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<document id="7CB4A298C5F465DDE3279A15193353B1" ID-DOI="10.5281/zenodo.4525859" ID-ISSN="1638-9387" ID-Zenodo-Dep="4525859" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612879736561" checkinUser="felipe" docAuthor="Ahrens, Dirk &amp; Fabrizi, Silvia" docDate="2008" docId="1354741BFFF6FFB5FF355473A118FED3" docLanguage="en" docName="Zoosystema.30.4.917-927.pdf" docOrigin="Zoosystema 30 (4)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Emphania ranomafanae Ahrens &amp; Fabrizi 2008, n. sp." docType="treatment" docVersion="4" lastPageNumber="926" masterDocId="EF6D0C63FFFFFFBCFFE25667A544FFC0" masterDocTitle="A taxonomic revision of the Malagasy genus Emphania Erichson, 1847 (Insecta, Coleoptera, Scarabaeidae, Sericini)" masterLastPageNumber="927" masterPageNumber="917" pageNumber="926" updateTime="1698869142263" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F4DCD98AC334E65FC1DB32A15287A093">A taxonomic revision of the Malagasy genus Emphania Erichson, 1847 (Insecta, Coleoptera, Scarabaeidae, Sericini)</mods:title>
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<mods:namePart id="D3E91A5C6F9363BDBA9D91D30EAF6AFD">Ahrens, Dirk</mods:namePart>
<mods:affiliation id="BA146D6B754597299751EDBCF05FCCC0">The Natural History Museum, Department of Entomology, Cromwell Road, London SW 7 5 BD (United Kingdom) d. ahrens @ nhm. ac. uk</mods:affiliation>
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<mods:namePart id="763ADA933FD4B649DAF30C63B35422CC">Fabrizi, Silvia</mods:namePart>
<mods:affiliation id="EDF9EE84099A31A80C712905E4FE511D">The Natural History Museum, Department of Entomology, Cromwell Road, London SW 7 5 BD (United Kingdom)</mods:affiliation>
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<mods:title id="2208B77B7A12314EBBA85197C81351AA">Zoosystema</mods:title>
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<mods:date id="EBFFE0F0ADFBC8EBF5221BBF0FBBEB60">2008</mods:date>
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<treatment id="1354741BFFF6FFB5FF355473A118FED3" ID-DOI="http://doi.org/10.5281/zenodo.4525717" ID-Zenodo-Dep="4525717" LSID="urn:lsid:plazi:treatment:1354741BFFF6FFB5FF355473A118FED3" httpUri="http://treatment.plazi.org/id/1354741BFFF6FFB5FF355473A118FED3" lastPageNumber="926" pageId="9" pageNumber="926">
<subSubSection id="D3E79686FFF6FFB5FF355473A765FDF0" box="[215,545,532,560]" pageId="9" pageNumber="926" type="nomenclature">
<paragraph id="9B42C50DFFF6FFB5FF355473A765FDF0" blockId="9.[215,545,532,560]" box="[215,545,532,560]" pageId="9" pageNumber="926">
<heading id="C00A7261FFF6FFB5FF355473A765FDF0" box="[215,545,532,560]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="926" reason="2">
<taxonomicName id="5CFDBE8EFFF6FFB5FF355473A49BFDEE" authority="Ahrens &amp; Fabrizi, 2008" authorityName="Ahrens &amp; Fabrizi" authorityYear="2008" box="[215,479,532,559]" class="Insecta" family="Melolonthidae" genus="Emphania" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="926" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="A989191FFFF6FFB5FF355473A49BFDEE" bold="true" box="[215,479,532,559]" italics="true" pageId="9" pageNumber="926">Emphania ranomafanae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2BAA464FFF6FFB5FE045472A765FDF0" box="[486,545,533,560]" pageId="9" pageNumber="926" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="D3E79686FFF6FFB5FF935429A470FCDF" pageId="9" pageNumber="926" type="materials_examined">
<paragraph id="9B42C50DFFF6FFB5FF935429A43FFD75" blockId="9.[112,648,590,799]" pageId="9" pageNumber="926">
<materialsCitation id="2B95CF50FFF6FFB5FF935429A43FFD75" collectingDate="1998-05" collectionCode="TYPE, MATERIAL, CASC" collectorName="F. Lee &amp; K. J. Ribardo &amp; Calif. Acad. Sci. Collection" country="Madagascar" location="Talatakely" municipality="Ranomafana National Park" pageId="9" pageNumber="926" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<collectionCode id="FDEC5DC8FFF6FFB5FF935429A5BAFDA4" box="[113,254,590,613]" pageId="9" pageNumber="926">TYPE MATERIAL</collectionCode>
. —
<typeStatus id="44467BAFFFF6FFB5FEC45429A4C7FDA6" box="[294,387,590,614]" pageId="9" pageNumber="926" type="holotype">Holotype</typeStatus>
:
<specimenCount id="8DFB0E84FFF6FFB5FE685428A4DAFDA6" box="[394,414,591,614]" pageId="9" pageNumber="926" type="male"></specimenCount>
<collectingCountry id="E3EA859DFFF6FFB5FE485429A75AFDA6" box="[426,542,590,614]" name="Madagascar" pageId="9" pageNumber="926">Madagascar</collectingCountry>
:
<collectingRegion id="59390BEFFFF6FFB5FDC75429A5D6FD40" country="Madagascar" name="Fianarantsoa" pageId="9" pageNumber="926">Fianarantsoa</collectingRegion>
:
<collectingMunicipality id="7B265F77FFF6FFB5FF7B540FA4DDFD40" box="[153,409,616,640]" pageId="9" pageNumber="926">Ranomafana National Park</collectingMunicipality>
,
<location id="9E2293D6FFF6FFB5FE7D540FA4BAFD40" LSID="urn:lsid:plazi:treatment:1354741BFFF6FFB5FF355473A118FED3:9E2293D6FFF6FFB5FE7D540FA4BAFD40" box="[415,510,616,640]" country="Madagascar" municipality="Ranomafana National Park" name="Talatakely" pageId="9" pageNumber="926" stateProvince="Fianarantsoa">Talatakely</location>
, research
<date id="EF43E3CDFFF6FFB5FDB7540FA5C1FD5B" pageId="9" pageNumber="926" value="1998-05">
<collectingDate id="FF071A25FFF6FFB5FDB7540FA5C1FD5B" pageId="9" pageNumber="926" value="1998-05">1998 V.</collectingDate>
</date>
<collectorName id="3608A0DBFFF6FFB5FF6A54E4A586FD5B" box="[136,194,643,667]" pageId="9" pageNumber="926">F. Lee</collectorName>
,
<collectorName id="3608A0DBFFF6FFB5FF2854E4A40FFD5B" box="[202,331,643,667]" pageId="9" pageNumber="926">K. J. Ribardo</collectorName>
, leg.
<collectorName id="3608A0DBFFF6FFB5FE9B54E4A739FD5B" box="[377,637,643,667]" pageId="9" pageNumber="926">Calif. Acad. Sci. Collection</collectorName>
/ Casent 8006559” (
<collectionCode id="FDEC5DC8FFF6FFB5FEC954FAA434FD75" box="[299,368,669,693]" pageId="9" pageNumber="926">CASC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="9B42C50DFFF6FFB5FF9354DFA470FCDF" blockId="9.[112,648,590,799]" pageId="9" pageNumber="926">
<typeStatus id="44467BAFFFF6FFB5FF9354DFA58CFD10" box="[113,200,696,720]" pageId="9" pageNumber="926" type="paratype">Paratype</typeStatus>
:
<specimenCount id="8DFB0E84FFF6FFB5FF3354DFA5B0FD10" box="[209,244,696,720]" pageId="9" pageNumber="926" type="female">1 ♀</specimenCount>
<collectingCountry id="E3EA859DFFF6FFB5FEE154DFA43FFD10" box="[259,379,696,720]" name="Madagascar" pageId="9" pageNumber="926">Madagascar</collectingCountry>
:
<collectingRegion id="59390BEFFFF6FFB5FE6154DFA746FD10" box="[387,514,696,720]" country="Madagascar" name="Fianarantsoa" pageId="9" pageNumber="926">Fianarantsoa</collectingRegion>
: Ranomafana National Park, Talatakely, research
<date id="EF43E3CDFFF6FFB5FE2C54B5A75AFD2A" box="[462,542,722,746]" pageId="9" pageNumber="926" value="1998-05">
1998
<collectionCode id="FDEC5DC8FFF6FFB5FDEA54B5A75EFD2A" box="[520,538,722,746]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="926" type="Museum">V</collectionCode>
.
