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<document id="2A37A1AB3CEBC79C19B1BDCA72E4BFB2" ID-CLB-Dataset="21673" ID-DOI="10.11646/zootaxa.4789.1.1" ID-GBIF-Dataset="c12aaed8-264c-459c-b1aa-09eaeb686670" ID-ISSN="1175-5326" ID-Zenodo-Dep="3884610" ID-ZooBank="311A27A2-A053-4D20-A43E-EF57DBE55166" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591601966137" checkinUser="plazi" docAuthor="Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit &amp; Katoh, Toru" docDate="2020" docId="03C86C16EB285754FF6EDD91FD89FD65" docLanguage="en" docName="zootaxa.4789.1.1.pdf" docOrigin="Zootaxa 4789 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phortica (Ashima) foliiseta Duda (Duda 1923" docType="treatment" docVersion="10" lastPageNumber="15" masterDocId="FFF1146EEB26575AFFF9DD08FFEBFFAA" masterDocTitle="Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="15" updateTime="1698842174822" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="431766226920B7C9218F2BAF39023C24">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:namePart id="2BB281F0C87380E2653F32235AEB6F17">Toda, Masanori J.</mods:namePart>
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<mods:namePart id="64FBB2AEF1D5D9C2713B2A6F70D295E3">Bänziger, Hans</mods:namePart>
<mods:affiliation id="77D605F578AC507B030AFB0E0B8B68DF">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="BB6133C8826C817DB7D3957B93E0D1E5">Sati, Pradeep C.</mods:namePart>
<mods:affiliation id="9A6CC4FF6616B3F62BA8D6105BD5F379">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="46C989565D5861316326CD4EF64F2564">Fartyal, Rajendra S.</mods:namePart>
<mods:affiliation id="EB23DD2D252B767FC1931FCCAAFAC04A">Systematic &amp; Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. &amp; rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:namePart id="BCBC981D849A48E5779974BAE1F3667D">Suwito, Awit</mods:namePart>
<mods:affiliation id="80E0CCC8AACC684D8C3C777653DC38CA">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<mods:namePart id="386EE6F408E193257F8A1E8CA54A4493">Katoh, Toru</mods:namePart>
<mods:affiliation id="0EA9DF08CD6D3164A132B65F64FA9713">Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545</mods:affiliation>
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<mods:date id="F3ED2B01F1EF736C3D120665EC87405C">2020</mods:date>
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<treatment id="03C86C16EB285754FF6EDD91FD89FD65" ID-DOI="http://doi.org/10.5281/zenodo.5221278" ID-GBIF-Taxon="164612808" ID-Zenodo-Dep="5221278" LSID="urn:lsid:plazi:treatment:03C86C16EB285754FF6EDD91FD89FD65" httpUri="http://treatment.plazi.org/id/03C86C16EB285754FF6EDD91FD89FD65" lastPageNumber="15" pageId="14" pageNumber="15">
<subSubSection id="C37B8E8BEB285754FF6EDD91FCC4FF1E" box="[151,815,153,180]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BDEDD00EB285754FF6EDD91FCC4FF1E" blockId="14.[151,815,153,180]" box="[151,815,153,180]" pageId="14" pageNumber="15">
<heading id="D0966A6CEB285754FF6EDD91FCC4FF1E" bold="true" box="[151,815,153,180]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B9150112EB285754FF6EDD91FCC4FF1E" bold="true" box="[151,815,153,180]" pageId="14" pageNumber="15">
2.1.
<taxonomicName id="4C61A683EB285754FF32DD91FDF9FF1E" ID-CoL="4GS4N" authorityName="Duda (Duda" authorityYear="1923" box="[203,530,153,180]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="foliiseta" subGenus="Ashima">
<emphasis id="B9150112EB285754FF32DD91FED8FF1E" bold="true" box="[203,307,153,180]" italics="true" pageId="14" pageNumber="15">Phortica</emphasis>
(
<emphasis id="B9150112EB285754FEBDDD91FE4AFF1E" bold="true" box="[324,417,153,180]" italics="true" pageId="14" pageNumber="15">Ashima</emphasis>
)
<emphasis id="B9150112EB285754FE4BDD91FDF9FF1E" bold="true" box="[434,530,153,180]" italics="true" pageId="14" pageNumber="15">foliiseta</emphasis>
</taxonomicName>
species subgroup, new
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37B8E8BEB285754FF6EDDE9FD89FD65" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BDEDD00EB285754FF6EDDE9FE76FEEE" blockId="14.[151,1436,225,720]" pageId="14" pageNumber="15">
<emphasis id="B9150112EB285754FF6EDDE9FEF9FF56" bold="true" box="[151,274,225,252]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
Surstylus without peg-like teeth (ch.37-1;
<figureCitation id="135AC185EB285754FD0DDDE9FCA5FF51" box="[756,846,225,251]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="14" pageNumber="15">Figs 4B</figureCitation>
,
<figureCitation id="135AC185EB285754FCA2DDE9FC95FF56" box="[859,894,225,252]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="14" pageNumber="15">5C</figureCitation>
); pregonite basally with recurved, apically truncate extension (ch.49-3;
<figureCitation id="135AC185EB285754FE50DC0DFDE9FEB5" box="[425,514,261,287]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="14" pageNumber="15">Figs 4C</figureCitation>
,
<figureCitation id="135AC185EB285754FDF4DC0DFDD9FE8A" box="[525,562,261,288]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="14" pageNumber="15">5D</figureCitation>
); posterior, rod-like process of phallapodeme longer than aedeagal median rod (ch.62-1;
<figureCitation id="135AC185EB285754FEFDDC21FEB5FEE9" box="[260,350,297,323]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="14" pageNumber="15">Figs 4C</figureCitation>
,
<figureCitation id="135AC185EB285754FE93DC21FE64FEEE" box="[362,399,297,324]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="14" pageNumber="15">5D</figureCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB285754FF3EDC45FDC9FE52" blockId="14.[151,1436,225,720]" pageId="14" pageNumber="15">
<emphasis id="B9150112EB285754FF3EDC45FE42FEC2" bold="true" box="[199,425,333,360]" pageId="14" pageNumber="15">Shared characters.</emphasis>
Prementum without pubescent patches (ch.12-0). Setulae on medial surface of abdominal sternites IV and V not so dense (ch.27,28-0;
<figureCitation id="135AC185EB285754FD7CDC79FD37FE26" box="[645,732,369,396]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="14" pageNumber="15">Fig. 5B</figureCitation>
). Setae on and/or near ventral margin of epandrium longer than others (ch.33-1;
<figureCitation id="135AC185EB285754FEB7DC9DFE4CFE05" box="[334,423,405,431]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="14" pageNumber="15">Figs 4B</figureCitation>
,
<figureCitation id="135AC185EB285754FE4ADC9DFE3CFE1A" box="[435,471,405,432]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="14" pageNumber="15">5C</figureCitation>
). Aedeagal, outer membrane without spiny ornamentation (ch.55-1;
<figureCitation id="135AC185EB285754FB36DC9DFAD4FE1A" box="[1231,1343,405,432]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="14" pageNumber="15">Fig. 5D,E</figureCitation>
); aedeagal, median rod less sclerotized (ch.58-2;
<figureCitation id="135AC185EB285754FD97DCB1FD2CFE7E" box="[622,711,441,468]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="14" pageNumber="15">Fig. 5D</figureCitation>
). Aedeagal sheath with lateral lobes bearing strongly sclerotized postgonites (ch.65-1;
<figureCitation id="135AC185EB285754FE72DCD5FE0CFE5D" box="[395,487,477,503]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda &amp; Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="14" pageNumber="15">Figs 4D</figureCitation>
,
<figureCitation id="135AC185EB285754FE0BDCD5FDFFFE52" box="[498,532,477,504]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati &amp; Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="14" pageNumber="15">5E</figureCitation>
).
</paragraph>
<paragraph id="8BDEDD00EB285754FF3EDF0AFD89FD65" blockId="14.[151,1436,225,720]" pageId="14" pageNumber="15">
<emphasis id="B9150112EB285754FF3EDF0AFE99FDB6" bold="true" box="[199,370,514,540]" pageId="14" pageNumber="15">Included taxa.</emphasis>
<taxonomicName id="4C61A683EB285754FE81DF09FD10FDB1" authority="Chen" authorityName="Chen in Chen &amp; Maca" authorityYear="2012" box="[376,763,513,540]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="biclavata" subGenus="Ashima">
<emphasis id="B9150112EB285754FE81DF09FE31FDB1" box="[376,474,513,539]" italics="true" pageId="14" pageNumber="15">Phortica</emphasis>
(
<emphasis id="B9150112EB285754FE13DF09FDD5FDB1" box="[490,574,513,539]" italics="true" pageId="14" pageNumber="15">Ashima</emphasis>
)
<emphasis id="B9150112EB285754FDB4DF09FD5EFDB1" box="[589,693,513,539]" italics="true" pageId="14" pageNumber="15">biclavata</emphasis>
Chen
</taxonomicName>
;
<emphasis id="B9150112EB285754FCFCDF0AFCFDFDB1" box="[773,790,514,539]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FCDDDF09FB3CFDB1" authority="Gong, Zhu &amp; Chen" authorityName="Gong, Zhu &amp; Chen" authorityYear="2018" box="[804,1239,513,540]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dianmianensis" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FCD4DF0AFCD5FDB6" box="[813,830,514,540]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FCACDF09FC1CFDB1" box="[853,1015,513,539]" italics="true" pageId="14" pageNumber="15">dianmianensis</emphasis>
Gong, Zhu &amp; Chen
</taxonomicName>
;
<emphasis id="B9150112EB285754FB18DF0AFB19FDB1" box="[1249,1266,514,539]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FAF9DF09FEB5FD95" authority="Chen &amp; Toda" authorityName="Chen &amp; Toda" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="foliisetoides" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FAF0DF0AFAF1FDB6" box="[1289,1306,514,540]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FAC8DF09FF57FD95" italics="true" pageId="14" pageNumber="15">foliisetoides</emphasis>
Chen &amp; Toda
</taxonomicName>
;
<emphasis id="B9150112EB285754FE90DF2EFE91FD95" box="[361,378,550,575]" italics="true" pageId="14" pageNumber="15">P</emphasis>
. (
<emphasis id="B9150112EB285754FE6BDF2EFE48FDEA" box="[402,419,550,576]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FE42DF2DFE0CFD95" box="[443,487,549,575]" italics="true" pageId="14" pageNumber="15">libo</emphasis>
Zhu, Qin &amp; Chen*;
<emphasis id="B9150112EB285754FD29DF2EFD0AFD95" box="[720,737,550,575]" italics="true" pageId="14" pageNumber="15">P</emphasis>
. (
<emphasis id="B9150112EB285754FD00DF2EFCE1FDEA" box="[761,778,550,576]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FCD8DF2DFC62FD95" box="[801,905,549,575]" italics="true" pageId="14" pageNumber="15">menglian</emphasis>
Gong, Zhu &amp; Chen*;
<emphasis id="B9150112EB285754FB7EDF2EFB73FD95" box="[1159,1176,550,575]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FB5FDF2DFA92FD95" authority="Chen" authorityName="Chen in Chen &amp; Maca" authorityYear="2012" box="[1190,1401,549,576]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nakanoi" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FB49DF2EFB2AFDEA" box="[1200,1217,550,576]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FB21DF2DFAD9FD95" box="[1240,1330,549,575]" italics="true" pageId="14" pageNumber="15">nakanoi</emphasis>
Chen
</taxonomicName>
;
<emphasis id="B9150112EB285754FA7DDF2EFA7EFD95" box="[1412,1429,550,575]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FF6EDF41FEB1FDC9" authorityName="Tsacas &amp; Okada" box="[151,346,585,612]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="phyllochaeta" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FF58DF42FF59FDCE" box="[161,178,586,612]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FF30DF41FEB1FDC9" box="[201,346,585,611]" italics="true" pageId="14" pageNumber="15">phyllochaeta</emphasis>
</taxonomicName>
;
<emphasis id="B9150112EB285754FE91DF42FE92FDC9" box="[360,377,586,611]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FE7EDF41FD03FDCE" authority="Chen &amp; Wen" authorityName="Chen &amp; Wen in Chen et al." authorityYear="2005" box="[391,744,585,612]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sagittaristula" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FE69DF42FE4AFDCE" box="[400,417,586,612]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FE41DF41FDA6FDC9" box="[440,589,585,611]" italics="true" pageId="14" pageNumber="15">sagittaristula</emphasis>
Chen &amp; Wen
</taxonomicName>
;
<emphasis id="B9150112EB285754FD0ADF42FCEFFDC9" box="[755,772,586,611]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FCEBDF41FB8DFDCE" authority="Chen &amp; Wen" authorityName="Chen &amp; Wen in Chen et al." authorityYear="2005" box="[786,1126,585,612]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="saltiaristula" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FCE5DF42FCC6FDCE" box="[796,813,586,612]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FCBDDF41FC20FDC9" box="[836,971,585,611]" italics="true" pageId="14" pageNumber="15">saltiaristula</emphasis>
Chen &amp; Wen
</taxonomicName>
;
<emphasis id="B9150112EB285754FB88DF42FB69FDC9" box="[1137,1154,586,611]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FB69DF41FADDFDC9" authorityName="Chen &amp; Toda" authorityYear="2005" box="[1168,1334,585,612]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="symmetria" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FB60DF42FB41FDCE" box="[1177,1194,586,612]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FB38DF42FADDFDC9" box="[1217,1334,586,611]" italics="true" pageId="14" pageNumber="15">symmetria</emphasis>
</taxonomicName>
*;
<emphasis id="B9150112EB285754FAAADF42FA8FFDC9" box="[1363,1380,586,611]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FA88DF41FED2FD2D" authorityName="Senior-White" authorityYear="1921" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="xyleboriphaga" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FA82DF42FA67FDCE" box="[1403,1420,586,612]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FF6EDF65FED2FD2D" box="[151,313,621,647]" italics="true" pageId="14" pageNumber="15">xyleboriphaga</emphasis>
</taxonomicName>
*;
<emphasis id="B9150112EB285754FEAFDF66FE8CFD2D" box="[342,359,622,647]" italics="true" pageId="14" pageNumber="15">P</emphasis>
. (
<emphasis id="B9150112EB285754FE87DF66FE64FD22" box="[382,399,622,648]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FE5EDF65FDECFD2D" box="[423,519,621,647]" italics="true" pageId="14" pageNumber="15">yalianae</emphasis>
Zhu, Qin &amp; Chen*;
<emphasis id="B9150112EB285754FD09DF66FCEAFD2D" box="[752,769,622,647]" italics="true" pageId="14" pageNumber="15">P</emphasis>
. (
<emphasis id="B9150112EB285754FCE1DF66FCC2FD22" box="[792,809,622,648]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FCB8DF66FC6AFD2D" box="[833,897,622,647]" italics="true" pageId="14" pageNumber="15">yiqini</emphasis>
Zhu, Qin &amp; Chen*; and
<emphasis id="B9150112EB285754FB60DF66FB41FD2D" box="[1177,1194,622,647]" italics="true" pageId="14" pageNumber="15">P</emphasis>
.
