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<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.30.35559" ID-GBIF-Dataset="e070f562-f13e-4194-8a42-1c196fad663e" ID-GBIF-Taxon="158498353" ID-Pensoft-Pub="1314-2615-30-57" ID-ZooBank="19205CA6468045D4A3A0F1640752D41B" LSID="urn:lsid:zoobank.org:act:5CDE1D3A-71A2-48EA-B4BB-F9FDD20D860D" ModsDocID="1314-2615-30-57" checkinTime="1565091320742" checkinUser="pensoft" docAuthor="Liu, Weixin &amp; Wynne, J. Judson" docDate="2019" docId="E483ACABE7A9F291D70E273157526E29" docLanguage="en" docName="SubterBiol 30: 57-94" docOrigin="Subterranean Biology 30" docSource="http://dx.doi.org/10.3897/subtbiol.30.35559" docTitle="Glyphiulus maocun Liu &amp; Wynne, 2019, sp. nov." docType="treatment" docUuid="5CDE1D3A-71A2-48EA-B4BB-F9FDD20D860D" docUuidSource="ZooBank" docVersion="5" lastPageId="24" lastPageNumber="81" masterDocId="FF9D4F49FFE472773E49FFCC1B62FFAC" masterDocTitle="Cave millipede diversity with the description of six new species from Guangxi, China" masterLastPageNumber="94" masterPageNumber="57" pageId="22" pageNumber="79" updateTime="1643473345400" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Cave millipede diversity with the description of six new species from Guangxi, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Weixin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wynne, J. Judson</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>30</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>57</mods:start>
<mods:end>94</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/subtbiol.30.35559</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.30.35559</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2615-30-57</mods:identifier>
<mods:identifier type="ZooBank">19205CA6468045D4A3A0F1640752D41B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="158498353" LSID="urn:lsid:zoobank.org:act:5CDE1D3A-71A2-48EA-B4BB-F9FDD20D860D" httpUri="http://treatment.plazi.org/id/E483ACABE7A9F291D70E273157526E29" lastPageId="24" lastPageNumber="81" pageId="22" pageNumber="79">
<subSubSection pageId="22" pageNumber="79" type="nomenclature">
<paragraph pageId="22" pageNumber="79">
<taxonomicName LSID="E483ACABE7A9F291D70E273157526E29" class="Diplopoda" family="Glomeridae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Glomerida" pageId="22" pageNumber="79" phylum="Arthropoda" rank="species" species="maocun">
<pageBreakToken pageId="22" pageNumber="79" start="start">Glyphiulus</pageBreakToken>
maocun
</taxonomicName>
<taxonomicNameLabel pageId="22" pageNumber="79">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. A Trichopeltis liangfengdong sp. nov. from Liangfeng Cave B Pacidesmus trifidus Golovatch &amp; Geoffroy, 2014 from Maomaotou Cave C Glyphiulus maocun sp. nov. from Liangfeng Cave." figureDoi="10.3897/subtbiol.30.35559.figure4" httpUri="https://binary.pensoft.net/fig/314552" pageId="22" pageNumber="79">Figs 4C</figureCitation>
,
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="22" pageNumber="79">18</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="22" pageNumber="79">19</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="22" pageNumber="79">20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="79" type="type material">
<paragraph pageId="22" pageNumber="79">Type material.</paragraph>
<paragraph pageId="22" pageNumber="79">
Holotype male, China, Guangxi Zhuang Autonomous Region, Lingchuan County, Maocun Village, Liangfeng Cave [
<geoCoordinate degrees="25" direction="north" minutes="12" orientation="latitude" precision="1" seconds="34.86" value="25.209684">25°12'34.86&quot;N</geoCoordinate>
,
<geoCoordinate degrees="110" direction="east" minutes="31" orientation="longitude" precision="1" seconds="56.8" value="110.53245">110°31'56.8&quot;E</geoCoordinate>
], 184 m el., deep zone, direct intuitive search, 18 November 2016, J.J. Wynne. Paratypes, 4 females, 4 juv. (SCAU), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="79" type="etymology">
<paragraph pageId="22" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="22" pageNumber="79">
The species name,
<emphasis italics="true" pageId="22" pageNumber="79">maocun</emphasis>
, is used as a noun in apposition and is the name of the village nearest to the type locality.