</date>
F. Lee, K. J. Ribardo, leg. Calif. Acad. Sci. Collection/Casent 8006560” (
<collectionCode id="FDEC5DC8FFF6FFB5FF065560A46DFCDF" box="[228,297,775,799]" pageId="9" pageNumber="926">CASC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D3E79686FFF6FFB5FF935550A770FCA9" pageId="9" pageNumber="926" type="etymology">
<paragraph id="9B42C50DFFF6FFB5FF935550A770FCA9" blockId="9.[113,646,823,873]" pageId="9" pageNumber="926">ETYMOLOGY. — The new species is named after its occurrence in the Ranomafana National Park.</paragraph>
</subSubSection>
<subSubSection id="D3E79686FFF6FFB5FF9355F6A70CFA26" pageId="9" pageNumber="926" type="description">
<paragraph id="9B42C50DFFF6FFB5FF9355F6A4D8FC69" blockId="9.[111,649,913,1320]" box="[113,412,913,939]" pageId="9" pageNumber="926">
DESCRIPTION OF
<typeStatus id="44467BAFFFF6FFB5FECF55F1A4D8FC69" box="[301,412,918,937]" pageId="9" pageNumber="926" type="holotype">HOLOTYPE</typeStatus>
</paragraph>
<paragraph id="9B42C50DFFF6FFB5FF9355D7A43BFAC8" blockId="9.[111,649,913,1320]" pageId="9" pageNumber="926">
Length:
<quantity id="5C0568E8FFF6FFB5FF2955D7A460FC0B" box="[203,292,944,971]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="9" pageNumber="926" unit="mm" value="8.5">8.5 mm</quantity>
; length of elytra:
<quantity id="5C0568E8FFF6FFB5FE3C55D7A772FC0B" box="[478,566,944,971]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="9" pageNumber="926" unit="mm" value="5.8">5.8 mm</quantity>
; width:
<quantity id="5C0568E8FFF6FFB5FF9355B7A596FC2B" box="[113,210,976,1003]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="926" unit="mm" value="5.3">5.3 mm</quantity>
. Body surface brown with reddish to greenish shine. Labroclypeus and antenna as in
<taxonomicName id="5CFDBE8EFFF6FFB5FD8D5596A58AFBE9" baseAuthorityName="Blanchard" baseAuthorityYear="1850" class="Insecta" family="Melolonthidae" genus="Heptomera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="926" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="A989191FFFF6FFB5FD8D5596A58AFBE9" italics="true" pageId="9" pageNumber="926">E. metallica</emphasis>
</taxonomicName>
; eyes, ratio of diameter/interocular width: 0.53; antennomeres 3-5 not fused. Pronotum, scutellum and elytra as in
<taxonomicName id="5CFDBE8EFFF6FFB5FE705237A74BFBA9" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[402,527,1103,1129]" class="Insecta" family="Melolonthidae" genus="Heptomera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="926" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="A989191FFFF6FFB5FE705237A74BFBA9" box="[402,527,1103,1129]" italics="true" pageId="9" pageNumber="926">E. metallica</emphasis>
</taxonomicName>
. Metacoxa and pygidium as in
<taxonomicName id="5CFDBE8EFFF6FFB5FEB25208A48BFB48" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[336,463,1134,1160]" class="Insecta" family="Melolonthidae" genus="Heptomera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="926" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="A989191FFFF6FFB5FEB25208A48BFB48" box="[336,463,1134,1160]" italics="true" pageId="9" pageNumber="926">E. metallica</emphasis>
</taxonomicName>
. Ratio of length of metepisternum/metacoxa: 1/1.87. Legs as in
<taxonomicName id="5CFDBE8EFFF6FFB5FD8D52E8A595FB08" baseAuthorityName="Blanchard" baseAuthorityYear="1850" class="Insecta" family="Melolonthidae" genus="Heptomera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="926" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="A989191FFFF6FFB5FD8D52E8A595FB08" italics="true" pageId="9" pageNumber="926">E. metallica</emphasis>
</taxonomicName>
; ratio metatibial width/length: 1/4.25, medial face of metatibia with one or two punctures beside the dorsal margin.
</paragraph>
<paragraph id="9B42C50DFFF6FFB5FF6A536AA4C2FAE8" blockId="9.[111,649,913,1320]" box="[136,390,1293,1320]" pageId="9" pageNumber="926">
Aedeagus:
<figureCitation id="03C6D988FFF6FFB5FF19536AA417FAE8" box="[251,339,1293,1320]" captionStart="FIG" captionStartId="8.[113,124,1612,1629]" captionTargetBox="[186,1133,716,1585]" captionTargetId="figure-229@8.[184,660,1124,1495]" captionTargetPageId="8" captionText="FIG. 2. — A-C, Emphania nitida Moser, 1911 (lectotype); D-F, E. sulcipennis Moser, 1911 (Madagascar Nord, Montagne dAmbre); G-I, E. semiviridis n. sp. (holotype); J-L, E. erichsoni n. sp. (holotype); M-O, E. subsmaragdina n. sp. (holotype); P-R, E. ranomafanae n. sp. (holotype); A, D, G, J, M, P, aedeagus,left side lateral views;B, E, H, K, N, Q, aedeagus,dorsal views;C, F, I, L, O, R, aedeagus, right side lateral views. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4525867" httpUri="https://zenodo.org/record/4525867/files/figure.png" pageId="9" pageNumber="926">Figure 2</figureCitation>
P-R.
</paragraph>
<paragraph id="9B42C50DFFF6FFB5FF93532AA5A3FAA5" blockId="9.[111,647,1357,1510]" box="[113,231,1357,1383]" pageId="9" pageNumber="926">VARIATION</paragraph>
<paragraph id="9B42C50DFFF6FFB5FF93530BA70CFA26" blockId="9.[111,647,1357,1510]" pageId="9" pageNumber="926">No variation in body measurements observed.Colour varies from blackish green to reddish purple. The female has the antennal club slightly shorter and the punctures on pygidium are less distinct.</paragraph>
</subSubSection>
<subSubSection id="D3E79686FFF6FFB5FF93506CA118FED3" pageId="9" pageNumber="926" type="discussion">
<paragraph id="9B42C50DFFF6FFB5FF93506CA118FED3" blockId="9.[111,647,1547,1700]" lastBlockId="9.[675,1208,216,275]" pageId="9" pageNumber="926">
REMARKS The species is very similar in external morphology to
<taxonomicName id="5CFDBE8EFFF6FFB5FF8D502CA5A7F9A4" baseAuthorityName="Blanchard" baseAuthorityYear="1850" box="[111,227,1610,1636]" class="Insecta" family="Melolonthidae" genus="Heptomera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="926" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="A989191FFFF6FFB5FF8D502CA5A7F9A4" box="[111,227,1610,1636]" italics="true" pageId="9" pageNumber="926">E. metallica</emphasis>
</taxonomicName>
, and may externally only be distinguished by the unfused antennomeres 3-5. The elongate shape of aedeagus and the weakly curved parameres provide best features for separating the taxon from all other so far known
<taxonomicName id="5CFDBE8EFFF6FFB5FC7A5690A140FED1" authorityName="Erichson" authorityYear="1847" box="[920,1028,247,273]" class="Insecta" family="Melolonthidae" genus="Emphania" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="926" phylum="Arthropoda" rank="genus">
<emphasis id="A989191FFFF6FFB5FC7A5690A140FED1" box="[920,1028,247,273]" italics="true" pageId="9" pageNumber="926">Emphania</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>