<taxonomicName id="4C61A683EB285754FB41DF65FAAFFD2D" authorityName="Duda (Duda" authorityYear="1923" box="[1208,1348,621,648]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="foliiseta" subGenus="Ashima">
(
<emphasis id="B9150112EB285754FB38DF66FB39FD22" box="[1217,1234,622,648]" italics="true" pageId="14" pageNumber="15">A</emphasis>
.)
<emphasis id="B9150112EB285754FB13DF65FAAFFD2D" box="[1258,1348,621,647]" italics="true" pageId="14" pageNumber="15">foliiseta</emphasis>
</taxonomicName>
species complex. * The assignment was judged from the original descriptions by
<bibRefCitation id="EFF0A0F1EB285754FC31DF99FB59FD01" author="Senior-White" box="[968,1202,657,683]" firstAuthor="Senior-White" pageId="14" pageNumber="15" pagination="381 - 396" refId="ref36831" refString="Senior-White, R. A. (1921) New Ceylon Diptera. Spolia Zeylanica, 11, 381 - 396. https: // doi. org / 10.5962 / bhl. part. 12654" type="journal article" year="1921">Senior-White (1921)</bibRefCitation>
,
<bibRefCitation id="EFF0A0F1EB285754FB44DF99FA73FD06" author="Chen" box="[1213,1432,657,684]" etAl="et al." firstAuthor="Chen" pageId="14" pageNumber="15" pagination="419 - 429" refId="ref34109" refString="Chen, H. W., Toda, M. J. &amp; Gao, J. J. (2005 b) The Phortica (s. str.) foliiseta species-complex (Diptera, Drosophilidae) from China and its adjacent countries. Acta Zootaxonomica Sinica, 30, 419 - 429." type="journal article" year="2005" yearSuffix="b">
Chen
<emphasis id="B9150112EB285754FB06DF9AFAC4FD01" box="[1279,1327,657,683]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2005b)
</bibRefCitation>
,
<bibRefCitation id="EFF0A0F1EB285754FF6EDFBDFEBEFD65" author="Zhu" box="[151,341,693,720]" etAl="et al." firstAuthor="Zhu" pageId="14" pageNumber="15" pagination="71 - 82" refId="ref38187" refString="Zhu, L., Qin, T. &amp; Chen, H. W. (2017) Three new species of the subgenus Ashima (Diptera, Drosophilidae, Phortica) from southwestern China, with DNA barcoding information. Journal of Natural History, 51, 71 - 82. https: // doi. org / 10.1080 / 00222933.2016.1247480" type="journal article" year="2017">
Zhu
<emphasis id="B9150112EB285754FF32DFBEFF16FD65" box="[203,253,693,719]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2017)
</bibRefCitation>
and
<bibRefCitation id="EFF0A0F1EB285754FE72DFBDFDB6FD65" author="Gong" box="[395,605,693,720]" etAl="et al." firstAuthor="Gong" pageId="14" pageNumber="15" pagination="434 - 446" refId="ref34842" refString="Gong, L., Zhu, L. &amp; Chen, H. W. (2018) Six new species of the subgenera Alloparadisa and Ashima (Diptera: Drosophilidae: Phortica) from Yunnan, China, with DNA barcoding information. Annales de la Societe entomologique de France, New Series, 54, 434 - 446. https: // doi. org / 10.1080 / 00379271.2018.1491327" type="journal article" year="2018">
Gong
<emphasis id="B9150112EB285754FE28DFBEFDE9FD65" box="[465,514,693,719]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (2018)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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