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79" type="diagnosis">
<paragraph pageId="22" pageNumber="79">Diagnosis.</paragraph>
<paragraph lastPageId="23" lastPageNumber="80" pageId="22" pageNumber="79">
This new species is similar to
<taxonomicName genus="G." lsidName="G. melanoporus" pageId="22" pageNumber="79" rank="species" species="melanoporus">
<emphasis italics="true" pageId="22" pageNumber="79">G. melanoporus</emphasis>
</taxonomicName>
(from caves in Guilin) in showing carinotaxy formula of collum
<normalizedToken originalValue="IIV">I-IV</normalizedToken>
+5c+6a+pc+ma. However,
<taxonomicName genus="G." lsidName="G. maocun" pageId="22" pageNumber="79" rank="species" species="maocun">
<emphasis italics="true" pageId="22" pageNumber="79">G. maocun</emphasis>
</taxonomicName>
sp. nov. is distinct from
<taxonomicName genus="G." lsidName="G. melanoporus" pageId="22" pageNumber="79" rank="species" species="melanoporus">
<emphasis italics="true" pageId="22" pageNumber="79">G. melanoporus</emphasis>
</taxonomicName>
by (1) carinotaxy formula of metaterga 2/2+I/
<pageBreakToken pageId="23" pageNumber="80" start="start">i</pageBreakToken>
+3/3; (2) anterior gonopod with a conspicuous, high, curved downward process on coxosternum (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="23" pageNumber="80">Figs 19F</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="23" pageNumber="80">20A</figureCitation>
); (3) posterior gonopod tip branch plumose flagellum (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="23" pageNumber="80">
Figs 19
<normalizedToken originalValue="GI">G-I</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="23" pageNumber="80">20B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="81" type="description">
<paragraph pageId="24" pageNumber="81">
<pageBreakToken pageId="24" pageNumber="81" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="24" pageNumber="81">
Based on the type specimens. Lengths of both sexes
<emphasis italics="true" pageId="24" pageNumber="81">ca</emphasis>
26-32 mm, mid-body rings round in cross-section, their widths and heights similar, 1.2-1.8 mm.
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Coloration</emphasis>
: yellow-brown to brownish in alcohol. Ommatidia blackish (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18A</figureCitation>
).
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Body</emphasis>
: with 46-54 podous rings + 2-1 apodous one(s) + telson.
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Head</emphasis>
: clypeus with 4 teeth anteromedially. Each eye patch with about 8-12 ommatidium arranged in three irregular vertical rows (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Figs 18A</figureCitation>
,
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">19A</figureCitation>
). Antennae long, antennomere 7 with four short apical cones (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18A, B</figureCitation>
). Gnathochilarium usually, with a separate promentum (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19B</figureCitation>
).
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Exoskeleton</emphasis>
: Carinotaxy formula of collum
<normalizedToken originalValue="IIV">I-IV</normalizedToken>
+5c+6a+pc+ma, anterior margin crests lower (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19A</figureCitation>
). Subsequent metaterga strongly crested, carinotaxy formula 2/2+I/i+3/3. Ozoporiferous tubercles round (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18</figureCitation>
). Prozonae delicately alveolate; strictures and metazonae fine longitudinal striations. Rings 2 and 3 with long pleural flaps. Epiproct simple, with a short, low, rounded tubercle medially. Paraprocts rather regularly convex. Hypoproct broadly emarginated (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18D</figureCitation>
). ♂ legs 1 very strongly reduced, represented only by a sternum devoid of any median or paramedian structures but carrying 1+1 strongly separated prongs both curved anteriad and bearing several strong setae (
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Glyphiulus maocun sp. nov., holotype. A, B Anterior part of body, sublateral views C mid-body rings, dorsal view D posterior part of body, ventral view." figureDoi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81">Fig. 18B, C</figureCitation>
). ♂ legs 2 with large coxae; penes oblong, each with two strong setae distolaterally (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19D</figureCitation>
). ♂ legs 3 with slender and elongate coxae (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Fig. 19E</figureCitation>
). Legs slender, about 1.2 times as long as mid-body ring height. Claw simple, without any spine at base.
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Anterior gonopods</emphasis>
: (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">Figs 19F</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81">20A</figureCitation>
) with a conspicuous, high, lobe-shaped, curved downward process on coxosternum, which is much higher than telopdite; telopodite slightly coiled, 1-segmented, lateral in position, with 2 or 3 strong apical setae.
<emphasis bold="true" italics="true" pageId="24" pageNumber="81">Posterior gonopods</emphasis>
: (
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Glyphiulus maocun sp. nov., holotype. A Head and collum, dorsal view B head, collum, and legs 1, ventral view C legs 1, anterior view D leg 2, caudal view E legs 3, frontal view F anterior gonopods, caudal view G posterior gonopods, anterior view H posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81">
Figs 19
<normalizedToken originalValue="GI">G-I</normalizedToken>
</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Glyphiulus maocun sp. nov., holotype. A Anterior gonopods, caudal view B posterior gonopods, anterior view C posterior gonopods, caudal view." figureDoi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81">20B, C</figureCitation>
) laterally with a low, lateral lobe carrying a dozen of long setae, tip branch very slender and high, plumose flagellum.
</paragraph>
<caption doi="10.3897/subtbiol.30.35559.figure18" httpUri="https://binary.pensoft.net/fig/314566" pageId="24" pageNumber="81" start="Figure 18" startId="F18">
<paragraph pageId="24" pageNumber="81">
Figure 18.
<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
</taxonomicName>
sp. nov., holotype.
<emphasis bold="true" pageId="24" pageNumber="81">A, B</emphasis>
Anterior part of body, sublateral views
<emphasis bold="true" pageId="24" pageNumber="81">C</emphasis>
mid-body rings, dorsal view
<emphasis bold="true" pageId="24" pageNumber="81">D</emphasis>
posterior part of body, ventral view.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.30.35559.figure19" httpUri="https://binary.pensoft.net/fig/314567" pageId="24" pageNumber="81" start="Figure 19" startId="F19">
<paragraph pageId="24" pageNumber="81">
Figure 19.
<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
</taxonomicName>
sp. nov., holotype.
<emphasis bold="true" pageId="24" pageNumber="81">A</emphasis>
Head and collum, dorsal view
<emphasis bold="true" pageId="24" pageNumber="81">B</emphasis>
head, collum, and legs 1, ventral view
<emphasis bold="true" pageId="24" pageNumber="81">C</emphasis>
legs 1, anterior view
<emphasis bold="true" pageId="24" pageNumber="81">D</emphasis>
leg 2, caudal view
<emphasis bold="true" pageId="24" pageNumber="81">E</emphasis>
legs 3, frontal view
<emphasis bold="true" pageId="24" pageNumber="81">F</emphasis>
anterior gonopods, caudal view
<emphasis bold="true" pageId="24" pageNumber="81">G</emphasis>
posterior gonopods, anterior view
<emphasis bold="true" pageId="24" pageNumber="81">H</emphasis>
posterior gonopods, caudal view.
</paragraph>
</caption>
<caption doi="10.3897/subtbiol.30.35559.figure20" httpUri="https://binary.pensoft.net/fig/314568" pageId="24" pageNumber="81" start="Figure 20" startId="F20">
<paragraph pageId="24" pageNumber="81">
Figure 20.
<taxonomicName class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus maocun" order="Spirostreptida" pageId="24" pageNumber="81" phylum="Arthropoda" rank="species" species="maocun">
<emphasis italics="true" pageId="24" pageNumber="81">Glyphiulus maocun</emphasis>
</taxonomicName>
sp. nov., holotype.
<emphasis bold="true" pageId="24" pageNumber="81">A</emphasis>
Anterior gonopods, caudal view
<emphasis bold="true" pageId="24" pageNumber="81">B</emphasis>
posterior gonopods, anterior view
<emphasis bold="true" pageId="24" pageNumber="81">C</emphasis>
posterior gonopods, caudal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="24" pageNumber="81" type="habitat">
<paragraph pageId="24" pageNumber="81">Habitat.</paragraph>
<paragraph pageId="24" pageNumber="81">Specimens were collected within or at proximity to flood detritus.</paragraph>
</subSubSection>
<subSubSection pageId="24" pageNumber="81" type="notes">
<paragraph pageId="24" pageNumber="81">Notes.</paragraph>
<paragraph pageId="24" pageNumber="81">Specimens exhibited no characteristics suggestive of cave adaptation. It had a pigmented cuticle and well-developed blackish ommatidia. We consider this species to be troglophillic within Liangfeng Cave. Subsequently, this species may have a larger, more regional distribution.</paragraph>
</subSubSection>
</treatment>
</document>