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<mods:title id="5830844142FAF8F40C0D105AB94E1C50">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species</mods:title>
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<mods:namePart id="59A12B67FB6D3EB23D340F99957F3A69">Basílio, Daniel Silva</mods:namePart>
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<mods:affiliation id="94195F23173DCB67DF668A7C63D04FEF">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; basilio. dsb @ gmail. com; https: // orcid. org / 0000 - 0003 - 3556 - 5880</mods:affiliation>
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<mods:namePart id="900DD1BE95F7556FE25B7A3E28D5AA9F">Mise, Kleber Makoto</mods:namePart>
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<mods:affiliation id="0F9D692B7FC9E565620265E58D7C4437">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; klebermise @ yahoo. com. br; https: // orcid. org / 0000 - 0003 - 4391 - 9589</mods:affiliation>
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<taxonomicName id="82EE94F3EC0F5A766BBF19FEFE58FA7D" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[151,498,1446,1473]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="779A3362EC0F5A766BBF19FEFE58FA7D" bold="true" box="[151,498,1446,1473]" pageId="10" pageNumber="11">
<emphasis id="779A3362EC0F5A766BBF19FEFEC3FA7C" bold="true" box="[151,361,1446,1473]" italics="true" pageId="10" pageNumber="11">Liogenys densata</emphasis>
<bibRefCitation id="217F9281EC0F5A766A5819FEFE58FA7D" author="Frey, G." box="[368,498,1446,1473]" pageId="10" pageNumber="11" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
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<figureCitation id="DDD5F3F5EC0F5A766BBF1992FF4CFA58" box="[151,230,1483,1509]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="10" pageNumber="11">Figs. 4</figureCitation>
;
<figureCitation id="DDD5F3F5EC0F5A766BDA1992FEBBFA58" box="[242,273,1483,1509]" captionStart="FIGURE 24" captionStartId="52.[151,250,1459,1483]" captionTargetBox="[171,1416,196,1422]" captionTargetId="figure-21@52.[151,1435,181,1434]" captionTargetPageId="52" captionText="FIGURE 24. Geographical distribution of four known Liogenys species occurring in the Chacoan province and its boundaries. Shaped points: previously known species; colors: Neotropical provinces (Morrone 2014) and Monte province from the South American Transition Zone (Roig-Juñent et al. 2018)." figureDoi="http://doi.org/10.5281/zenodo.4561326" httpUri="https://zenodo.org/record/4561326/files/figure.png" pageId="10" pageNumber="11">24</figureCitation>
.
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<taxonomicName id="82EE94F3EC0F5A766BBF1A49FE46F99A" authority="Frey, 1969: 39" authorityName="Frey" authorityPageNumber="39" authorityYear="1969" box="[151,492,1551,1576]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="densatus">
<emphasis id="779A3362EC0F5A766BBF1A49FEF8F99A" box="[151,338,1551,1575]" italics="true" pageId="10" pageNumber="11">Liogenys densatus</emphasis>
<bibRefCitation id="217F9281EC0F5A766A701A49FE46F99A" author="Frey, G." box="[344,492,1551,1576]" pageId="10" pageNumber="11" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969: 39</bibRefCitation>
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<taxonomicName id="82EE94F3EC0F5A766BBF1A77FDABF9FB" authority=": Evans 2003: 207" authorityName="Evans" authorityPageNumber="207" authorityYear="2003" box="[151,513,1582,1606]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="779A3362EC0F5A766BBF1A77FEE2F9FB" box="[151,328,1582,1606]" italics="true" pageId="10" pageNumber="11">Liogenys densata</emphasis>
:
<bibRefCitation id="217F9281EC0F5A766A7E1A77FDABF9FB" author="Evans, A. V." box="[342,513,1582,1606]" pageId="10" pageNumber="11" pagination="1 - 458" refId="ref35927" refString="Evans, A. V. (2003) A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 207</bibRefCitation>
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<bibRefCitation id="217F9281EC0F5A7669A81A77FC22F9FB" author="Evans, A. V. &amp; Smith, A. B. T." box="[640,904,1582,1606]" pageId="10" pageNumber="11" refId="ref35974" refString="Evans, A. V. &amp; Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolon- thinae). Version 3. Available from: http: // unsm-ento. unl. edu / SSSA / nwmelos. htm (accessed 4 January 2021)" type="url" year="2009">Evans &amp; Smith 2009: 176</bibRefCitation>
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(checklist);
<treatmentCitation id="C44FC961EC0F5A766F2F1A77FB6AF9FB" author="Krajcik, M." box="[1031,1216,1582,1606]" page="144" pageId="10" pageNumber="11" year="2012">
<bibRefCitation id="217F9281EC0F5A766F2F1A77FB6AF9FB" author="Krajcik, M." box="[1031,1216,1582,1606]" pageId="10" pageNumber="11" refId="ref36405" refString="Krajcik, M. (2012) Checklist of the world Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp." type="book" year="2012">Krajčík 2012: 144</bibRefCitation>
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(checklist);
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<bibRefCitation id="217F9281EC0F5A766E171A77FEC2F9D9" author="Cherman, M. A. &amp; Moron, M. A. &amp; Almeida, L. M." pageId="10" pageNumber="11" pagination="744 - 770" refId="ref35581" refString="Cherman, M. A., Moron, M. A. &amp; Almeida, L. M. (2016) Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology, 41, 744 - 770. https: // doi. org / 10.1111 / syen. 12188" type="journal article" year="2016">
Cherman
<emphasis id="779A3362EC0F5A766BEF1A14FF50F9D9" box="[199,250,1612,1636]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2016: 764
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, 766 (systematics);
<treatmentCitation id="C44FC961EC0F5A76691A1A15FC83F9D8" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[562,809,1612,1637]" page="10" pageId="10" pageNumber="11" year="2017">
<bibRefCitation id="217F9281EC0F5A76691A1A15FC83F9D8" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[562,809,1612,1637]" pageId="10" pageNumber="11" pagination="1 - 120" refId="ref35643" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman
<emphasis id="779A3362EC0F5A7669BD1A14FD63F9D9" box="[661,713,1612,1636]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2017: 10
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, 16 (redescription).
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<emphasis id="779A3362EC0F5A766BBF1ACAFEEAF911" bold="true" box="[151,320,1682,1709]" pageId="10" pageNumber="11">Type material</emphasis>
.
<taxonomicName id="82EE94F3EC0F5A766A671ACAFDB5F910" authorityName="Frey" authorityYear="1969" box="[335,543,1683,1709]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="densatus">
<emphasis id="779A3362EC0F5A766A671ACAFDB5F910" box="[335,543,1683,1709]" italics="true" pageId="10" pageNumber="11">Liogenys densatus</emphasis>
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female
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(
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): [white typeset] “
<collectingCountry id="3DF9AFE0EC0F5A766F321ACAFB68F910" box="[1050,1218,1683,1709]" name="Argentina" pageId="10" pageNumber="11">ARGENTINE</collectingCountry>
: /
<collectingRegion id="872A2192EC0F5A766FF71ACAFA22F910" box="[1247,1416,1683,1709]" country="Argentina" name="La Rioja" pageId="10" pageNumber="11">La Rioja Prov.</collectingRegion>
, / Patquia, /
<collectorName id="E81B8AA6EC0F5A766A231AEEFE05F96C" box="[267,431,1719,1745]" pageId="10" pageNumber="11">K.J. Hayward.</collectorName>
/ B. M. 1933-333”, [red typeset] “TYPE”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC0F5A766E091AEEFA26F96C" authorityName="Guerin-Meneville" authorityYear="1831" box="[1313,1420,1719,1745]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ densatus / nov. sp. / Type / det.
<determinerName id="B08D03B4EC0F5A766ADF1A82FDE4F948" box="[503,590,1755,1781]" pageId="10" pageNumber="11">G. Frey</determinerName>
, 1968”.
</materialsCitation>
<materialsCitation id="F586E52DEC0F5A7669841A82FB9BF880" ID-GBIF-Occurrence="3047558317" collectingDate="1964-12-13" collectionCode="NHMB, USNM" collectorName="A. Martinez" country="Argentina" determinerName="G. Frey" location="One" pageId="10" pageNumber="11" specimenCount="958" stateProvince="La Rioja" typeStatus="paratype">
<typeStatus id="9A5551D2EC0F5A7669841A82FCB3F948" box="[684,793,1755,1781]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
(2):
<location id="4031B9ABEC0F5A7668781A82FCD5F948" LSID="urn:lsid:plazi:treatment:CD475E66EC0F5A706BBF19FEFDA3FB78:4031B9ABEC0F5A7668781A82FCD5F948" box="[848,895,1755,1781]" country="Argentina" name="One" pageId="10" pageNumber="11" stateProvince="La Rioja">One</location>
male
<typeStatus id="9A5551D2EC0F5A7668EB1A82FB89F948" box="[963,1059,1755,1781]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
(
<collectionCode id="23FF77B5EC0F5A766F1A1A82FB27F948" box="[1074,1165,1755,1781]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="10" pageNumber="11">NHMB</collectionCode>
): [white, outlined black, typeset] “Argentina / Pcia. San Juan / Marayes
<date id="3150C9B0EC0F5A76698E1AA6FC85F8A4" box="[678,815,1791,1817]" pageId="10" pageNumber="11" value="1964-12-13">
<collectingDate id="21143058EC0F5A76698E1AA6FC85F8A4" box="[678,815,1791,1817]" pageId="10" pageNumber="11" value="1964-12-13">13.XII.1964</collectingDate>
</date>
/
<collectorName id="E81B8AA6EC0F5A76686A1AA6FC63F8A4" box="[834,969,1791,1817]" pageId="10" pageNumber="11">A. Martinez</collectorName>
leg.”, [red handwritten] “
<collectorName id="E81B8AA6EC0F5A766FC01AA6FAEAF8A4" box="[1256,1344,1791,1817]" pageId="10" pageNumber="11">P. Type</collectorName>
”, [white handwritten] “
<taxonomicName id="82EE94F3EC0F5A766A111B7AFE0EF880" authorityName="Guerin-Meneville" authorityYear="1831" box="[313,420,1827,1853]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
♂opacicollis / Fairm / [typeset] det.
<determinerName id="B08D03B4EC0F5A7668671B7AFC0DF880" box="[847,935,1827,1853]" pageId="10" pageNumber="11">G. Frey</determinerName>
1967 / 68”.
</materialsCitation>
<materialsCitation id="F586E52DEC0F5A766F121B7AFCC8F814" ID-GBIF-Occurrence="3047558377" collectingDate="1964-12-13" collectionCode="NHMB, USNM" collectorName="P. Kohla" country="Argentina" determinerName="G. Frey" location="One" pageId="10" pageNumber="11" specimenCount="958" stateProvince="La Rioja" typeStatus="paratype">
One female
<typeStatus id="9A5551D2EC0F5A766FEF1B7AFA8DF880" box="[1223,1319,1827,1853]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
(
<collectionCode id="23FF77B5EC0F5A766E111B7AFA3BF880" box="[1337,1425,1827,1853]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
): [white handwritten] “
<collectingRegion id="872A2192EC0F5A766AA11B1EFE45F8DC" box="[393,495,1863,1889]" country="Argentina" name="La Rioja" pageId="10" pageNumber="11">La Rioja</collectingRegion>
/ Capital / XI.
<specimenCount id="53E824F9EC0F5A7669BC1B1FFD76F8DC" box="[660,732,1862,1889]" pageId="10" pageNumber="11" type="generic">958 P.</specimenCount>
Kohla”, [red, typeset and handwritten] “P. TYPE”, [light red, typeset and handwritten] “Type No / 74008 / USNM”, [white, typeset and handwritten] “
<taxonomicName id="82EE94F3EC0F5A766F451B32FB72F838" authorityName="Guerin-Meneville" authorityYear="1831" box="[1133,1240,1899,1925]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ densatus / n. sp. / P.type / det.
<determinerName id="B08D03B4EC0F5A766A181BD6FED5F814" box="[304,383,1935,1961]" pageId="10" pageNumber="11">G.Frey</determinerName>
, 1968”, [pink typeset] “Property / USNM”
</materialsCitation>
.
</paragraph>
<paragraph id="4551EF70EC0F5A706BEF1BEAFD23FEA0" blockId="10.[151,1437,1682,2033]" lastBlockId="12.[151,1436,150,1221]" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<emphasis id="779A3362EC0F5A766BEF1BEAFE0CF871" bold="true" box="[199,422,1970,1997]" pageId="10" pageNumber="11">Non-type material</emphasis>
(12).
<materialsCitation id="F586E52DEC0F5A766ADA1BEAFE1CF84C" ID-GBIF-Occurrence="3047558408" collectingDate="1939-12" collectionCode="MZUC" collectorName="Bosq" country="Argentina" location="Santiago del Estero" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" stateProvince="Santiago del Estero">
<collectingCountry id="3DF9AFE0EC0F5A766ADA1BEAFD33F870" box="[498,665,1971,1997]" name="Argentina" pageId="10" pageNumber="11">ARGENTINA</collectingCountry>
.
<collectingRegion id="872A2192EC0F5A76698F1BEAFC24F870" box="[679,910,1971,1997]" country="Argentina" name="Santiago del Estero" pageId="10" pageNumber="11">Santiago del Estero</collectingRegion>
:
<collectingRegion id="872A2192EC0F5A7668B41BEAFAA9F870" box="[924,1283,1971,1997]" country="Argentina" name="Santiago del Estero" pageId="10" pageNumber="11">Ciudad de Santiago del Estero</collectingRegion>
,
<date id="3150C9B0EC0F5A766E3A1BEAFAD0F870" box="[1298,1402,1971,1997]" pageId="10" pageNumber="11" value="1939-12">
<collectingDate id="21143058EC0F5A766E3A1BEAFAD0F870" box="[1298,1402,1971,1997]" pageId="10" pageNumber="11" value="1939-12">XII.1939</collectingDate>
</date>
, J.
<collectorName id="E81B8AA6EC0F5A766BBF1B8EFF79F84C" box="[151,211,2007,2033]" pageId="10" pageNumber="11">Bosq</collectorName>
,
<specimenCount id="53E824F9EC0F5A766BCA1B8EFEEFF84C" box="[226,325,2007,2033]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
(
<collectionCode id="23FF77B5EC0F5A766A7F1B8EFE04F84C" box="[343,430,2007,2033]" country="Chile" name="Museo de Zoologia, Universidad de Concepcion" pageId="10" pageNumber="11">MZUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F586E52DEC0F5A706AEC1B8EFCD1FF0C" ID-GBIF-Occurrence="3047558483" collectingDate="2005-12-16" collectionCode="IADIZA" collectorName="Lavalle" country="Argentina" lastPageId="12" lastPageNumber="13" latitude="-32.383213" location="Mendoza" longLatPrecision="1" longitude="-68.05394" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" specimenCount-male="1" stateProvince="Mendoza">
<collectingRegion id="872A2192EC0F5A766AEC1B8EFD9BF84C" box="[452,561,2007,2033]" country="Argentina" name="Mendoza" pageId="10" pageNumber="11">Mendoza</collectingRegion>
:
<collectorName id="E81B8AA6EC0F5A7669161B8EFD3FF84C" box="[574,661,2007,2033]" pageId="10" pageNumber="11">Lavalle</collectorName>
, Telteca,
<date id="3150C9B0EC0F5A76682C1B8FFC25F84C" box="[772,911,2006,2033]" pageId="10" pageNumber="11" value="2005-12-16">
<collectingDate id="21143058EC0F5A76682C1B8FFC25F84C" box="[772,911,2006,2033]" pageId="10" pageNumber="11" value="2005-12-16">16.XII.2005</collectingDate>
</date>
,
<collectorName id="E81B8AA6EC0F5A7668B51B8EFBADF84C" box="[925,1031,2007,2033]" pageId="10" pageNumber="11">Marvaldi</collectorName>
,
<collectorName id="E81B8AA6EC0F5A766F3D1B8EFBE7F84C" box="[1045,1101,2007,2033]" pageId="10" pageNumber="11">Ruiz</collectorName>
,
<collectorName id="E81B8AA6EC0F5A766F721B8EFB09F84C" box="[1114,1187,2007,2033]" pageId="10" pageNumber="11">Flores</collectorName>
, mercury vapor light,
<specimenCount id="53E824F9EC095A706BBF1CCEFF4FFF0C" box="[151,229,151,177]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
and
<specimenCount id="53E824F9EC095A706A091CCEFE2FFF0C" box="[289,389,151,177]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectionCode id="23FF77B5EC095A706ABF1CCEFE54FF0C" box="[407,510,151,177]" pageId="12" pageNumber="13">IADIZA</collectionCode>
);
<geoCoordinate id="20DA89B7EC095A70693C1CCFFD17FF0C" box="[532,701,150,177]" degrees="32" direction="south" minutes="22" orientation="latitude" pageId="12" pageNumber="13" precision="1" seconds="59.58" value="-32.383213">32º22’59.58’’S</geoCoordinate>
<geoCoordinate id="20DA89B7EC095A7069EF1CCEFCD1FF0C" box="[711,891,151,177]" degrees="68" direction="west" minutes="03" orientation="longitude" pageId="12" pageNumber="13" precision="1" seconds="14.16" value="-68.05394">68º03’14.16’’W</geoCoordinate>
</materialsCitation>
,
<materialsCitation id="F586E52DEC095A7068A11CCFFD5FFF68" ID-GBIF-Occurrence="3047558446" collectingDate="2008-02-05" collectionCode="IADIZA" collectorName="Ocampo &amp; Ruiz &amp; San Blas" country="Argentina" location="Reserva de Telteca" pageId="12" pageNumber="13" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Mendoza">
<date id="3150C9B0EC095A7068A11CCFFC59FF0C" box="[905,1011,150,177]" pageId="12" pageNumber="13" value="2008-02-05">
<collectingDate id="21143058EC095A7068A11CCFFC59FF0C" box="[905,1011,150,177]" pageId="12" pageNumber="13" value="2008-02-05">
5.II.
<quantity id="82164295EC095A70689B1CCFFC59FF0C" box="[947,1011,150,177]" metricMagnitude="6" metricUnit="m" metricValue="2.008548" pageId="12" pageNumber="13" unit="m" value="2008548.0">2008</quantity>
</collectingDate>
</date>
, 548 m,
<collectorName id="E81B8AA6EC095A706F731CCEFB17FF0C" box="[1115,1213,151,177]" pageId="12" pageNumber="13">Ocampo</collectorName>
,
<collectorName id="E81B8AA6EC095A706FE21CCEFAA8FF0C" box="[1226,1282,151,177]" pageId="12" pageNumber="13">Ruiz</collectorName>
, &amp;
<collectorName id="E81B8AA6EC095A706E181CCEFA3DFF0C" box="[1328,1431,151,177]" pageId="12" pageNumber="13">San Blas</collectorName>
, mercury vapor light,
<specimenCount id="53E824F9EC095A706AA81CE2FE7FFF68" box="[384,469,187,213]" pageId="12" pageNumber="13" type="male">2 males</specimenCount>
and
<specimenCount id="53E824F9EC095A7069221CE2FDDFFF68" box="[522,629,187,213]" pageId="12" pageNumber="13" type="female">2 females</specimenCount>
(
<collectionCode id="23FF77B5EC095A7069AA1CE2FD43FF68" box="[642,745,187,213]" pageId="12" pageNumber="13">IADIZA</collectionCode>
);
</materialsCitation>
<materialsCitation id="F586E52DEC095A7069D31CE2FD0BFF44" ID-GBIF-Occurrence="3047558466" collectingDate="2008-02-05" collectingDateMax="2002-12-10" collectingDateMin="2002-12-08" collectionCode="IADIZA" collectorName="V.M. Diegues" country="Argentina" latitude="-34.0494" location="Reserva de Telteca" longLatPrecision="6" longitude="-67.9668" pageId="12" pageNumber="13" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Mendoza">
<location id="4031B9ABEC095A7069D31CE2FC64FF68" LSID="urn:lsid:plazi:treatment:CD475E66EC0F5A706BBF19FEFDA3FB78:4031B9ABEC095A7069D31CE2FC64FF68" box="[763,974,187,213]" country="Argentina" name="Reserva de Telteca" pageId="12" pageNumber="13" stateProvince="Mendoza">Reserva de Telteca</location>
,
<date id="3150C9B0EC095A7068F01CE2FB2AFF68" box="[984,1152,187,213]" pageId="12" pageNumber="13" value="2002-12-08" valueMax="2002-12-10" valueMin="2002-12-08">
<collectingDate id="21143058EC095A7068F01CE2FB2AFF68" box="[984,1152,187,213]" pageId="12" pageNumber="13" value="2002-12-08" valueMax="2002-12-10" valueMin="2002-12-08">8–10.XII.2002</collectingDate>
</date>
,
<collectorName id="E81B8AA6EC095A706FA21CE2FA8DFF68" box="[1162,1319,187,213]" pageId="12" pageNumber="13">V.M. Diéguez</collectorName>
,
<specimenCount id="53E824F9EC095A706E191CE2FA36FF68" box="[1329,1436,187,213]" pageId="12" pageNumber="13" type="female">3 females</specimenCount>
(
<collectionCode id="23FF77B5EC095A706BB71C86FF53FF44" box="[159,249,223,249]" pageId="12" pageNumber="13">CMVD</collectionCode>
),
<location id="4031B9ABEC095A706A241CB9FE2AFF46" LSID="urn:lsid:plazi:treatment:CD475E66EC0F5A706BBF19FEFDA3FB78:4031B9ABEC095A706A241CB9FE2AFF46" box="[268,384,224,251]" country="Argentina" latitude="-34.0494" longLatPrecision="6" longitude="-67.9668" name="Nancunan" pageId="12" pageNumber="13" stateProvince="Mendoza">Ñancuñan</location>
, [
<geoCoordinate id="20DA89B7EC095A706ABB1C86FDA6FF44" box="[403,524,223,249]" degrees="34.0494" direction="south" orientation="latitude" pageId="12" pageNumber="13" precision="5" value="-34.0494">34.0494ºS</geoCoordinate>
,
<geoCoordinate id="20DA89B7EC095A70693F1C87FD31FF44" box="[535,667,222,249]" degrees="67.9668" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="5" value="-67.9668">67.9668ºW</geoCoordinate>
]
</materialsCitation>
,
<materialsCitation id="F586E52DEC095A7069851C87FF15FEA0" ID-GBIF-Occurrence="3047558410" collectingDate="1975-12" collectionCode="IADIZA" collectorName="Roig" country="Argentina" location="San Rafael" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" stateProvince="Mendoza">
<date id="3150C9B0EC095A7069851C87FCBCFF44" box="[685,790,222,249]" pageId="12" pageNumber="13" value="1975-12">
<collectingDate id="21143058EC095A7069851C87FCBCFF44" box="[685,790,222,249]" pageId="12" pageNumber="13" value="1975-12">XII.1975</collectingDate>
</date>
, S.
<collectorName id="E81B8AA6EC095A7068681C86FCD3FF44" box="[832,889,223,249]" pageId="12" pageNumber="13">Roig</collectorName>
,
<specimenCount id="53E824F9EC095A7068AD1C86FC7AFF44" box="[901,976,223,249]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(
<collectionCode id="23FF77B5EC095A7068C81C86FBEDFF44" box="[992,1095,223,249]" pageId="12" pageNumber="13">IADIZA</collectionCode>
);
<location id="4031B9ABEC095A706F721C86FB72FF44" LSID="urn:lsid:plazi:treatment:CD475E66EC0F5A706BBF19FEFDA3FB78:4031B9ABEC095A706F721C86FB72FF44" box="[1114,1240,223,249]" country="Argentina" name="San Rafael" pageId="12" pageNumber="13" stateProvince="Mendoza">San Rafael</location>
,
<locationDeviation id="54E3F652EC095A706FCC1C86FACFFF44" box="[1252,1381,223,249]" location="Pedro" pageId="12" pageNumber="13">
<quantity id="82164295EC095A706FCC1C86FAB7FF44" box="[1252,1309,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="km" value="2.0">2 km</quantity>
Pedro
</locationDeviation>
Vargas
</materialsCitation>
,
<materialsCitation id="F586E52DEC095A706BE31D5AFD2FFEA0" ID-GBIF-Occurrence="3047558419" box="[203,645,259,285]" collectingDate="1979-01-31" collectionCode="IADIZA" collectorName="Roig" country="Argentina" pageId="12" pageNumber="13" specimenCount="1" specimenCount-female="1" stateProvince="Mendoza">
<date id="3150C9B0EC095A706BE31D5AFE92FEA0" box="[203,312,259,285]" pageId="12" pageNumber="13" value="1979-01-31">
<collectingDate id="21143058EC095A706BE31D5AFE92FEA0" box="[203,312,259,285]" pageId="12" pageNumber="13" value="1979-01-31">31.I.1979</collectingDate>
</date>
, S.
<collectorName id="E81B8AA6EC095A706A491D5AFE30FEA0" box="[353,410,259,285]" pageId="12" pageNumber="13">Roig</collectorName>
,
<specimenCount id="53E824F9EC095A706A8E1D5AFDADFEA0" box="[422,519,259,285]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(
<collectionCode id="23FF77B5EC095A70693E1D5AFDD7FEA0" box="[534,637,259,285]" pageId="12" pageNumber="13">IADIZA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="1191BFF8EC0E5A776BBF1BCEFD27F869" ID-DOI="http://doi.org/10.5281/zenodo.4561270" ID-Zenodo-Dep="4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="11">
<paragraph id="4551EF70EC0E5A776BBF1BCEFD27F869" blockId="11.[151,1437,1943,2004]" pageId="11" pageNumber="12">
<emphasis id="779A3362EC0E5A776BBF1BCEFEB9F812" bold="true" box="[151,275,1943,1967]" pageId="11" pageNumber="12">FIGURE 4.</emphasis>
<taxonomicName id="82EE94F3EC0E5A776A321BC1FD97F80D" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[282,573,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="779A3362EC0E5A776A321BC1FE60F80D" box="[282,458,1944,1968]" italics="true" pageId="11" pageNumber="12">Liogenys densata</emphasis>
<bibRefCitation id="217F9281EC0E5A776AF91BC1FD97F80D" author="Frey, G." box="[465,573,1944,1968]" pageId="11" pageNumber="12" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral.
</paragraph>
</caption>
<subSubSection id="0DF4BCFBEC095A706BEF1D7FFD82FD4C" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="4551EF70EC095A706BEF1D7FFD82FD4C" blockId="12.[151,1436,150,1221]" pageId="12" pageNumber="13">
<emphasis id="779A3362EC095A706BEF1D7FFE90FEFD" bold="true" box="[199,314,294,320]" pageId="12" pageNumber="13">Diagnosis</emphasis>
. Length:
<quantity id="82164295EC095A706A8F1D7EFDE6FEFC" box="[423,588,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" metricValueMax="1.39" metricValueMin="1.19" pageId="12" pageNumber="13" unit="mm" value="12.9" valueMax="13.9" valueMin="11.9">11.9–13.9 mm</quantity>
; width: 6.0–
<quantity id="82164295EC095A7069F01D7EFC9AFEFC" box="[728,816,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="12" pageNumber="13" unit="mm" value="6.9">6.9 mm</quantity>
. Body and elytra purplish brown, elongate, shiny, sides parallel; pronotum slightly darker than elytra; elytra glabrous (
<figureCitation id="DDD5F3F5EC095A70685B1D12FC78FED8" box="[883,978,331,357]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="12" pageNumber="13">Fig. 4A</figureCitation>
); head and pronotum densely punctate; clypeal emargination deep, rounded and narrow; outer sides of anterior teeth subparallel; clypeal lateral margin strongly convex, with a weak, blunt projection, obtuse angle between this projection and the anterior teeth; antennae with 10 antennomeres; pronotal posterior corners sharp, in obtuse angle (
<figureCitation id="DDD5F3F5EC095A7068D21DEEFBF9FE6C" box="[1018,1107,439,465]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="12" pageNumber="13">Fig. 4B</figureCitation>
); hypomere scaly abundantly and with long bristles, mesepisternum, sides of metaventrite, metacoxae and sides of ventrites scaly abundantly; protibial teeth equally spaced; mesotibia cylindrical in cross section, in males longer; two transverse carinae posteriorly, the apical complete (
<figureCitation id="DDD5F3F5EC095A706AE41E7AFDEEFD80" box="[460,580,547,573]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="12" pageNumber="13">Fig. 4C, E</figureCitation>
); pygidium flat; subquadrate; pygidial disc with abundant scales throughout except on the apex, with long bristles (
<figureCitation id="DDD5F3F5EC095A7069171E1EFD33FDDC" box="[575,665,583,609]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="12" pageNumber="13">Fig. 4D</figureCitation>
); in males, inner margin of metatibia weakly carinate, protarsi weakly enlarged, mesotarsi not enlarged; basal region of parameres punctate, slightly narrower than both sections of the parameres at the transverse midline, parameral split at 1/3; inner margins strongly concave on inner margin, opened; apex harpoon shaped, angles very sharp, parameres four times longer than apex (
<figureCitation id="DDD5F3F5EC095A706F331EEAFBDAFD70" box="[1051,1136,691,717]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="12" pageNumber="13">Fig. 4F</figureCitation>
); parameres in lateral view slightly convex, coplanar (
<figureCitation id="DDD5F3F5EC095A706AE91E8EFDB6FD4C" box="[449,540,727,753]" captionStart="FIGURE 4" captionStartId="11.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@11.[203,1385,181,1920]" captionTargetPageId="11" captionText="FIGURE 4. Liogenys densata Frey, 1969 male views: A) dorsal, B) head and pronotum, C) lateral, D) pygidium, E) metatibia, and parameres in views: F) dorsal, and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561270" httpUri="https://zenodo.org/record/4561270/files/figure.png" pageId="12" pageNumber="13">Fig. 4G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC095A706BEF1EA2FAB8FC84" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="4551EF70EC095A706BEF1EA2FBE5FCA8" blockId="12.[151,1436,150,1221]" box="[199,1103,762,789]" pageId="12" pageNumber="13">
<materialsCitation id="F586E52DEC095A706BEF1EA2FBE1FCA8" ID-GBIF-Occurrence="3047558481" box="[199,1099,762,789]" country="Argentina" latitude="-30.043554" location="Patquia" longLatPrecision="1" longitude="-66.883995" pageId="12" pageNumber="13" specimenCount="1" stateProvince="La Rioja" typeStatus="holotype">
<emphasis id="779A3362EC095A706BEF1EA2FECAFCA9" bold="true" box="[199,352,762,789]" pageId="12" pageNumber="13">
<typeStatus id="9A5551D2EC095A706BEF1EA2FEA8FCA8" box="[199,258,763,789]" pageId="12" pageNumber="13">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="3DF9AFE0EC095A706A441EA2FDB8FCA8" box="[364,530,763,789]" name="Argentina" pageId="12" pageNumber="13">ARGENTINA</collectingCountry>
,
<collectingRegion id="872A2192EC095A7069361EA2FD28FCA8" box="[542,642,763,789]" country="Argentina" name="La Rioja" pageId="12" pageNumber="13">La Rioja</collectingRegion>
,
<location id="4031B9ABEC095A7069A51EA2FD4BFCA8" LSID="urn:lsid:plazi:treatment:CD475E66EC0F5A706BBF19FEFDA3FB78:4031B9ABEC095A7069A51EA2FD4BFCA8" box="[653,737,763,789]" country="Argentina" latitude="-30.043554" longLatPrecision="1" longitude="-66.883995" name="Patquia" pageId="12" pageNumber="13" stateProvince="La Rioja">Patquía</location>
[
<geoCoordinate id="20DA89B7EC095A7069C71EA2FC25FCA8" box="[751,911,763,789]" degrees="30" direction="south" minutes="02" orientation="latitude" pageId="12" pageNumber="13" precision="1" seconds="36.8" value="-30.043554">30°02’36.8’’S</geoCoordinate>
<geoCoordinate id="20DA89B7EC095A7068BE1EA3FBEEFCA8" box="[918,1092,762,789]" degrees="66" direction="west" minutes="53" orientation="longitude" pageId="12" pageNumber="13" precision="1" seconds="02.4" value="-66.883995">66°53’02.4’’W</geoCoordinate>
]
</materialsCitation>
.
</paragraph>
<paragraph id="4551EF70EC095A706BEF1F47FAB8FC84" blockId="12.[151,1436,150,1221]" box="[199,1298,798,825]" pageId="12" pageNumber="13">
<emphasis id="779A3362EC095A706BEF1F47FE54FC85" bold="true" box="[199,510,798,824]" pageId="12" pageNumber="13">Geographical distribution</emphasis>
.
<materialsCitation id="F586E52DEC095A7069221F46FAA6FC84" ID-GBIF-Occurrence="3047558441" box="[522,1292,798,825]" country="Argentina" location="Santiago del Estero" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Santiago del Estero">
<collectingCountry id="3DF9AFE0EC095A7069221F46FD06FC84" box="[522,684,799,825]" name="Argentina" pageId="12" pageNumber="13">ARGENTINA</collectingCountry>
(
<emphasis id="779A3362EC095A7069941F47FC00FC84" bold="true" box="[700,938,798,825]" pageId="12" pageNumber="13">
<collectingRegion id="872A2192EC095A7069941F47FC0FFC84" box="[700,933,798,825]" country="Argentina" name="Santiago del Estero" pageId="12" pageNumber="13">Santiago del Estero</collectingRegion>
,
</emphasis>
<collectingRegion id="872A2192EC095A7068991F46FBBCFC84" box="[945,1046,799,825]" country="Argentina" name="La Rioja" pageId="12" pageNumber="13">La Rioja</collectingRegion>
, San Juan,
<emphasis id="779A3362EC095A706FBB1F46FAABFC84" bold="true" box="[1171,1281,799,825]" pageId="12" pageNumber="13">
<collectingRegion id="872A2192EC095A706FBB1F46FAABFC84" box="[1171,1281,799,825]" country="Argentina" name="Mendoza" pageId="12" pageNumber="13">Mendoza</collectingRegion>
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0DF4BCFBEC095A706BEF1F1AFDA3FB78" pageId="12" pageNumber="13" type="discussion">
<paragraph id="4551EF70EC095A706BEF1F1AFDA3FB78" blockId="12.[151,1436,150,1221]" pageId="12" pageNumber="13">
<emphasis id="779A3362EC095A706BEF1F1AFE91FCE0" bold="true" box="[199,315,835,861]" pageId="12" pageNumber="13">Remarks.</emphasis>
<taxonomicName id="82EE94F3EC095A706A691F1AFDAEFCE0" authorityName="Frey" authorityYear="1969" box="[321,516,835,861]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="779A3362EC095A706A691F1AFDAEFCE0" box="[321,516,835,861]" italics="true" pageId="12" pageNumber="13">Liogenys densata</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="82EE94F3EC095A7069BB1F1AFC81FCE0" authorityName="Fairmaire" authorityYear="1892" box="[659,811,835,861]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="opacicollis">
<emphasis id="779A3362EC095A7069BB1F1AFC81FCE0" box="[659,811,835,861]" italics="true" pageId="12" pageNumber="13">L. opacicollis</emphasis>
</taxonomicName>
(
<figureCitation id="DDD5F3F5EC095A7068121F1AFC20FCE3" box="[826,906,835,862]" captionStart="FIGURE 16" captionStartId="37.[151,250,1904,1928]" captionTargetBox="[216,1372,181,1880]" captionTargetId="figure-20@37.[216,1372,181,1880]" captionTargetPageId="37" captionText="FIGURE 16. Liogenys opacicollis Fairmaire, 1892 female lectotype views: A) dorsal, C) labels; L. densicollis Moser, 1921 male lectotype views: B) dorsal, D) labels, E) lateral, F) mesotibia and metatibia, white arrow indicates shape of metatibial margin, G) mesotarsomere I, arrow indicates modified inner margin and H) its variation, I) apex of metatibia, white arrow indicates projection below the spurs, J) pygidium, and parameres in views: K) dorsal and L) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561304" httpUri="https://zenodo.org/record/4561304/files/figure.png" pageId="12" pageNumber="13">Fig. 16</figureCitation>
) in the body color; the shape of the clypeus and in the head and pronotum densely punctate. This species differs from
<taxonomicName id="82EE94F3EC095A7068861F3EFBE2FC3C" authorityName="Fairmaire" authorityYear="1892" box="[942,1096,871,897]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="opacicollis">
<emphasis id="779A3362EC095A7068861F3EFBE2FC3C" box="[942,1096,871,897]" italics="true" pageId="12" pageNumber="13">L. opacicollis</emphasis>
</taxonomicName>
(in parenthesis) mainly in the punctures of pronotum finer (cribrose-like); the pronotum and elytra glabrous (with bristles), shinier; in the shape of tibiae; and in the parameres.
<taxonomicName id="82EE94F3EC095A70692C1FF6FD63FC74" authorityName="Frey" authorityYear="1969" box="[516,713,943,969]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="779A3362EC095A70692C1FF6FD63FC74" box="[516,713,943,969]" italics="true" pageId="12" pageNumber="13">Liogenys densata</emphasis>
</taxonomicName>
also resembles
<taxonomicName id="82EE94F3EC095A7068A91FF6FB14FC74" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[897,1214,942,969]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="779A3362EC095A7068A91FF6FC50FC74" box="[897,1018,943,969]" italics="true" pageId="12" pageNumber="13">L. obscura</emphasis>
<bibRefCitation id="217F9281EC095A706F2B1FF6FB14FC74" author="Blanchard, E." box="[1027,1214,942,969]" pageId="12" pageNumber="13" pagination="129 - 240" refId="ref35099" refString="Blanchard, E. (1851) s. n. In: Museum D'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Part 2. Gide et Baudry, Paris, pp. 129 - 240." type="book chapter" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
(
<figureCitation id="DDD5F3F5EC095A706FE71FF6FA8EFC74" box="[1231,1316,942,969]" captionStart="FIGURE 15" captionStartId="34.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-21@34.[203,1385,181,1920]" captionTargetPageId="34" captionText="FIGURE 15. Liogenys obscura Blanchard, 1851 female lectotype views: A) dorsal, C) labels; L. bruchi Moser, 1921 male lectotype views: B) dorsal, D) labels, E) head and pronotum, F) lateral, G) pygidium, and parameres in views: H) dorsal, I) dorsolateral and J) lateral." figureDoi="http://doi.org/10.5281/zenodo.4561302" httpUri="https://zenodo.org/record/4561302/files/figure.png" pageId="12" pageNumber="13">Fig. 15</figureCitation>
), differing mainly in the shape of the pygidium, as wide as it is long and covered by scales abundantly (wider than it is long and also setose, but the integument is seen below the bristles).
<bibRefCitation id="217F9281EC095A70686F1FAEFC60FBAF" author="Frey, G." box="[839,970,1015,1042]" pageId="12" pageNumber="13" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
indicated that the male
<typeStatus id="9A5551D2EC095A706FF81FAEFA9AFBAC" box="[1232,1328,1015,1041]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
from San Juan is at MZSP, but we found it at NHMB instead.
<bibRefCitation id="217F9281EC095A7069F41842FCCBFB8B" author="Frey, G." box="[732,865,1051,1078]" pageId="12" pageNumber="13" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
mentioned that he received a male specimen under the name of
<taxonomicName id="82EE94F3EC095A706A0A1866FE10FBE4" authorityName="Fairmaire" authorityYear="1892" box="[290,442,1087,1113]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="opacicollis">
<emphasis id="779A3362EC095A706A0A1866FE10FBE4" box="[290,442,1087,1113]" italics="true" pageId="12" pageNumber="13">L. opacicollis</emphasis>
</taxonomicName>
from The Natural History Museum (London,
<collectingCountry id="3DF9AFE0EC095A7068951866FB2AFBE7" box="[957,1152,1087,1114]" name="United Kingdom" pageId="12" pageNumber="13">United Kingdom</collectingCountry>
). The specimen remained at the NHMB and currently bears only one determination label written by Frey, so we suppose he removed the label of The Natural History Museum from the specimen. In the key,
<bibRefCitation id="217F9281EC095A70684A18DEFC4DFB1F" author="Frey, G." box="[866,999,1159,1186]" pageId="12" pageNumber="13" pagination="38 - 64" refId="ref36163" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten Aus Dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
illustrated the parameres of
<taxonomicName id="82EE94F3EC095A706E0F18DEFA36FB1C" authorityName="Frey" authorityYear="1969" box="[1319,1436,1159,1185]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="779A3362EC095A706E0F18DEFA36FB1C" box="[1319,1436,1159,1185]" italics="true" pageId="12" pageNumber="13">L. densata</emphasis>
</taxonomicName>
under the name of
<taxonomicName id="82EE94F3EC095A706A4118F2FDA8FB78" authorityName="Fairmaire" authorityYear="1892" box="[361,514,1195,1221]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="opacicollis">
<emphasis id="779A3362EC095A706A4118F2FDA8FB78" box="[361,514,1195,1221]" italics="true" pageId="12" pageNumber="13">L. opacicollis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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