263 lines
28 KiB
XML
263 lines
28 KiB
XML
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<mods:titleInfo id="861996359A6D04ABA371560515F21B4C">
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<mods:title id="1ED01ABF67EFF0525327F05357CB0E98">Stone loaches of Choman River system, Kurdistan, Iran (Teleostei: Cypriniformes: Nemacheilidae)</mods:title>
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<mods:namePart id="04BE77B328D6B2364B5F099AFC071391">Kamangar, Barzan Bahrami</mods:namePart>
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<mods:namePart id="8E72B7C18743673CEF6E5261C75477A2">Prokofiev, Artem M.</mods:namePart>
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<mods:namePart id="D5E54CFC4BC23B24D17A5CFB0AAEB5A4">Ghaderi, Edris</mods:namePart>
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<mods:namePart id="ADA994408010004892D99B8AAFF5EB93">Nalbant, Theodore T.</mods:namePart>
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<mods:title id="05C4A202508165C07C8BE7FC87FB10D0">Zootaxa</mods:title>
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<mods:date id="9DA9C46962D2D96C0E11720A31D0E8AB">2014</mods:date>
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<mods:number id="C58CE026EA6BE681DCB1ECF6658D755C">3755</mods:number>
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<treatment id="33348786FF97FFFCFF20FD89FE6E7B9A" ID-DOI="http://doi.org/10.5281/zenodo.5672508" ID-GBIF-Taxon="127666877" ID-Zenodo-Dep="5672508" LSID="urn:lsid:plazi:treatment:33348786FF97FFFCFF20FD89FE6E7B9A" httpUri="http://treatment.plazi.org/id/33348786FF97FFFCFF20FD89FE6E7B9A" lastPageId="5" lastPageNumber="37" pageId="4" pageNumber="37">
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<subSubSection id="F387651BFF97FFFDFF20FD89FD537971" box="[151,694,549,574]" pageId="4" pageNumber="37" type="nomenclature">
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<paragraph id="BB223690FF97FFFDFF20FD89FD537971" blockId="4.[151,694,549,574]" box="[151,694,549,574]" pageId="4" pageNumber="37">
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<heading id="E06A81FCFF97FFFDFF20FD89FD537971" bold="true" box="[151,694,549,574]" fontSize="10" level="2" pageId="4" pageNumber="37" reason="2">
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<taxonomicName id="7C9D4D13FF97FFFDFF20FD89FD537971" ID-CoL="6BDV" ID-ENA="417298" authority="Banarescu & Nalbant, 1966" authorityName="Banarescu & Nalbant" authorityYear="1966" box="[151,694,549,574]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
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<emphasis id="89E9EA82FF97FFFDFF20FD89FD537971" bold="true" box="[151,694,549,574]" pageId="4" pageNumber="37">
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<emphasis id="89E9EA82FF97FFFDFF20FD89FE837971" bold="true" box="[151,358,549,574]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
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<bibRefCitation id="DF0C4B61FF97FFFDFEDBFD89FD537971" author="Banarescu" box="[364,694,549,574]" pageId="4" pageNumber="37" refString="Banarescu, P. & Nalbant, T. T. (1966) The 3 rd Danish Expedition to central Asia. Zoological results 34. Cobitidae (Pisces) from Afghanistan and Iran. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kjobenhavn, 129, 149 - 186, pls. 19 - 21." type="journal article" year="1966">Bănărescu & Nalbant, 1966</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F387651BFF97FFFDFF20FDC1FC9A78CD" pageId="4" pageNumber="37" type="materials_examined">
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||
<paragraph id="BB223690FF97FFFDFF20FDC1FEAF79E5" blockId="4.[151,1437,621,2014]" pageId="4" pageNumber="37">
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<emphasis id="89E9EA82FF97FFFDFF20FDC1FEDF79C9" bold="true" box="[151,314,621,646]" pageId="4" pageNumber="37">
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<typeStatus id="64268832FF97FFFDFF20FDC1FF3779C9" box="[151,210,621,646]" pageId="4" pageNumber="37">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="7C9D4D13FF97FFFDFEF1FDC1FE0779CA" box="[326,482,621,646]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="persa">
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<emphasis id="89E9EA82FF97FFFDFEF1FDC1FE0779CA" box="[326,482,621,646]" italics="true" pageId="4" pageNumber="37">Cobitis persa</emphasis>
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</taxonomicName>
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(nec Heckel, 1848)
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<bibRefCitation id="DF0C4B61FF97FFFDFD6BFDC2FBC679C9" author="Banarescu" box="[732,1059,621,646]" pageId="4" pageNumber="37" refString="Banarescu, P. & Nalbant, T. T. (1966) The 3 rd Danish Expedition to central Asia. Zoological results 34. Cobitidae (Pisces) from Afghanistan and Iran. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kjobenhavn, 129, 149 - 186, pls. 19 - 21." type="journal article" year="1966">Bănărescu & Nalbant, 1966</bibRefCitation>
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=
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<taxonomicName id="7C9D4D13FF97FFFDFBFBFDC1FEA079E5" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="brandti">
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<emphasis id="89E9EA82FF97FFFDFBFBFDC1FAF679C9" box="[1100,1299,621,646]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
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cf.
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<emphasis id="89E9EA82FF97FFFDFAFEFDC1FA7979C9" box="[1353,1436,621,646]" italics="true" pageId="4" pageNumber="37">brandti</emphasis>
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(Kessler, 1877)
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="BB223690FF97FFFDFF70FD19FC9A78CD" blockId="4.[151,1437,621,2014]" pageId="4" pageNumber="37">
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<emphasis id="89E9EA82FF97FFFDFF70FD19FE5A7981" bold="true" box="[199,447,693,718]" pageId="4" pageNumber="37">Emended diagnosis:</emphasis>
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This genus can be distinguished from all other nemacheilid genera based on the following combination of characters: mouth unmodified, with lips furrowed to nearly smooth, processus dentiformis present or absent; preethmoidei-I absent; four cylindrical pectoral bony radials; mature males with epidermal tubercles widely distributed on head, body and paired fins, not forming separate pads; anus close to analfin origin; dorsal adipose crest present or absent, when present, of uniform height or gradually increasing in height towards base of caudal-fin; no jet-black mark at base of dorsal fin.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="F387651BFF97FFFCFF70FC21FE6E7B9A" lastPageId="5" lastPageNumber="38" pageId="4" pageNumber="37" type="discussion">
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<paragraph id="BB223690FF97FFFDFF70FC21FDDC7E36" blockId="4.[151,1437,621,2014]" pageId="4" pageNumber="37">
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<emphasis id="89E9EA82FF97FFFDFF70FC21FED878E9" bold="true" box="[199,317,909,934]" pageId="4" pageNumber="37">Remarks:</emphasis>
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<bibRefCitation id="DF0C4B61FF97FFFDFEF2FC21FDE378E9" author="Prokofiev" box="[325,518,909,934]" pageId="4" pageNumber="37" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49 (10), 874 - 898. http: // dx. doi. org / 10.1134 / s 0032945209100051" type="journal article" year="2009">Prokofiev (2009)</bibRefCitation>
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transferred species having the free pelvic axillary lobe from
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<taxonomicName id="7C9D4D13FF97FFFDFB00FC21FA9B78E9" box="[1207,1406,909,934]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
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<emphasis id="89E9EA82FF97FFFDFB00FC21FA9B78E9" box="[1207,1406,909,934]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
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</taxonomicName>
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to
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<taxonomicName id="7C9D4D13FF97FFFDFF20FC1DFE227885" authority="Bleeker, 1863" authorityName="Bleeker" authorityYear="1863" box="[151,455,945,970]" class="Actinopterygii" family="Nemacheilidae" genus="Paracobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
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<emphasis id="89E9EA82FF97FFFDFF20FC1DFEFF7885" box="[151,282,945,970]" italics="true" pageId="4" pageNumber="37">Paracobitis</emphasis>
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Bleeker, 1863
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</taxonomicName>
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, which was not supported by Freyhof
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<emphasis id="89E9EA82FF97FFFDFC27FC1EFC297885" box="[912,972,945,970]" italics="true" pageId="4" pageNumber="37">et al.</emphasis>
