phocoenidae" genus="Phocoena" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="sinus">
|
||
sinus (
|
||
<bibRefCitation id="4A4CC0C8FFFEFFB3FA50FB2AEE96FB37" author="Fraser, F. C. & Purves, P. E." pageId="5" pageNumber="6" pagination="1 - 140" refId="ref8718" refString="Fraser, F. C. and Purves, P. E. 1960. Hearing in cetaceans: Evolution of the accessory air sacs and the structure of the outer and middle ear in recent cetaceans. Bulletin of the British Museum (Natural History), Zoology 7: 1 - 140." type="journal article" year="1960">Fraser and Purves 1960</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, alternatively it may mark the origin of part of the pterygoid muscles involved in the closure of the mandibles (for delphinids see
|
||
<bibRefCitation id="4A4CC0C8FFFEFFB3FB80FA8DE99FFAF7" author="Seagars, D. J." box="[1105,1261,1300,1325]" pageId="5" pageNumber="6" refId="ref9683" refString="Seagars, D. J. 1982. Jaw Structure and Functional Mechanics of Six Delphinids (Cetacea, Odontoceti). 179 pp. Unpublished M. Sc. thesis, San Diego State University, San Diego." type="book" year="1982">Seagars 1982</bibRefCitation>
|
||
). Only a small part of the lateral lamina of the palatine is preserved. Although both pterygoids are lost, the well-preserved medial part of the right palatine provides a rough estimate of the palatine-pterygoid suture and of the extent of the pterygoid sinus in this area. The pterygoid sinus fossa excavates the palatine up to the level of the antorbital notch, whereas the sinus is more posterior in extant phocoenids (and some extinct taxa like
|
||
<taxonomicName id="E9DDC6BAFFFEFFB3FCF9F98CEEA9F9F5" authorityName="Lambert" authorityYear="2008" box="[808,987,1557,1583]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFEFFB3FCF9F98CEEA9F9F5" box="[808,987,1557,1583]" italics="true" pageId="5" pageNumber="6">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
). A few small pockets in the dorsal plate of the palatine anterior to each choana presumably correspond to extensions of the pterygoid sinus system.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFFEFFB3FCF9F919E827F8C6" blockId="5.[808,1481,1664,1820]" pageId="5" pageNumber="6">
|
||
<emphasis id="1CA9612BFFFEFFB3FCF9F919EEF6F940" box="[808,900,1664,1690]" italics="true" pageId="5" pageNumber="6">Parietal</emphasis>
|
||
.—Only a short portion of the thin parietal is preserved on the mediodorsal wall of each temporal fossa. The wider dorsal exposure of the left parietal in this area (
|
||
<figureCitation id="B6E6A1BCFFFEFFB3FAACF95AE8B3F906" box="[1405,1473,1731,1756]" captionStart="Fig" captionStartId="2.[153,186,680,700]" captionTargetBox="[154,1530,165,658]" captionTargetId="graphics-729@2.[770,1528,176,649]" captionTargetPageId="2" captionText="Fig. 2. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype) in dorsal view. Photo (A) and corresponding line drawing (B). Dotted lines indicate reconstructed parts and poorly visible structures. Dark grey surfaces indicate fracture surfaces. Light grey surface indicates the original position of the lost right nasal." figureDoi="http://doi.org/10.5281/zenodo.10626271" httpUri="https://zenodo.org/record/10626271/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
|
||
) is almost certainly an artefact introduced during reconstruction of the skull and/or post-mortem deformation.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFFEFFB0FCF9F8B5EE41FB7C" blockId="5.[808,1481,1836,1960]" lastBlockId="6.[151,825,612,1190]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
<emphasis id="1CA9612BFFFEFFB3FCF9F8B5EED0F89C" box="[808,930,1836,1862]" italics="true" pageId="5" pageNumber="6">Squamosal</emphasis>
|
||
.—The zygomatic process is proportionally longer than in extant phocoenids and the extinct
|
||
<taxonomicName id="E9DDC6BAFFFEFFB3FAD8F8D4E8B0F8BD" authorityName="Barnes" authorityYear="1985" box="[1289,1474,1869,1895]" class="Mammalia" family="Phocoenidae" genus="Salumiphocaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="5" pageNumber="6" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFEFFB3FAD8F8D4E8B0F8BD" box="[1289,1474,1869,1895]" italics="true" pageId="5" pageNumber="6">Salumiphocaena</emphasis>
|
||
</taxonomicName>
|
||
. In lateral view, this process has a rectangular outline (
|
||
<figureCitation id="B6E6A1BCFFFEFFB3FAA9F8F6E8C9F852" box="[1400,1467,1903,1928]" captionStart="Fig" captionStartId="5.[98,131,929,949]" captionTargetBox="[98,1477,158,906]" captionTargetId="figure-619@5.[97,1481,157,580]" captionTargetPageId="5" captionText="Fig. 5. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Right lateral view, photo (A1) and corresponding line drawing (A2). Dotted lines indicate the dorsal margin of the temporal fossa (A1), the reconstructed right postorbital process, and poorly visible sutures (A2). B. Left lateral view." figureDoi="http://doi.org/10.5281/zenodo.10626277" httpUri="https://zenodo.org/record/10626277/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
|
||
). The fossae for the sternomastoideus muscles, deeper than in extant phocoenids, occupy more than two thirds of the lateral surface of the process. The postglenoid process is a short and rounded protuberance, more prominent than in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FD0EFD3DED8AFD04" authorityName="Lambert" authorityYear="2008" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFDFFB0FD0EFD3DED8AFD04" italics="true" pageId="6" pageNumber="7">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
. The distance between the postglenoid process and the spiny process is longer in the latter, corresponding to a posterior portion of the tympanosquamosal recess (for the middle sinus see
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FE88FCBEEF16FC9A" author="Fraser, F. C. & Purves, P. E." box="[345,612,807,832]" pageId="6" pageNumber="7" pagination="1 - 140" refId="ref8718" refString="Fraser, F. C. and Purves, P. E. 1960. Hearing in cetaceans: Evolution of the accessory air sacs and the structure of the outer and middle ear in recent cetaceans. Bulletin of the British Museum (Natural History), Zoology 7: 1 - 140." type="journal article" year="1960">Fraser and Purves 1960</bibRefCitation>
|
||
) that is larger than in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FF64FCDFECCDFCBA" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[181,447,838,864]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gigaseorum">
|
||
<emphasis id="1CA9612BFFFDFFB0FF64FCDFECCDFCBA" box="[181,447,838,864]" italics="true" pageId="6" pageNumber="7">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
. The mandibular fossa is proportionally narrower, less rounded than in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FD81FCFEEE73FC5B" authorityName="Lambert" authorityYear="2008" box="[592,769,871,897]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFDFFB0FD81FCFEEE73FC5B" box="[592,769,871,897]" italics="true" pageId="6" pageNumber="7">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
, and medially margined by a wider and longer anterior part of the tympanosquamosal recess, on a broad medial flange along the zygomatic process (
|
||
<figureCitation id="B6E6A1BCFFFDFFB0FEA3FC53ECA0FC39" box="[370,466,970,995]" captionStart="Fig" captionStartId="4.[862,894,973,993]" captionTargetBox="[868,1529,164,947]" captionTargetId="figure-663@4.[867,1528,166,911]" captionTargetPageId="4" captionText="Fig. 3. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Anterior view of the facial region. B. Posterior view. C. Ventral view of the right squamosal." figureDoi="http://doi.org/10.5281/zenodo.10626273" httpUri="https://zenodo.org/record/10626273/files/figure.png" pageId="6" pageNumber="7">Figs. 3C</figureCitation>
|
||
,
|
||
<figureCitation id="B6E6A1BCFFFDFFB0FE0CFC53EC9FFC39" box="[477,493,970,995]" captionStart="Fig" captionStartId="6.[862,895,1824,1844]" captionTargetBox="[920,1476,1335,1799]" captionTargetId="figure-810@6.[1016,1310,1335,1765]" captionTargetPageId="6" captionText="Fig. 7. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Fragmentary basicranium in ventral view. Dotted lines indicate poorly visible sutures, the right tympanosquamosal recess, the groove for the mandibular nerve V3, and the presumed area of origin for scalenus medius." figureDoi="http://doi.org/10.5281/zenodo.10626283" httpUri="https://zenodo.org/record/10626283/files/figure.png" pageId="6" pageNumber="7">7</figureCitation>
|
||
). Between the zygomatic process and the squamosal plate, a deep and narrow squamosal fossa forms the floor of the temporal fossa. In extant phocoenids, the shallower squamosal fossa is more open laterally, and not, or only slightly concave longitudinally. Present in the
|
||
<typeStatus id="F166039BFFFDFFB0FF11FBF4EC53FB5C" box="[192,289,1133,1158]" pageId="6" pageNumber="7">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FE94FBF5EF0AFB5F" authorityName="Lambert" authorityYear="2008" box="[325,632,1131,1158]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="bosselaersi">
|
||
<emphasis id="1CA9612BFFFDFFB0FE94FBF5EF0AFB5F" box="[325,632,1131,1158]" italics="true" pageId="6" pageNumber="7">Septemtriocetus bosselaersi</emphasis>
|
||
</taxonomicName>
|
||
, the fossa is nevertheless posteriorly shorter than in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FDF5FB15EE5EFB7C" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[548,812,1164,1190]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gigaseorum">
|
||
<emphasis id="1CA9612BFFFDFFB0FDF5FB15EE5EFB7C" box="[548,812,1164,1190]" italics="true" pageId="6" pageNumber="7">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="7AA2EDB1FFFDFFB0FF46FE60ECA0FD9F" ID-DOI="http://doi.org/10.5281/zenodo.10626279" ID-Zenodo-Dep="10626279" httpUri="https://zenodo.org/record/10626279/files/figure.png" pageId="6" pageNumber="7" startId="6.[151,184,505,525]" targetBox="[162,1521,162,482]" targetPageId="6" targetType="figure">
|
||
<paragraph id="2E62BD39FFFDFFB0FF46FE60ECA0FD9F" blockId="6.[151,1534,504,581]" pageId="6" pageNumber="7">
|
||
Fig. 6. Skull of the phocoenid cetacean
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FE2DFE61EFA6FDD7" authority="Colpaert & Bosselaers & Lambert, 2015" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[508,724,504,525]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gigaseorum" status="gen. et sp. nov.">
|
||
<emphasis id="1CA9612BFFFDFFB0FE2DFE61EFA6FDD7" box="[508,724,504,525]" italics="true" pageId="6" pageNumber="7">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="079ADC50FFFDFFB0FD08FE60EE13FDD7" box="[729,865,505,525]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Right ventrolateral view of the palate and orbital regions. Photo (
|
||
<emphasis id="1CA9612BFFFDFFB0FD7AFD8DEFC9FDF3" bold="true" box="[683,699,532,553]" pageId="6" pageNumber="7">A</emphasis>
|
||
) and corresponding line drawing (
|
||
<emphasis id="1CA9612BFFFDFFB0FC25FD8DE971FDF3" bold="true" box="[1012,1027,532,553]" pageId="6" pageNumber="7">B</emphasis>
|
||
). Dotted lines indicate poorly visible sutures. Dark grey surface indicates fracture surfaces.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2E62BD39FFFDFFB0FF46FB2EEE44F921" blockId="6.[151,825,1207,1787]" pageId="6" pageNumber="7">
|
||
<emphasis id="1CA9612BFFFDFFB0FF46FB2EEC65FB0B" box="[151,279,1207,1233]" italics="true" pageId="6" pageNumber="7">Exoccipital</emphasis>
|
||
.—The occipital condyles protrude only weakly from the posterior wall of the braincase and a shallow dorsal condyloid fossa is present (
|
||
<figureCitation id="B6E6A1BCFFFDFFB0FE14FB60EF6EFAC8" box="[453,540,1273,1298]" captionStart="Fig" captionStartId="4.[862,894,973,993]" captionTargetBox="[868,1529,164,947]" captionTargetId="figure-663@4.[867,1528,166,911]" captionTargetPageId="4" captionText="Fig. 3. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). A. Anterior view of the facial region. B. Posterior view. C. Ventral view of the right squamosal." figureDoi="http://doi.org/10.5281/zenodo.10626273" httpUri="https://zenodo.org/record/10626273/files/figure.png" pageId="6" pageNumber="7">Fig. 3B</figureCitation>
|
||
). The long axis of the left condyle diverges more from the sagittal plane than on the right side (difference of ca. 5°). In addition, the left exoccipital is wider than the right exoccipital along its ventral margin. Not due to reconstruction approximations, this asymmetry may indicate either an asymmetrical position, or even preferential directions of movement, for the head relative to the vertebral column (a hypothesis that should be tested with a larger sample, including extant phocoenids). The foramen magnum is subcircular. As in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FE35F984EFE5F9ED" authorityName="Lambert" authorityYear="2008" box="[484,663,1565,1591]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFDFFB0FE35F984EFE5F9ED" box="[484,663,1565,1591]" italics="true" pageId="6" pageNumber="7">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
and most other fossil phocoenids, a shallow depression is present on the anterior surface of the paroccipital process, probably corresponding to the fossa for the posterior sinus. The fossa is deeper in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FED6F906EDABF900" authority="(Ichishima and Kimura 2009)" baseAuthorityName="Ichishima and Kimura" baseAuthorityYear="2009" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="minutus">
|
||
<emphasis id="1CA9612BFFFDFFB0FED6F906EF59F963" box="[263,555,1695,1721]" italics="true" pageId="6" pageNumber="7">Haborophocoena minutus</emphasis>
|
||
(
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FDEBF939EDA3F900" author="Ichishima, H. & Kimura, M." pageId="6" pageNumber="7" pagination="855 - 874" refId="ref8993" refString="Ichishima, H. and Kimura, M. 2009. A new species of Haborophocoena, an Early Pliocene phocoenid cetacean from Hokkaido, Japan. Marine Mammal Science 25: 855 - 874." type="journal article" year="2009">Ichishima and Kimura 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and extant phocoenids, in which it is sometimes only closed posteriorly by a thin, translucent plate of exoccipital.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFFDFFB0FF46F892E822FD07" blockId="6.[151,824,1803,1960]" lastBlockId="6.[862,1535,612,733]" pageId="6" pageNumber="7">
|
||
<emphasis id="1CA9612BFFFDFFB0FF46F892EC59F8FF" box="[151,299,1803,1829]" italics="true" pageId="6" pageNumber="7">Basioccipital</emphasis>
|
||
.—The left basioccipital crest is preserved. In lateral view, a shallow notch cuts the crest before a transversely swollen region anterior to the jugular notch. As in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FF46F8F4EC38F85D" authorityName="Lambert" authorityYear="2008" box="[151,330,1901,1927]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFDFFB0FF46F8F4EC38F85D" box="[151,330,1901,1927]" italics="true" pageId="6" pageNumber="7">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
, a subcircular depression is present on the medial wall of the basioccipital crest, next to the jugular notch (
|
||
<figureCitation id="B6E6A1BCFFFDFFB0FC60FDFDEE88FDA7" box="[945,1018,612,637]" captionStart="Fig" captionStartId="6.[862,895,1824,1844]" captionTargetBox="[920,1476,1335,1799]" captionTargetId="figure-810@6.[1016,1310,1335,1765]" captionTargetPageId="6" captionText="Fig. 7. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Fragmentary basicranium in ventral view. Dotted lines indicate poorly visible sutures, the right tympanosquamosal recess, the groove for the mandibular nerve V3, and the presumed area of origin for scalenus medius." figureDoi="http://doi.org/10.5281/zenodo.10626283" httpUri="https://zenodo.org/record/10626283/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
|
||
), probably for the scalenus medius muscle, which when activated draws the head downward (
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FA4FFD1DEEE9FD67" author="Lambert, O." pageId="6" pageNumber="7" pagination="863 - 872" refId="ref9029" refString="Lambert, O. 2008. A new porpoise (Cetacea, Odontoceti, Phocoenidae) from the Pliocene of the North Sea. Journal of Vertebrate Paleontology 28: 863 - 872." type="journal article" year="2008">Lambert 2008</bibRefCitation>
|
||
). In ventral view, the angle drawn by the two basioccipital crests is roughly estimated at 45–50°.
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFFDFFB0FC8FFD74E831FB92" blockId="6.[862,1535,749,1096]" pageId="6" pageNumber="7">
|
||
<emphasis id="1CA9612BFFFDFFB0FC8FFD74EE87FCDD" box="[862,1013,749,775]" italics="true" pageId="6" pageNumber="7">Basisphenoid</emphasis>
|
||
.—The basisphenoid crest is damaged by erosion. Its lateral wall is slightly excavated at its base, but not as much as in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FC24FCB7E9DAFC92" authorityName="Lambert" authorityYear="2008" box="[1013,1192,814,840]" class="Mammalia" family="Phocoenidae" genus="Septemtriocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFDFFB0FC24FCB7E9DAFC92" box="[1013,1192,814,840]" italics="true" pageId="6" pageNumber="7">Septemtriocetus</emphasis>
|
||
</taxonomicName>
|
||
and extant phocoenids. If present, the basisphenoidal sinus (sensu
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FB3AFCD6E888FCB2" author="Mead, J. G. & Fordyce, R. E." box="[1259,1530,847,872]" pageId="6" pageNumber="7" pagination="1 - 248" refId="ref9166" refString="Mead, J. G. and Fordyce, R. E. 2009. The therian skull: a lexicon with emphasis on the odontocetes. Smithsonian Contributions to Zoology 627: 1 - 248." type="journal article" year="2009">Mead and Fordyce 2009</bibRefCitation>
|
||
; basisphenoid lobe of the pterygoid
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FB31FCF6EE86FC72" authority="sensu Ichishima and Kimura 2005" authorityName="Ichishima and Kimura" authorityYear="2005" class="Mammalia" family="Phocoenidae" genus="Phocoena" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="sinus">
|
||
sinus sensu
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FAB3FCF6EE86FC72" author="Ichishima, H. & Kimura, M." pageId="6" pageNumber="7" pagination="655 - 664" refId="ref8954" refString="Ichishima, H. and Kimura, M. 2005. Haborophocoena toyoshimai, a new early Pliocene porpoise (Cetacea: Phocoenidae) from Hokkaido, Japan. Journal of Vertebrate Paleontology 25: 655 - 664." type="journal article" year="2005">Ichishima and Kimura 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FC2CFC16E94AFC72" author="Ichishima, H. & Kimura, M." box="[1021,1080,911,936]" pageId="6" pageNumber="7" pagination="855 - 874" refId="ref8993" refString="Ichishima, H. and Kimura, M. 2009. A new species of Haborophocoena, an Early Pliocene phocoenid cetacean from Hokkaido, Japan. Marine Mammal Science 25: 855 - 874." type="journal article" year="2009">2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FB92FC16E993FC72" author="Lambert, O." box="[1091,1249,911,936]" pageId="6" pageNumber="7" pagination="863 - 872" refId="ref9029" refString="Lambert, O. 2008. A new porpoise (Cetacea, Odontoceti, Phocoenidae) from the Pliocene of the North Sea. Journal of Vertebrate Paleontology 28: 863 - 872." type="journal article" year="2008">Lambert 2008</bibRefCitation>
|
||
) was poorly developed.A deeper fossa is reported in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FB42FC34E8EEFC1D" box="[1171,1436,941,967]" class="Mammalia" family="Phocoenidae" genus="Piscolithax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="longirostris">
|
||
<emphasis id="1CA9612BFFFDFFB0FB42FC34E8EEFC1D" box="[1171,1436,941,967]" italics="true" pageId="6" pageNumber="7">Piscolithax longirostris</emphasis>
|
||
</taxonomicName>
|
||
, and the condition in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FC21FC54E98FFC3D" authorityName="Colpaert & Bosselaers & Lambert" authorityYear="2015" box="[1008,1277,973,999]" class="Mammalia" genus="Brabocetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="gigaseorum">
|
||
<emphasis id="1CA9612BFFFDFFB0FC21FC54E98FFC3D" box="[1008,1277,973,999]" italics="true" pageId="6" pageNumber="7">Brabocetus gigaseorum</emphasis>
|
||
</taxonomicName>
|
||
more closely matches the condition in
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FBC9FC74E9D0FBDD" authorityName="de Muizon" authorityYear="1988" box="[1048,1186,1005,1031]" class="Mammalia" family="Phocoenidae" genus="Australithax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">
|
||
<emphasis id="1CA9612BFFFDFFB0FBC9FC74E9D0FBDD" box="[1048,1186,1005,1031]" italics="true" pageId="6" pageNumber="7">Australithax</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FB63FC74E88AFBDD" box="[1202,1528,1005,1031]" class="Mammalia" family="Phocoenidae" genus="Haborophocoena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="toyoshimai">
|
||
<emphasis id="1CA9612BFFFDFFB0FB63FC74E88AFBDD" box="[1202,1528,1005,1031]" italics="true" pageId="6" pageNumber="7">Haborophocoena toyoshimai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="1CA9612BFFFDFFB0FC8FFB97EEACFBFD" box="[862,990,1038,1063]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FC8FFB97EEA8FBFD" authorityName="de Muizon" authorityYear="1986" box="[862,986,1038,1063]" class="Mammalia" family="Phocoenidae" genus="Lomacetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="genus">Lomacetus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="E9DDC6BAFFFDFFB0FBC9FB94E985FBFD" box="[1048,1271,1037,1063]" class="Mammalia" family="Phocoenidae" genus="Piscolithax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tedfordi">
|
||
<emphasis id="1CA9612BFFFDFFB0FBC9FB94E985FBFD" box="[1048,1271,1037,1063]" italics="true" pageId="6" pageNumber="7">Piscolithax tedfordi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FAD6FB96E8E8FBF2" author="Barnes, L. G." box="[1287,1434,1039,1064]" pageId="6" pageNumber="7" pagination="1 - 46" refId="ref8032" refString="Barnes, L. G. 1984. Fossil odontocetes (Mammalia: Cetacea) from the Almejas Formation, Isla Cedros, Mexico. PaleoBios 42: 1 - 46." type="journal article" year="1984">Barnes 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FA76FB96EEEBFB92" author="Muizon, C. de" pageId="6" pageNumber="7" pagination="1 - 188" refId="ref9203" refString="Muizon, C. de 1984. Les Vertebres de la Formation Pisco (Perou). Deuxieme partie: Les Odontocetes (Cetacea, Mammalia) du Pliocene inferieur du Sud-Sacaco. Travaux de l'Institut Francais d'Etudes Andines 27: 1 - 188." type="journal article" year="1984">Muizon 1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FC75FBB6EE9FFB92" author="Muizon, C. de" box="[932,1005,1071,1096]" pageId="6" pageNumber="7" pagination="1 - 244" refId="ref9271" refString="Muizon, C. de 1988 b. Les Vertebres fossiles de la Formation Pisco (Perou). Troisieme partie: Les Odontocetes (Cetacea, Mammalia) du Miocene. Travaux de l'Institut Francais d'Etudes Andines 42: 1 - 244." type="journal article" year="1988">1988b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4A4CC0C8FFFDFFB0FC28FBB6E845FB92" author="Ichishima, H. & Kimura, M." box="[1017,1335,1071,1096]" pageId="6" pageNumber="7" pagination="655 - 664" refId="ref8954" refString="Ichishima, H. and Kimura, M. 2005. Haborophocoena toyoshimai, a new early Pliocene porpoise (Cetacea: Phocoenidae) from Hokkaido, Japan. Journal of Vertebrate Paleontology 25: 655 - 664." type="journal article" year="2005">Ichishima and Kimura 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2E62BD39FFFDFFB0FC8FFBC1E842FAC9" blockId="6.[862,1535,1112,1299]" pageId="6" pageNumber="7">
|
||
<emphasis id="1CA9612BFFFDFFB0FC8FFBC1EE96FBA8" box="[862,996,1112,1138]" italics="true" pageId="6" pageNumber="7">Alisphenoid</emphasis>
|
||
.—Better preserved on the left side of IRSNB M.2171, the foramen ovale is small (maximum diameter
|
||
<quantity id="E92510DCFFFDFFB0FA20FBE3EEFCFB69" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="7" unit="mm" value="7.0">7 mm</quantity>
|
||
). It is bordered laterally by a short trough that held the mandibular nerve V3 (
|
||
<figureCitation id="B6E6A1BCFFFDFFB0FB87FB23E9EAFB09" box="[1110,1176,1210,1235]" captionStart="Fig" captionStartId="6.[862,895,1824,1844]" captionTargetBox="[920,1476,1335,1799]" captionTargetId="figure-810@6.[1016,1310,1335,1765]" captionTargetPageId="6" captionText="Fig. 7. Skull of the phocoenid cetacean Brabocetus gigaseorum gen. et sp. nov. from the Pliocene of northern Belgium IRSNB M. 2171 (holotype). Fragmentary basicranium in ventral view. Dotted lines indicate poorly visible sutures, the right tympanosquamosal recess, the groove for the mandibular nerve V3, and the presumed area of origin for scalenus medius." figureDoi="http://doi.org/10.5281/zenodo.10626283" httpUri="https://zenodo.org/record/10626283/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
|
||
). The anterior ridge of the aforementioned trough corresponds to the posterior limit of a wide and shallow fossa for the pterygoid sinus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |