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<document id="381B0922C383174C36B3B8D2B082F31F" ID-DOI="10.5281/zenodo.5330604" ID-ISSN="0374-1036" ID-Zenodo-Dep="5330604" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168017471" checkinUser="felipe" docAuthor="Fikáček, Martin, Prokin, Alexander, Angus, Robert B., Pono, Alexander, Marenko, Yue, Yanli, Ren, Dong &amp; Prokop, Jakub" docDate="2012" docId="B970A055FFC7FFE2FEA3278DFBB8FD5C" docLanguage="en" docName="ActaEntMusNatPra.2012.52.1.89-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Laetopsia bontsaganica Fikáček &amp; Prokin &amp; Angus &amp; Pono &amp; Marenko &amp; Yue &amp; Ren &amp; Prokop 2012, comb. nov." docType="treatment" docVersion="4" lastPageNumber="96" masterDocId="4549D82DFFC0FFE5FF9B2756FFB9FFFB" masterDocTitle="Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)" masterLastPageNumber="127" masterPageNumber="89" pageNumber="96" updateTime="1699171640213" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="71F3A536ECFB6FB221769B0AF4CCC7AE">Revision of Mesozoic fossils of the helophorid lineage of the superfamily Hydrophiloidea (Coleoptera: Polyphaga)</mods:title>
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<mods:namePart id="E869302240897CE73E0F288D43D646A7">Fikáček, Martin</mods:namePart>
<mods:affiliation id="B599C074D35C367D80AA8ADDAB5645B1">Department of Entomology, National Museum, Kunratice 1, CZ- 148 00 Praha 4, Czech Republic; e-mail: mfikacek @ gmail. com &amp; Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:namePart id="D6D9D7378E9F7C74EFF98D560381757A">Prokin, Alexander</mods:namePart>
<mods:affiliation id="9BACC1B5055146674BD621BC1B169503">Papanin Institute for Inland Water Biology, Russian Academy of Science, Borok, Russia; e-mail: prokina @ mail. ru &amp; Voronezh State University, Research-Educational Centre “ Venevitinovo ”, Voronezh, Russia</mods:affiliation>
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<mods:namePart id="E41E3139D237DE6A1BAA406E39F90EF2">Angus, Robert B.</mods:namePart>
<mods:affiliation id="616E1280B375ADCC1519F0D5EB8034FD">Department of Entomology, The Natural History Museum, London, UK; e-mail: r. angus @ rhul. ac. uk</mods:affiliation>
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<mods:namePart id="7A70932D3EBB1631D18F4DDDB8202A7B">Pono, Alexander</mods:namePart>
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<mods:namePart id="BD14DDC30EDE87BD9DC258FF4A537526">Marenko</mods:namePart>
<mods:affiliation id="4965D90B063132692BDC023BEF138BA7">Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, Russia; e-mail: aponom @ paleo. ru</mods:affiliation>
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<mods:namePart id="96D8EE006CBA24CF28EE02F239C6D6FE">Yue, Yanli</mods:namePart>
<mods:affiliation id="FFA5CCF7A268E8F17B0014E8CA632617">School of Life Science, Ningxia University, Ningxia, China; e-mail: yueyanli 0407 @ yahoo. cn</mods:affiliation>
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<mods:namePart id="453D7E0D2892B5AC8C18937B48C4883D">Ren, Dong</mods:namePart>
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<mods:namePart id="CBF8E7600AC9E82862F5FAABFE36FC50">Prokop, Jakub</mods:namePart>
<mods:affiliation id="7367BFC27B0940E20FA6D08580982526">Department of Zoology, Faculty of Science, Charles University in Prague, Viničná 7, CZ- 128 44 Praha 2, Czech Republic; e-mail: jprokop @ natur. cuni. cz</mods:affiliation>
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<mods:title id="165FFEEB5387B280A82B4BB3744ACAAD">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="D7289D19A7A06A5C8D8D9474ED0867C1">2012</mods:date>
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<mods:identifier id="339B142DBC326C097F10209ED881FAB9" type="ISSN">0374-1036</mods:identifier>
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<paragraph id="31661143FFC7FFE2FEA3278DFCC3FF08" blockId="7.[312,890,218,243]" box="[312,890,218,243]" pageId="7" pageNumber="96">
<heading id="6A2EA62FFFC7FFE2FEA3278DFCC3FF08" bold="true" box="[312,890,218,243]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="96" reason="3">
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<taxonomicName id="F6D96AC0FFC7FFE2FED7278DFD40FF08" ID-CoL="6NT75" authority="(Prokin, 2009)" authorityName="Fikáček &amp; Prokin &amp; Angus &amp; Pono &amp; Marenko &amp; Yue &amp; Ren &amp; Prokop" authorityYear="2012" baseAuthorityName="Prokin" baseAuthorityYear="2009" box="[332,761,218,243]" class="Insecta" family="Helophoridae" genus="Laetopsia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="bontsaganica" status="comb. nov.">
<emphasis id="03ADCD51FFC7FFE2FED7278DFDE9FF08" bold="true" box="[332,592,218,243]" italics="true" pageId="7" pageNumber="96">Laetopsia bontsaganica</emphasis>
(
<bibRefCitation id="55486CB2FFC7FFE2FDC5278DFD4BFF08" author="PROKIN A." box="[606,754,219,243]" pageId="7" pageNumber="96" pagination="660 - 663" refId="ref15856" refString="PROKIN A. 2009: New water scavenger beetles (Coleoptera: Hydrophilidae) from the Mesozoic of Mongolia. Paleontological Journal 43: 660 - 663." type="journal article" year="2009">Prokin, 2009</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="189E702AFFC7FFE2FC9B278DFCC3FF08" box="[768,890,219,243]" pageId="7" pageNumber="96" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="79C342C8FFC7FFE2FDAF27AFFDC7FEF7" box="[564,638,249,268]" pageId="7" pageNumber="96" type="description">
<paragraph id="31661143FFC7FFE2FDAF27AFFDC7FEF7" blockId="7.[113,638,249,330]" box="[564,638,249,268]" pageId="7" pageNumber="96">
(
<figureCitation id="A9E20DC6FFC7FFE2FDA127AFFDC1FEF7" box="[570,632,249,268]" captionStart="Figs" captionStartId="23.[113,152,1537,1556]" captionTargetBox="[136,1066,219,1523]" captionTargetId="figure-15@23.[136,1066,219,1523]" captionTargetPageId="23" captionText="Figs. 5362.Representatives of Laetopsia gen.nov.5356 L.baissensis (Ponomarenko, 1986) (53 PIN 3064/7016; 5455 PIN 3064/6987, piece and counterpiece; 56 detail of PIN 3064/6987). 57, 61 L. hydraenoides (Prokin, Ren &amp; Fikáček, 2010), CNU 2010021, holotype (57 detail of head and pronotum; 61 general view). 5860 L. mongolica (Ponomarenko, 1987) (58 PIN 3664/1661, detail of head and pronotum; 59 PIN 3664/1772; 60 PIN 3664/1567, holotype). 62 L. bontsaganica (Prokin, 2009), holotype. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5330645" httpUri="https://zenodo.org/record/5330645/files/figure.png" pageId="7" pageNumber="96">Fig. 62</figureCitation>
)
</paragraph>
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<paragraph id="31661143FFC7FFE2FFEA264BFD89FEB2" blockId="7.[113,638,249,330]" pageId="7" pageNumber="96">
<taxonomicName id="F6D96AC0FFC7FFE2FFEA264BFDB6FED4" ID-CoL="6MJRX" authority="Prokin, 2009 a: 54" authorityName=": PROKIN" authorityPageNumber="54" authorityYear="2009" box="[113,527,284,303]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="bontsaganica">
<emphasis id="03ADCD51FFC7FFE2FFEA264BFED6FED4" box="[113,367,285,303]" italics="true" pageId="7" pageNumber="96">Hydrophilopsia bontsaganica</emphasis>
<bibRefCitation id="55486CB2FFC7FFE2FEEF264AFE5FFED4" author="PROKIN A." box="[372,486,284,303]" pageId="7" pageNumber="96" pagination="660 - 663" refId="ref15856" refString="PROKIN A. 2009: New water scavenger beetles (Coleoptera: Hydrophilidae) from the Mesozoic of Mongolia. Paleontological Journal 43: 660 - 663." type="journal article" year="2009">Prokin, 2009</bibRefCitation>
a: 54
</taxonomicName>
.
<taxonomicName id="F6D96AC0FFC7FFE2FFEA2661FD95FEB2" ID-CoL="6MJRX" authority=": PROKIN (2009 b: 660)" authorityName=": PROKIN" authorityPageNumber="660" authorityYear="2009" box="[113,556,310,330]" class="Insecta" family="Helophoridae" genus="Hydrophilopsia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="species" species="bontsaganica">
<emphasis id="03ADCD51FFC7FFE2FFEA2661FED6FEB2" box="[113,367,311,329]" italics="true" pageId="7" pageNumber="96">Hydrophilopsia bontsaganica</emphasis>
:
<bibRefCitation id="55486CB2FFC7FFE2FEE12661FE56FEB2" author="PROKIN A." box="[378,495,310,330]" pageId="7" pageNumber="96" pagination="660 - 663" refId="ref15856" refString="PROKIN A. 2009: New water scavenger beetles (Coleoptera: Hydrophilidae) from the Mesozoic of Mongolia. Paleontological Journal 43: 660 - 663." type="journal article" year="2009">PROKIN (2009</bibRefCitation>
b: 660)
</taxonomicName>
.
</paragraph>
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<subSubSection id="79C342C8FFC7FFE2FFEA2636FC1AFE43" pageId="7" pageNumber="96" type="materials_examined">
<paragraph id="31661143FFC7FFE2FFEA2636FCCFFE75" blockId="7.[113,1090,351,398]" pageId="7" pageNumber="96">
<emphasis id="03ADCD51FFC7FFE2FFEA2636FE8CFE88" bold="true" box="[113,309,351,371]" pageId="7" pageNumber="96">
<typeStatus id="EE62AFE1FFC7FFE2FFEA2636FF27FE88" box="[113,158,352,371]" pageId="7" pageNumber="96">Type</typeStatus>
locality and age.
</emphasis>
<collectingCountry id="49CE51D3FFC7FFE2FEA12636FE36FE88" box="[314,399,352,371]" name="Mongolia" pageId="7" pageNumber="96">Mongolia</collectingCountry>
,
<collectingRegion id="F31DDFA1FFC7FFE2FE0C2636FDDFFE88" box="[407,614,352,371]" country="Mongolia" name="Bayanhongor" pageId="7" pageNumber="96">Bayankhongor province</collectingRegion>
, foothills of the Dund Ula Mts., S of Bööntsagaan Lake. Lower Cretaceous, Barremian or Aptian, ca. 130112 mya (
<bibRefCitation id="55486CB2FFC7FFE2FDF4262DFCD5FE75" author="RASNITSYN A. P. &amp; ZHERIKHIN V. V." box="[623,876,379,398]" pageId="7" pageNumber="96" pagination="437 - 444" refId="ref15948" refString="RASNITSYN A. P. &amp; ZHERIKHIN V. V. 2002: Alphabetic list of selected insect fossil sites - Impression fossils. Pp. 437 - 444. In: RASNITSYN A. P. &amp; QUICKE D. L. J. (eds.): History of Insects. Kluwer Academic Publishers, Norwell, 517 pp." type="book chapter" year="2002">RASNITSYN &amp; ZHERIKHIN 2002</bibRefCitation>
).
</paragraph>
<paragraph id="31661143FFC7FFE2FFEA26F2FC1AFE43" blockId="7.[113,931,420,440]" box="[113,931,420,440]" pageId="7" pageNumber="96">
<emphasis id="03ADCD51FFC7FFE2FFEA26F2FEA7FE43" bold="true" box="[113,286,420,440]" pageId="7" pageNumber="96">Material examined</emphasis>
(
<specimenCount id="27DFDACAFFC7FFE2FEB026F3FED1FE43" box="[299,360,421,440]" pageId="7" pageNumber="96" type="generic">1 spec.</specimenCount>
).
<typeStatus id="EE62AFE1FFC7FFE2FEEC26F3FE72FE4D" box="[375,459,421,440]" pageId="7" pageNumber="96" type="holotype">HOLOTYPE</typeStatus>
: PIN 3559/6370 (piece only, specimen in dorsal view).
</paragraph>
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<subSubSection id="79C342C8FFC7FFE2FFEA2686FBB8FD5C" pageId="7" pageNumber="96" type="description">
<paragraph id="31661143FFC7FFE2FFEA2686FE56FDB3" blockId="7.[113,1089,464,680]" pageId="7" pageNumber="96">
<emphasis id="03ADCD51FFC7FFE2FFEA2686FEAEFE13" bold="true" box="[113,279,464,488]" pageId="7" pageNumber="96">Redescription.</emphasis>
Body elongate oval,
<quantity id="F621BCA6FFC7FFE2FD9C2686FDE2FE1C" box="[519,603,464,488]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="7" pageNumber="96" unit="mm" value="8.8">8.8 mm</quantity>
long. Head with very distinct frontoclypeal suture, its median portion grooved, anterior portion of clypeus straight. Pronotum slightly transverse, bearing median groove, lateral margins smooth, arcuate. Elytron with darkened striae. Abdomen with five ventrites.
</paragraph>
<paragraph id="31661143FFC7FFE2FFEA2506FBB8FD5C" blockId="7.[113,1089,464,680]" pageId="7" pageNumber="96">
<emphasis id="03ADCD51FFC7FFE2FFEA2506FF17FD93" bold="true" box="[113,174,592,616]" pageId="7" pageNumber="96">Note.</emphasis>
Although the only known specimen is poorly preserved, the combination of elytra with dark striae, the distinct frontoclypeal suture with grooved median portion and the presence of the median groove on the pronotum clearly assign the species to the genus
<taxonomicName id="F6D96AC0FFC7FFE2FC0B25C7FC40FD53" authorityName="Fikáček &amp; Prokin &amp; Angus &amp; Pono &amp; Marenko &amp; Yue &amp; Ren &amp; Prokop" authorityYear="2012" box="[912,1017,657,680]" class="Insecta" family="Helophoridae" genus="Laetopsia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="03ADCD51FFC7FFE2FC0B25C7FC40FD53" box="[912,1017,657,680]" italics="true" pageId="7" pageNumber="96">Laetopsia</emphasis>
</taxonomicName>
.
</paragraph>
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