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(2011). Furthermore, in addition to the species listed by
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<bibRefCitation id="DF0C4B61FF97FFFDFEEAFC79FDFA78A1" author="Prokofiev" box="[349,543,981,1006]" pageId="4" pageNumber="37" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49 (10), 874 - 898. http: // dx. doi. org / 10.1134 / s 0032945209100051" type="journal article" year="2009">Prokofiev (2009)</bibRefCitation>
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Freyhof
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<emphasis id="89E9EA82FF97FFFDFD3DFC7AFD2078A1" box="[650,709,981,1006]" italics="true" pageId="4" pageNumber="37">et al.</emphasis>
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(2011: 303) reported the presence of the “small pelvic axillary lobe” in
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<taxonomicName id="7C9D4D13FF97FFFDFF4BFC56FEBA7F5E" box="[252,351,1018,1041]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="persa">
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<emphasis id="89E9EA82FF97FFFDFF4BFC56FEBA7F5E" box="[252,351,1018,1041]" italics="true" pageId="4" pageNumber="37">
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<collectingCountry id="C38A7600FF97FFFDFF4BFC56FEF67F5E" box="[252,275,1018,1041]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
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. persa
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</emphasis>
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</taxonomicName>
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, the
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<typeStatus id="64268832FF97FFFDFE2BFC56FE297F5D" box="[412,460,1018,1042]" pageId="4" pageNumber="37">type</typeStatus>
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species of the genus. It should be noted, however, that
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<bibRefCitation id="DF0C4B61FF97FFFDFBE4FC56FEE77F79" author="Banarescu" pageId="4" pageNumber="37" refString="Banarescu, P. & Nalbant, T. T. (1966) The 3 rd Danish Expedition to central Asia. Zoological results 34. Cobitidae (Pisces) from Afghanistan and Iran. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kjobenhavn, 129, 149 - 186, pls. 19 - 21." type="journal article" year="1966">Bănărescu & Nalbant (1966: 153–154)</bibRefCitation>
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misidentified
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<taxonomicName id="7C9D4D13FF97FFFDFE10FBB2FDEF7F7A" box="[423,522,1054,1077]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="persa">
|
||
<emphasis id="89E9EA82FF97FFFDFE10FBB2FDEF7F7A" box="[423,522,1054,1077]" italics="true" pageId="4" pageNumber="37">
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||
<collectingCountry id="C38A7600FF97FFFDFE10FBB2FE5B7F7A" box="[423,446,1054,1077]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
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. persa
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</emphasis>
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</taxonomicName>
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with a species distributed in the Urmia Lake basin when they erected the taxon
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<taxonomicName id="7C9D4D13FF97FFFDFF20FBEDFEBB7F15" box="[151,350,1089,1114]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
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||
<emphasis id="89E9EA82FF97FFFDFF20FBEDFEBB7F15" box="[151,350,1089,1114]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
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||
</taxonomicName>
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||
. Later, the true
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||
<taxonomicName id="7C9D4D13FF97FFFDFD96FBEEFD637F16" box="[545,646,1090,1113]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="persa">
|
||
<emphasis id="89E9EA82FF97FFFDFD96FBEEFD637F16" box="[545,646,1090,1113]" italics="true" pageId="4" pageNumber="37">
|
||
<collectingCountry id="C38A7600FF97FFFDFD96FBEEFDDD7F16" box="[545,568,1090,1113]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
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. persa
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</emphasis>
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</taxonomicName>
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was described by
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<bibRefCitation id="DF0C4B61FF97FFFDFCD0FBEDFB747F15" author="Nalbant" box="[871,1169,1089,1114]" pageId="4" pageNumber="37" refString="Nalbant, T. T. & Bianco, P. G. (1998) The loaches of Iran and adjacent regions with description of six new species (Cobitoidea). Italian Journal of Zoology, 65, Supplement 1, 109 - 123. http: // dx. doi. org / 10.1080 / 11250009809386803" type="journal article" year="1998">Nalbant & Bianco (1998)</bibRefCitation>
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||
under the name of
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<taxonomicName id="7C9D4D13FF97FFFDFA36FBEEFF157F31" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="farsicus">
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||
<emphasis id="89E9EA82FF97FFFDFA36FBEEFF157F31" italics="true" pageId="4" pageNumber="37">
|
||
<collectingCountry id="C38A7600FF97FFFDFA36FBEEFA7D7F16" box="[1409,1432,1090,1113]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
|
||
. farsicus
|
||
</emphasis>
|
||
</taxonomicName>
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||
, as clearly shown by Freyhof
|
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<emphasis id="89E9EA82FF97FFFDFDF9FBCAFD6C7F31" box="[590,649,1125,1150]" italics="true" pageId="4" pageNumber="37">et al.</emphasis>
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||
(2011: 309). Thus, the taxon
|
||
<taxonomicName id="7C9D4D13FF97FFFDFC56FBC9FB4D7F31" box="[993,1192,1125,1150]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFC56FBC9FB4D7F31" box="[993,1192,1125,1150]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
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||
</taxonomicName>
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||
was based not on
|
||
<taxonomicName id="7C9D4D13FF97FFFDFA36FBCAFF307FEE" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="persa">
|
||
<emphasis id="89E9EA82FF97FFFDFA36FBCAFF307FEE" italics="true" pageId="4" pageNumber="37">
|
||
<collectingCountry id="C38A7600FF97FFFDFA36FBCAFA7D7F32" box="[1409,1432,1126,1149]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
|
||
. persa
|
||
</emphasis>
|
||
</taxonomicName>
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||
, but on a species similar or identical with
|
||
<taxonomicName id="7C9D4D13FF97FFFDFD52FB26FBFB7FED" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" box="[741,1054,1161,1186]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="brandti">
|
||
<emphasis id="89E9EA82FF97FFFDFD52FB26FCBA7FED" box="[741,863,1161,1186]" italics="true" pageId="4" pageNumber="37">
|
||
<collectingCountry id="C38A7600FF97FFFDFD52FB26FD197FEE" box="[741,764,1162,1185]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
|
||
. brandti
|
||
</emphasis>
|
||
(Kessler, 1877)
|
||
</taxonomicName>
|
||
according to the morphological description and Figure of the Urmian loaches in
|
||
<bibRefCitation id="DF0C4B61FF97FFFDFD75FB02FBE87F89" author="Banarescu" box="[706,1037,1197,1222]" pageId="4" pageNumber="37" refString="Banarescu, P. & Nalbant, T. T. (1966) The 3 rd Danish Expedition to central Asia. Zoological results 34. Cobitidae (Pisces) from Afghanistan and Iran. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kjobenhavn, 129, 149 - 186, pls. 19 - 21." type="journal article" year="1966">Bănărescu & Nalbant (1966)</bibRefCitation>
|
||
. Our personal data suppose a quite complex taxonomical situation in
|
||
<emphasis id="89E9EA82FF97FFFDFD9FFB7EFCF47FA5" box="[552,785,1233,1258]" italics="true" pageId="4" pageNumber="37">
|
||
<taxonomicName id="7C9D4D13FF97FFFDFD9FFB7EFD7C7FA5" box="[552,665,1233,1258]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="brandti">
|
||
<collectingCountry id="C38A7600FF97FFFDFD9FFB7EFDD97FA6" box="[552,572,1234,1257]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
|
||
brandti
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="7C9D4D13FF97FFFDFD11FB7DFCF47FA5" box="[678,785,1233,1258]" family="Nemacheilidae" genus="Nemachilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="bergianus">bergianus</taxonomicName>
|
||
</emphasis>
|
||
–complex which may represent more than two species; however, the results of the molecular analysis (I-Shiung Chen, B.A. Levin, personal communication 2012) does not correspond well with the morphological data; thus, it is not possible to separate reasonably the species within this group at current state of knowledge.
|
||
<collectingCountry id="C38A7600FF97FFFDFD81FA92FDB07E19" box="[566,597,1342,1366]" name="American Samoa" pageId="4" pageNumber="37">As</collectingCountry>
|
||
a result, the taxonomic position of the fishes from the Urmia Lake cannot be satisfactory determined at present.
|
||
</paragraph>
|
||
<paragraph id="BB223690FF97FFFDFF70FA2AFECA7C49" blockId="4.[151,1437,621,2014]" pageId="4" pageNumber="37">
|
||
In the present paper we accept the limits of the genera
|
||
<taxonomicName id="7C9D4D13FF97FFFDFC9CFA29FC177ED1" box="[811,1010,1413,1438]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFC9CFA29FC177ED1" box="[811,1010,1413,1438]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7C9D4D13FF97FFFDFB9EFA29FB497ED1" box="[1065,1196,1413,1438]" class="Actinopterygii" family="Nemacheilidae" genus="Paracobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFB9EFA29FB497ED1" box="[1065,1196,1413,1438]" italics="true" pageId="4" pageNumber="37">Paracobitis</emphasis>
|
||
</taxonomicName>
|
||
proposed by Freyhof
|
||
<emphasis id="89E9EA82FF97FFFDFF20FA06FF2A7E8D" box="[151,207,1449,1474]" italics="true" pageId="4" pageNumber="37">et al.</emphasis>
|
||
(2011) due to the following reasons: (1) though we had not found other species having the free pelvic axillary lobe, we found a skin fold lacking the free posterior tip in the several specimens of the species described below. This fold can be homologous to the axillary lobe (reduced or incompletely developed), although further study is required; thus, this feature may have the mosaic distribution within the species of
|
||
<taxonomicName id="7C9D4D13FF97FFFDFB93F9B9FB0E7D61" box="[1060,1259,1557,1582]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFB93F9B9FB0E7D61" box="[1060,1259,1557,1582]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
. (2) We studied specimens of
|
||
<taxonomicName id="7C9D4D13FF97FFFDFE83F996FDA07D1D" authority="Heckel, 1843" authorityName="Heckel" authorityYear="1843" box="[308,581,1593,1618]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="tigris">
|
||
<emphasis id="89E9EA82FF97FFFDFE83F996FE767D1E" box="[308,403,1594,1617]" italics="true" pageId="4" pageNumber="37">
|
||
<collectingCountry id="C38A7600FF97FFFDFE83F996FEAE7D1E" box="[308,331,1594,1617]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
|
||
. tigris
|
||
</emphasis>
|
||
(Heckel, 1843)
|
||
</taxonomicName>
|
||
from Lake Kinneret, which possess the
|
||
<typeStatus id="64268832FF97FFFDFBA3F996FBA17D1D" box="[1044,1092,1594,1618]" pageId="4" pageNumber="37">type</typeStatus>
|
||
of male breeding tubercles of
|
||
<taxonomicName id="7C9D4D13FF97FFFDFF20F9F1FEBB7D39" box="[151,350,1629,1654]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFF20F9F1FEBB7D39" box="[151,350,1629,1654]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
. We also found both muscular cheeks and breeding tubercles in males of
|
||
<taxonomicName id="7C9D4D13FF97FFFDFB7FF9F1FF127DD5" class="Actinopterygii" family="Nemacheilidae" genus="Turcinoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="kosswigi">
|
||
<emphasis id="89E9EA82FF97FFFDFB7FF9F1FF127DD5" italics="true" pageId="4" pageNumber="37">Turcinoemacheilus kosswigi</emphasis>
|
||
</taxonomicName>
|
||
(see below). Thus, the differences in the sexual dimorphic features among the genera
|
||
<taxonomicName id="7C9D4D13FF97FFFDFB62F92DFA797DD5" box="[1237,1436,1665,1690]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFB62F92DFA797DD5" box="[1237,1436,1665,1690]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7C9D4D13FF97FFFDFF71F909FEAC7DF1" box="[198,329,1701,1726]" class="Actinopterygii" family="Nemacheilidae" genus="Paracobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFF71F909FEAC7DF1" box="[198,329,1701,1726]" italics="true" pageId="4" pageNumber="37">Paracobitis</emphasis>
|
||
</taxonomicName>
|
||
as proposed by
|
||
<bibRefCitation id="DF0C4B61FF97FFFDFDB7F909FD257DF1" author="Prokofiev" box="[512,704,1701,1726]" pageId="4" pageNumber="37" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49 (10), 874 - 898. http: // dx. doi. org / 10.1134 / s 0032945209100051" type="journal article" year="2009">Prokofiev (2009)</bibRefCitation>
|
||
may not be stable, and the interrelationships between the species of
|
||
<taxonomicName id="7C9D4D13FF97FFFDFF0AF965FE617DAD" box="[189,388,1737,1762]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFF0AF965FE617DAD" box="[189,388,1737,1762]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
with and without pelvic axillary lobe and the species of
|
||
<taxonomicName id="7C9D4D13FF97FFFDFBE2F965FB3D7DAD" box="[1109,1240,1737,1762]" class="Actinopterygii" family="Nemacheilidae" genus="Paracobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFBE2F965FB3D7DAD" box="[1109,1240,1737,1762]" italics="true" pageId="4" pageNumber="37">Paracobitis</emphasis>
|
||
</taxonomicName>
|
||
requires further investigation.
|
||
</paragraph>
|
||
<paragraph id="BB223690FF97FFFCFF70F8BEFE6E7B9A" blockId="4.[151,1437,621,2014]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="38" pageId="4" pageNumber="37">
|
||
In addition, we found high variability in the distribution of the breeding tubercles on the head and body among the species of
|
||
<taxonomicName id="7C9D4D13FF97FFFDFE82F899FE197C01" box="[309,508,1845,1870]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF97FFFDFE82F899FE197C01" box="[309,508,1845,1870]" italics="true" pageId="4" pageNumber="37">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
. In the some species (i.e., the members of the
|
||
<emphasis id="89E9EA82FF97FFFDFC49F89AFB3C7C01" box="[1022,1241,1845,1870]" italics="true" pageId="4" pageNumber="37">
|
||
<taxonomicName id="7C9D4D13FF97FFFDFC49F89AFB987C01" box="[1022,1149,1846,1870]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="angorae">
|
||
<collectingCountry id="C38A7600FF97FFFDFC49F89AFBF07C02" box="[1022,1045,1846,1869]" name="Iceland" pageId="4" pageNumber="37">O</collectingCountry>
|
||
. angorae
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName id="7C9D4D13FF97FFFDFB30F899FB3C7C01" box="[1159,1241,1845,1870]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="brandti">brandti</taxonomicName>
|
||
</emphasis>
|
||
–complexes from the Caspian Sea drainage) the breeding tubercles are rather homogeneously distributed on the head and body of the nuptial males, while in the others (including the species described in the present paper) the breeding tubercles are aggregated mostly on the snout, paired fins, ventral surface of body and on the area near the base of the dorsal fin. The phylogenetic value of these differences requires further research. The interrelationships of the species of
|
||
<taxonomicName id="7C9D4D13FF96FFFCFF20FF3BFEBB7BFF" box="[151,350,151,176]" class="Actinopterygii" family="Nemacheilidae" genus="Oxynoemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="89E9EA82FF96FFFCFF20FF3BFEBB7BFF" box="[151,350,151,176]" italics="true" pageId="5" pageNumber="38">Oxynoemacheilus</emphasis>
|
||
</taxonomicName>
|
||
and similar genera will be discussed in a separate paper after revision of the additional materials from the other basins.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |