207 lines
25 KiB
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207 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.536.5982" ID-GBIF-Dataset="f3546db1-d96a-40d8-ac8c-fe806547f201" ID-PMC="PMC4714036" ID-Pensoft-Pub="1313-2970-536-1" ID-PubMed="26798233" ID-ZBK="A8B92CDD0B8C4384AAC559648BB45AA5" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-536-1" ModsDocOrigin="ZooKeys 536" ModsDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" checkinTime="1451243741663" checkinUser="pensoft" docAuthor="Shi, Hongliang & Liang, Hongbin" docDate="2015" docId="F3CB5F6DC736B44211A17739B9F0C402" docLanguage="en" docName="ZooKeys 536: 1-92" docOrigin="ZooKeys 536" docSource="http://dx.doi.org/10.3897/zookeys.536.5982" docTitle="Pterostichus (Circinatus) zhygealu Shi & Liang, 2015, sp. n." docType="treatment" docUuid="BB708E40-C604-41BD-AFD2-2389FD3162C7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="29" masterDocId="FFEAFF976442467ED413FFC17531FF81" masterDocTitle="The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)" masterLastPageNumber="92" masterPageNumber="1" pageNumber="25" updateTime="1668160951778" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shi, Hongliang</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liang, Hongbin</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>536</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>92</mods:end>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.536.5982</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.536.5982</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-536-1</mods:identifier>
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<mods:identifier type="ZBK">A8B92CDD0B8C4384AAC559648BB45AA5</mods:identifier>
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<mods:identifier type="ZooBank">A8B92CDD0B8C4384AAC559648BB45AA5</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152063157" LSID="urn:lsid:zoobank.org:act:BB708E40-C604-41BD-AFD2-2389FD3162C7" httpUri="http://treatment.plazi.org/id/F3CB5F6DC736B44211A17739B9F0C402" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25">
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<subSubSection pageId="24" pageNumber="25" type="multiple">
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<paragraph pageId="24" pageNumber="25">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection pageId="24" pageNumber="25" type="nomenclature">
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<paragraph pageId="24" pageNumber="25">
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<taxonomicName LSID="http://zoobank.org/BB708E40-C604-41BD-AFD2-2389FD3162C7" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus) zhygealu" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="zhygealu" subGenus="Circinatus">Pterostichus (Circinatus) zhygealu</taxonomicName>
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<taxonomicNameLabel pageId="24" pageNumber="25">sp. n.</taxonomicNameLabel>
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Figures 8, 38, 54, 75, 94, 117
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="type locality">
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<paragraph pageId="25" pageNumber="26">
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<pageBreakToken pageId="25" pageNumber="26" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph pageId="25" pageNumber="26">
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Sichuan: Dafengding national reserve, Yizi pass between Meigu and Ebian County (
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.67477">N28.67477°</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="103.05248">E103.05248°</geoCoordinate>
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), altitude 2923 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="type material">
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<paragraph pageId="25" pageNumber="26">Type material.</paragraph>
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<paragraph pageId="25" pageNumber="26">
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Holotype (IZAS): male, body length = 13.5 mm, pin mounted, genitalia dissected and glued on plastic film pinned under specimen, "CHINA, Sichuan prov., Yizi / pass btw. Meigu county and / Ebian county, mixed forest, / 2923 m, /
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.67477">N28.67477</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="103.05248">E103.05248</geoCoordinate>
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"; "2012.VI.12, in dead log, / SHI Hongliang, LIU Ye leg., / Institute of Zoology, CAS. /
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<normalizedToken originalValue="美姑县-峨边县椅子垭口”">美姑县-峨边县椅子垭口"</normalizedToken>
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; "HOLOTYPE ♂/ Pterostichus (Circinatus) / zhygealu new species / des. SHI H.L. 2015" [red label]. Paratypes, a total of 3 males and 7 females: 1 male, 2 females (IZAS): the same data as holotype. 2 males, 5 females (IZAS): "CHINA, Sichuan Prov., Yizi pass btw. Meigu county and Ebian county, mixed forest;
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.67477">N28.67477</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="103.05248">E103.05248</geoCoordinate>
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, 2923 m; 2012.VI.15; by pit fall trap; SHI Hongliang & LIU Ye leg.".
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</paragraph>
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</subSubSection>
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<subSubSection pageId="25" pageNumber="26" type="type material">
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<paragraph pageId="25" pageNumber="26">Non-type material.</paragraph>
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<paragraph pageId="25" pageNumber="26">
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1 female (IZAS): "CHINA, Sichuan, Mianning county, Yele conservation, mixed forest,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="28.96508">N28.96508</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="102.16137">E102.16137</geoCoordinate>
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; 2988 m, 2012.VI.22, under stone, YANG Xiaodong lgt."
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="diagnosis">
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<paragraph pageId="25" pageNumber="26">Diagnosis.</paragraph>
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<paragraph lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
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Relatively large-sized species within subgenus; pronotum with three or four mid-lateral setae; hind angle completely rounded; outer basal foveal groove faintly defined, approx half length of inner one, outer area of inner groove distinctly concave; elytron with distinct transverse or isodiametric microsculpture; humeral tooth indis
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<pageBreakToken pageId="26" pageNumber="27" start="start">tinct</pageBreakToken>
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, not pointed; males with terminal sternum unmodified, penultimate sternum weakly tumid on each side; apical lamella of aedeagus long, slightly hooked to left.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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This new species can be easily distinguished from all other species of the
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">agilis</taxonomicName>
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-group by its distinct larger size. Including the new species, a total of three known
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus)" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Circinatus</taxonomicName>
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species (the other two are
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">Pterostichus agilis</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus camelus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="camelus">Pterostichus camelus</taxonomicName>
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sp. n.) have their male secondary sexual characters on the penultimate sternum.
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus camelus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="camelus">Pterostichus camelus</taxonomicName>
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sp. n. is distinguishable from the others by its fifth tarsomeres setose, and different male penultimate sternum.
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">Pterostichus agilis</taxonomicName>
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is the species most similar to
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
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sp. n. in their similar male secondary sexual characters on penultimate sternum (weakly tumid on each side). These two species can be separated by: (1)
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
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sp. n. is much larger than
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">Pterostichus agilis</taxonomicName>
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; (2)
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
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sp. n. have the outer groove of the pronotal basal foveal present, but completely vanished in
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">Pterostichus agilis</taxonomicName>
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; (3) elytral humeral tooth almost invisible in
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
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sp. n., but sharp and distinct in
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">Pterostichus agilis</taxonomicName>
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; (4) apical lamella of aedeagus with obtuse hook in
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
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sp. n., but simple in
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="agilis">Pterostichus agilis</taxonomicName>
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.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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Among its sympatric species,
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
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sp. n. is most similar to
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus adelphus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="adelphus">Pterostichus adelphus</taxonomicName>
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sp. n. In addition to the difference in size, they also differ in elytral microsculpture: isodiametric or transverse in
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
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sp. n., very faint and linear in
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus adelphus" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="adelphus">Pterostichus adelphus</taxonomicName>
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sp. n.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
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<paragraph pageId="26" pageNumber="27">Description.</paragraph>
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<paragraph lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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Body form stout, body length 13.5-14.5 mm; dorsal side black, shining; elytron without iridescent shine; mouthparts, antenna and tarsus dark brown; ventral side blackish. Both sexes with distinct elytral microsculpture, isodiametric on disc, gradually turned to transverse on elytral apex and outer intervals. Head. Frons without punctures; antenna reaching elytron basal fifth; gena approx same length as eye, briefly tumid behind eye. Pronotum round, widest at approx anterior third, PW/PL = 1.22-1.28; usually four (sometimes three) mid-lateral setae present, first one close to anterior angle, last one near middle of lateral margin, a little distant from others; posterior seta distant from hind angle, distance between seta and hind angle approx same as distance between hind angle and inner basal foveal groove; hind angle completely rounded; basal fovea shallow, faintly defined; inner groove subparallel to median line, slightly curved outwards; outer groove faint, approx half length of inner one, outer area of inner groove distinctly concave; basal foveal area usually impunctate, sometimes with sparse punctures on inner side of inner groove. Elytron oviform, with basal ridge weakly oblique; elytral shoulder slightly narrowed, basal ridge and lateral margin forming obtuse angle, humeral tooth indistinct, not pointed; intervals feebly convex; striae moderately deep, without punctures; scutellar stria short, usually incomplete; third interval with two setigerous pores adjacent to second stria; umbilical pore series on ninth interval sparse in middle, composed of 15-18 pores (6, 1-3, 7-9). Ventral side. Proepisternum impunctate; mesepisternum finely punctate on anterior half; metepisternum impunctate or very sparsely punctate; penultimate sternum of males very faintly tumid on each side, tumidities close to primary setae, middle region between tumidities faintly depressed (Fig. 117); terminal sternum of males without special structure. Legs. Fifth tarsomeres without setae beneath; males with apical half of mesotibia slightly widened, inner margin slightly crenulate; first metatarsomere with distinct carina on outer surface, such carina on second metatarsomere superficial. Male
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<pageBreakToken pageId="27" pageNumber="28" start="start">genitalia</pageBreakToken>
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. Median lobe of male genitalia bent approx 90 degrees, apex slightly bent ventrally (Fig. 38A); ventral margin straight in middle, slightly sinuate before apex; dorsal margin gradually curved; apical orifice large, slightly turned to left side, not opened on ventral side; apical lamella long, approx half length of apical orifice, laminate, not thickened; in dorsal view, apical lamella distinctly inclined to right, nearly triangular with rounded apex, apex a little enlarged, slightly hooked to left; apical lamella length approx 1.5 times basal width (Fig. 38B). Right paramere straight and fine, length approx 3.5 times greatest width, apex rounded (Fig. 38C). Endophallus (Fig. 38 D, E, F) bent to ventral side across apical lamella; gonopore (gp, gonopore lobe folded in Fig. 38) located at level of apical lamella, pointing to aedeagal base. Four distinct lobes recognized, the three (vb-I, vb-II, va) located on ventral side of endophallus strongly chitinized: ventral-basal lobe I (vb-I) close to base of apical lamella, very small, covered by vb-II, upper surface strongly chitinized, lower surface membranous; ventral-basal lobe II (vb-II) larger than vb-I, upper surface strongly chitinized, lower surface membranous, apex of lower surface distinctly projected; ventral-apical lobe (va) close to apex of apical lamella, approx same length as vb-II, upper surface strongly chitinized, its apex strongly prolonged and sinuate, lower surface membranous; right lobe (rl) small and rounded, decorated with fine scales. Female genitalia. Spermatheca (Fig. 54) with seminal canal approx 5.5 times as long as receptaculum; receptaculum capitate, club approx half length of receptaculum; spermathecal gland slightly expanded; seminal canal inserted at base of common oviduct, base of seminal canal sclerotized. Stylomere II with two ensiform setae at outer margin, and one near middle of inner margin; two short nematiform setae located in a furrow near apex. Female sternum VIII (Fig. 75B) with dense and fine spines on posterior margin; posterior margin rounded, shallowly notched in middle; posterior region well chitinized, anterior region also, middle transparent region V-shaped, almost adjacent to posterior notch in middle. Female tergum VIII (Fig. 75A) with anterior half chitinized, posterior half semi-chitinized pigmented with sparse spots, without transparent region in middle.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="distribution">
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<paragraph pageId="27" pageNumber="28">Distribution.</paragraph>
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<paragraph pageId="27" pageNumber="28">Most specimens of this species were collected from the type locality (Yizi pass) on the border between Meigu and Ebian County (Sichuan, Liangshan Yi Autonomous Prefecture). One female, slightly different from other specimens, was collected in Yele Reserve in Mianning County. (Map 2)</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="etymology">
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<paragraph pageId="27" pageNumber="28">Etymology.</paragraph>
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<paragraph pageId="27" pageNumber="28">
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The scientific name of the new species comes from Zhygealu, an honored hero in the Yi
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<normalizedToken originalValue="people’s">people's</normalizedToken>
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legend. Yi is the local nation living in Liangshan Yi Autonomous Prefecture (Sichuan Province), where all known species of the
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="agilis">agilis</taxonomicName>
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-group are distributed. This species deserves a
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<normalizedToken originalValue="hero’s">hero's</normalizedToken>
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name as the largest-sized of all known
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="agilis">agilis</taxonomicName>
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-group species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="affinities">
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<paragraph pageId="27" pageNumber="28">Affinities.</paragraph>
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<paragraph pageId="27" pageNumber="28">
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This new species is closest to
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="agilis">Pterostichus agilis</taxonomicName>
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. Despite the difference in size, these two species are very similar in the following characters: (1) elytral microsculpture transverse or isodiametric; (2) male secondary sexual characters on the penultimate sternum; and (3) apical lamella of the aedeagus long and bent to the right. The type locality of
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="agilis">Pterostichus agilis</taxonomicName>
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is Luojishan (Sichuan), a mountain approximately 150 km southwest of the Yizi Pass, which is the type locality of
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<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
|
||
sp. n.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="28" pageNumber="29" type="habitat">
|
||
<paragraph pageId="28" pageNumber="29">
|
||
<pageBreakToken pageId="28" pageNumber="29" start="start">Habitat</pageBreakToken>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="28" pageNumber="29">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
|
||
sp. n. was collected in mixed forest with dominant large pines, and rich in dead logs. In daytime, they hide in dead logs, usually under bark. In nighttime, they come out, and can be trapped by pitfalls.
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
|
||
sp. n. lives together with three other
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus)" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Circinatus</taxonomicName>
|
||
species. Among them,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus zhygealu" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="zhygealu">Pterostichus zhygealu</taxonomicName>
|
||
sp. n. and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus adelphus" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="adelphus">Pterostichus adelphus</taxonomicName>
|
||
sp. n. are rarer than the other two species (
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus bullatus" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bullatus">Pterostichus bullatus</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus cavazzutianus" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cavazzutianus">Pterostichus cavazzutianus</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="28" pageNumber="29" type="variation">
|
||
<paragraph pageId="28" pageNumber="29">Variation.</paragraph>
|
||
<paragraph pageId="28" pageNumber="29">One female collected from Mianning County is slightly different from the holotype and the paratypes in the following characters: pronotum narrower, strongly narrowed to base, and body size small with a body length of 12.7 mm. Because of these morphological differences and a different locality (approximately 100 km NW of the type locality), we suspected that it might represent a different subspecies and excluded it from the type series.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |