237 lines
32 KiB
XML
237 lines
32 KiB
XML
<document id="7AD20A53A0161682308BEF1FF757C019" ID-DOI="10.11646/zootaxa.3957.3.1" ID-GBIF-Dataset="31f8999c-e65b-4e76-93be-09f17c90bffe" ID-ISSN="1175-5326" ID-Zenodo-Dep="233401" ID-ZooBank="6CC62878-DBBD-4478-AAF2-9403B46C8C36" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461061343700" checkinUser="plazi" docAuthor="Zhao, Zong-Yi & Zhou, Hong-Zhang" docDate="2015" docId="A81387A576142044F9AAFC7612C74CD6" docLanguage="en" docName="zt03957p276.pdf" docOrigin="Zootaxa 3957 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius" docType="treatment" docVersion="10" lastPageNumber="275" masterDocId="542AFFDD7602205CF93DFF8E16684D51" masterDocTitle="Phylogeny and taxonomic revision of the subgenus Velleius Leach (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="276" masterPageNumber="251" pageNumber="273" updateTime="1698582440283" updateUser="plazi">
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<mods:titleInfo id="EEA16EF108279F87F94C56D86DCC3609">
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<mods:title id="BF992DFEB1B9EC78DAC8AB9FFD1A29F8">Phylogeny and taxonomic revision of the subgenus Velleius Leach (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="3C190FE1133BAC73975A3544074F6158">Zhao, Zong-Yi</mods:namePart>
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<mods:namePart id="C0FA8998701A536AA66F9DE0315CC3CE">Zhou, Hong-Zhang</mods:namePart>
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<mods:typeOfResource id="75433BA291E6DC464DB1A56568F336AE">text</mods:typeOfResource>
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<mods:title id="7F7D25612B2AEEC981AC9B393C95A31D">Zootaxa</mods:title>
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<mods:part id="307BA6C47AD89931B367B2FDA634A647">
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<mods:date id="457B1E2E6D4F56C7386C3346FE58110B">2015</mods:date>
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<mods:number id="CB16C78409D945808BAAE3719D20A274">3957</mods:number>
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<treatment id="A81387A576142044F9AAFC7612C74CD6" ID-DOI="http://doi.org/10.5281/zenodo.5663295" ID-GBIF-Taxon="119614163" ID-Zenodo-Dep="5663295" LSID="urn:lsid:plazi:treatment:A81387A576142044F9AAFC7612C74CD6" httpUri="http://treatment.plazi.org/id/A81387A576142044F9AAFC7612C74CD6" lastPageId="24" lastPageNumber="275" pageId="22" pageNumber="273">
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<subSubSection id="68A065387614204AF9AAFC7614F14943" box="[151,665,1015,1042]" pageId="22" pageNumber="273" type="nomenclature">
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<paragraph id="200536B37614204AF9AAFC7614F14943" blockId="22.[151,665,1015,1075]" box="[151,665,1015,1042]" pageId="22" pageNumber="273">
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<heading id="7B4D81DF7614204AF9AAFC7614F14943" bold="true" box="[151,665,1015,1042]" fontSize="11" level="1" pageId="22" pageNumber="273" reason="1">
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<emphasis id="12CEEAA17614204AF9AAFC7614F14943" bold="true" box="[151,665,1015,1042]" pageId="22" pageNumber="273">
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9.
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<taxonomicName id="E7BA4D307614204AF989FC7917714940" ID-CoL="8VS2M" box="[180,281,1015,1041]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="genus">
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<emphasis id="12CEEAA17614204AF989FC7917714940" bold="true" box="[180,281,1015,1041]" italics="true" pageId="22" pageNumber="273">Quedius</emphasis>
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</taxonomicName>
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(
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<emphasis id="12CEEAA17614204AF817FC7917ED4940" bold="true" box="[298,389,1015,1041]" italics="true" pageId="22" pageNumber="273">Ve l le iu s</emphasis>
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)
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<taxonomicName id="E7BA4D307614204AF8A7FC7914F14943" authority="Sharp 1889" authorityName="Sharp" authorityYear="1889" box="[410,665,1015,1042]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="setosus">
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<emphasis id="12CEEAA17614204AF8A7FC7917984940" bold="true" box="[410,496,1015,1041]" italics="true" pageId="22" pageNumber="273">setosus</emphasis>
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(
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<bibRefCitation id="442B4B427614204AF8C2FC7614F94943" author="Sharp" box="[511,657,1016,1042]" pageId="22" pageNumber="273" refString="Sharp, D. S. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3, 28 - 44." type="journal article" year="1889">Sharp 1889</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="68A065387614204BF9AAFB9412634ABF" lastPageId="23" lastPageNumber="274" pageId="22" pageNumber="273" type="description">
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<paragraph id="200536B37614204AF9AAFB9417684962" blockId="22.[151,665,1015,1075]" box="[151,256,1050,1075]" pageId="22" pageNumber="273">
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(
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<figureCitation id="B8812A367614204AF9A2FB94169A4962" box="[159,242,1050,1075]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="22" pageNumber="273">Fig. 13</figureCitation>
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).
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</paragraph>
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<paragraph id="200536B37614204AF9AAFBEC171849C6" blockId="22.[151,1436,1122,1175]" pageId="22" pageNumber="273">
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<bibRefCitation id="442B4B427614204AF9AAFBEC17784929" author="Sharp" box="[151,272,1122,1144]" pageId="22" pageNumber="273" refString="Sharp, D. S. (1889) The Staphylinidae of Japan. The Annals and Magazine of Natural History, Series 6, 3, 28 - 44." type="journal article" year="1889">Sharp 1889</bibRefCitation>
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: 29 (originally in
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<taxonomicName id="E7BA4D307614204AF8F0FBEC14744929" box="[461,540,1122,1144]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="genus">
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<emphasis id="12CEEAA17614204AF8F0FBEC14744929" box="[461,540,1122,1144]" italics="true" pageId="22" pageNumber="273">Velleius</emphasis>
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</taxonomicName>
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;
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<typeStatus id="FF0188117614204AFB16FBED14354929" box="[555,605,1123,1144]" pageId="22" pageNumber="273">Type</typeStatus>
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locality:
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<collectingCountry id="58AD76237614204AFBFCFBED14974929" box="[705,767,1123,1144]" name="Japan" pageId="22" pageNumber="273">Japan</collectingCountry>
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: Junsai);
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<bibRefCitation id="442B4B427614204AFA5CFBEC158C4929" author="Naomi" box="[865,996,1122,1144]" pageId="22" pageNumber="273" refString="Naomi, S. I. (1986) Taxonomic study on the genus Velleius Mannerheim of Japan, with description of a new species (Coleoptera: Staphylinidae). Transactions of the Shikoku Entomological Society, 17 (4), 239 - 246." type="journal article" year="1986">Naomi 1986</bibRefCitation>
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: 243;
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<bibRefCitation id="442B4B427614204AFD18FBEC12AF4929" author="Watanabe" box="[1061,1223,1122,1144]" pageId="22" pageNumber="273" refString="Watanabe, Y. (1990) A revision of the Japanese species of the genus Velleius (Coleoptera, Staphylinidae). Elytra, 18 (1), 59 - 72." type="journal article" year="1990">Watanabe 1990</bibRefCitation>
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: 65;
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<bibRefCitation id="442B4B427614204AFDC6FBEC130C4929" author="Cho" box="[1275,1380,1122,1144]" pageId="22" pageNumber="273" refString="Cho, Y. B. (1996) Taxonomic study on the tribe Quediini (Coleoptera, Staphylinidae) from Korea. The Korean Journal of Systematic Zoology, 12 (2), 107 - 119." type="journal article" year="1996">Cho 1996</bibRefCitation>
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: 116;
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<bibRefCitation id="442B4B427614204AF9FAFB0F175149C6" author="Zhou" box="[199,313,1153,1175]" pageId="22" pageNumber="273" refString="Zhou, H. Z. (1999) Staphylinidae, Pselaphidae, Histeridae. In: Chen, S. C. (Ed.), Pictorial Handbook of Rare and Precious Insects in China. Forestry Publishing House, Beijing, pp. 119 - 123." type="book chapter" year="1999">Zhou 1999</bibRefCitation>
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: 121.
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</paragraph>
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<paragraph id="200536B37614204AF9AAFB4A17C44851" blockId="22.[151,1436,1220,1857]" pageId="22" pageNumber="273">
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<emphasis id="12CEEAA17614204AF9AAFB4A172C498C" bold="true" box="[151,324,1220,1245]" pageId="22" pageNumber="273">Redescription.</emphasis>
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Body length about 19.0 mm, body width about
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<quantity id="E7429B567614204AFA60FB4A15DB498D" box="[861,947,1220,1245]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="22" pageNumber="273" unit="mm" value="4.6">4.6 mm</quantity>
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(EW), HW/PW/EW/AW = 1.0:1.4:1.6:1.3, HL/PL/EL = 1.0:1.4:1.7.
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</paragraph>
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<paragraph id="200536B37614204AF9FAFA8212154874" blockId="22.[151,1436,1220,1857]" box="[199,1149,1292,1317]" pageId="22" pageNumber="273">Body entirely very dark brown, pronotum iridescent, hypomeron of elytra yellowish.</paragraph>
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<paragraph id="200536B37614204AF9FAFAA1134A48AC" blockId="22.[151,1436,1220,1857]" pageId="22" pageNumber="273">Head nearly quadrate, wider than long, HW/HL = 1.0:0.8, eyes large, in dorsal view longer than tempora, HEL/HTL = 1.0:0.7; posterior frontal setiferous puncture behind level of posterior edge of eye and separated from posteromedian edge of eye by about two diameters of puncture, one basal setiferous puncture on each side and much smaller than posterior frontal setiferous puncture, temporal setiferous puncture closer to nuchal constriction than to posterior edge of eye; one oblique narrow depression just behind insertion of antenna on each side, dorsal surface covered with sparse fine punctures and dense microsculpture consisting of isodiametric meshes.</paragraph>
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<paragraph id="200536B37614204AF9FAF98617144B39" blockId="22.[151,1436,1220,1857]" pageId="22" pageNumber="273">Antennae with apex of segment III about equal wide to segment I, segment IV shallowly, segments V–X each deeply bifurcated, segment III longer than segment II, last segment longer than 2 but shorter than 3 preceding segments combined.</paragraph>
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<paragraph id="200536B37614204AF9FAF9FA12A34BE1" blockId="22.[151,1436,1220,1857]" pageId="22" pageNumber="273">
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Left mandible (
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<figureCitation id="B8812A367614204AF84AF9FA17A04BDD" box="[375,456,1652,1677]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="22" pageNumber="273">Fig. 13</figureCitation>
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E) with anterior tooth divided into two subteeth with posterior one larger, edge before anterior tooth smoothly curved, posterior tooth absent, right mandible (
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<figureCitation id="B8812A367614204AFA8EF919126A4BE1" box="[947,1026,1687,1712]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="22" pageNumber="273">Fig. 13</figureCitation>
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F) with two teeth.
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</paragraph>
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<paragraph id="200536B37614204AF9FAF93213F44BA9" blockId="22.[151,1436,1220,1857]" pageId="22" pageNumber="273">Maxillary palpus with last segment nearly parallel-sided and lacking seta, surface covered with many longitudinal foveae on basal half and with many pores on apical half, ratio of length of segments II–IV 1.0:0.8:1.1.</paragraph>
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<paragraph id="200536B37614204AF9FAF88A14EC4A11" blockId="22.[151,1436,1220,1857]" pageId="22" pageNumber="273">Labial palpus with last segment fusiform and lacking seta, surface covered with many pores on apical portion, ratio of length of segments I–III 1.0:1.0:1.2.</paragraph>
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<caption id="74C5663B7615204BF9AAFC1F13704EB5" httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274" targetBox="[227,1359,187,883]" targetPageId="23">
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<paragraph id="200536B37615204BF9AAFC1F13704EB5" blockId="23.[151,1436,913,996]" pageId="23" pageNumber="274">
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<emphasis id="12CEEAA17615204BF9AAFC1F174B4EF6" bold="true" box="[151,291,913,935]" pageId="23" pageNumber="274">FIGURE 13.</emphasis>
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<taxonomicName id="E7BA4D307615204BF811FC1C17AC4EF6" box="[300,452,913,935]" class="Insecta" family="Staphylinidae" genus="Velleius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="setosus" subGenus="Velleius">
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<emphasis id="12CEEAA17615204BF811FC1C172D4EF6" box="[300,325,914,935]" italics="true" pageId="23" pageNumber="274">Q.</emphasis>
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(
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<emphasis id="12CEEAA17615204BF86BFC1C17004EF6" box="[342,360,914,935]" italics="true" pageId="23" pageNumber="274">V.</emphasis>
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)
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<emphasis id="12CEEAA17615204BF846FC1C17AC4EF6" box="[379,452,914,935]" italics="true" pageId="23" pageNumber="274">setosus</emphasis>
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</taxonomicName>
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. A–D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E–F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm.
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</paragraph>
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</caption>
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<paragraph id="200536B37615204BF9FAFB84144B4917" blockId="23.[151,1436,1034,2031]" pageId="23" pageNumber="274">Neck covered with dense microsculpture consisting meshes more transverse than those on head, and also covered with sparse fine punctures.</paragraph>
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<paragraph id="200536B37615204BF9FAFBDD12124987" blockId="23.[151,1436,1034,2031]" pageId="23" pageNumber="274">Pronotum wider than long, PW/PL = 1.0:0.8; two large setiferous punctures in each dorsal row, sometimes additional small puncture accompanying around, one large setiferous puncture in each sublateral row, large lateral setiferous puncture behind level of sublateral row but before posterior puncture in dorsal row; surface covered with dense but very vague transverse microsculpture, also covered with sparse fine punctures.</paragraph>
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<paragraph id="200536B37615204BF9FAFB6C15F5484E" blockId="23.[151,1436,1034,2031]" pageId="23" pageNumber="274">Scutellum setose, surface covered with very vague transverse microsculpture, anterior basal ridge with middle portion nearly straight, posterior basal ridge slightly arced backward.</paragraph>
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<paragraph id="200536B37615204BF9FAFAA417A44836" blockId="23.[151,1436,1034,2031]" pageId="23" pageNumber="274">Elytra. EW/EL = 1.0:0.9, EL/ESL = 1.0:0.5, surface evenly and densely setose, surface between setae covered with dense micropunctures.</paragraph>
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<paragraph id="200536B37615204BF9FAFAFC14FF48FE" blockId="23.[151,1436,1034,2031]" pageId="23" pageNumber="274">Abdomen with each tergite evenly setose, tergites III–VII all bearing one large middle lateral setae on each side, tergite VII bearing whitish apical fringe.</paragraph>
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<paragraph id="200536B37615204BF9FAFA3414B84882" blockId="23.[151,1436,1034,2031]" box="[199,720,1466,1491]" pageId="23" pageNumber="274">Apices of meso- and metatarsomere V dilated.</paragraph>
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<paragraph id="200536B37615204BF9FAFA5016884AD2" blockId="23.[151,1436,1034,2031]" pageId="23" pageNumber="274">
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Male. Sternite VIII with a middle emargination on apical margin, a small area before it glabrous; sternite IX (
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<figureCitation id="B8812A367615204BF9A2F98C16864B4A" box="[159,238,1538,1563]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274">Fig. 13</figureCitation>
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G) with basal portion very slender and short, apical portion without emargination on apical margin, surface from widest portion to apex covered with dense setiferous punctures, several larger setae on apical margin; tergite X (
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<figureCitation id="B8812A367615204BF982F9C4177B4B32" box="[191,275,1610,1635]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274">Fig. 13</figureCitation>
|
||
H) with basal edge arcuated backwards, apical portion nearly triangle-shaped, apex obtuse but not truncated, surface of apical 1/3 bearing many large setae; aedeagus in lateral view (
|
||
<figureCitation id="B8812A367615204BFDB9F9E012B24BD6" box="[1156,1242,1646,1671]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274">Fig. 13</figureCitation>
|
||
A) with apex of paramere protruding slightly beyond apex of median lobe, paramere nearly equally wide through whole length, not suddenly widened below apex, median lobe with a hook facing paramere below apex and concave below the hook (
|
||
<figureCitation id="B8812A367615204BF9A2F95416994BA2" box="[159,241,1754,1779]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274">Fig. 13</figureCitation>
|
||
D), in dorsal view (
|
||
<figureCitation id="B8812A367615204BF8EEF954144E4BA2" box="[467,550,1754,1779]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274">Fig. 13</figureCitation>
|
||
B) paramere wider than median lobe through whole length, paramere slightly dilated at about apical 1/3, apical portion of paramere (
|
||
<figureCitation id="B8812A367615204BFA29F970150C4A46" box="[788,868,1790,1815]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274">Fig. 13</figureCitation>
|
||
C) divided into two branches separated from each other, 2 apical and 2 smaller lateral setae on each side, sensory peg setae on underside of paramere forming two groups extending from apex towards base and beyond insertions of lateral setae, 14 or 15 sensory peg setae in each group.
|
||
</paragraph>
|
||
<paragraph id="200536B37615204BF9FAF80012634ABF" blockId="23.[151,1436,1034,2031]" pageId="23" pageNumber="274">
|
||
Female. Sternite IX (
|
||
<figureCitation id="B8812A367615204BF88AF80014624AF6" box="[439,522,1934,1959]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274">
|
||
Fig.
|
||
<date id="540410737615204BF8D3F80114624AF6" box="[494,522,1935,1959]" pageId="23" pageNumber="274" value="2013-01">13</date>
|
||
</figureCitation>
|
||
I) bearing several large apical setae; tergite X (
|
||
<figureCitation id="B8812A367615204BFD19F800121E4AF6" box="[1060,1142,1934,1959]" captionStart="FIGURE 13. Q" captionStartId="23.[151,250,913,935]" captionTargetBox="[227,1359,187,883]" captionTargetId="figure@23.[221,1366,183,893]" captionTargetPageId="23" captionText="FIGURE 13. Q. (V.) setosus. A – D, aedeagus, (A) lateral view, (B) dorsal view, (C) underside of paramere apex, showing sensory peg setae, (D) apical portion of median lobe in lateral view; E – F, mandibles, dorsal view; G, male abdominal sternite IX; H, male abdominal tergite X; I, female sternite IX, right part; J, female abdominal tergite X. Scales = 0.5 mm." httpUri="https://zenodo.org/record/233414/files/figure.png" pageId="23" pageNumber="274">Fig. 13</figureCitation>
|
||
J) with basal side broadly and smoothly concaved, apical portion trilobed, the two lateral lobes smaller than the middle lobe, several large setae on and before apical margin of middle lobe, apex obtuse but not truncate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68A06538761A2044F9FAFF1915B84C35" pageId="24" pageNumber="275" type="materials_examined">
|
||
<paragraph id="200536B3761A2044F9FAFF1912094DFE" blockId="24.[151,1436,151,392]" box="[199,1121,151,176]" pageId="24" pageNumber="275">
|
||
<emphasis id="12CEEAA1761A2044F9FAFF1917814DE1" bold="true" box="[199,489,151,176]" pageId="24" pageNumber="275">
|
||
<typeStatus id="FF018811761A2044F9FAFF19176A4DE1" box="[199,258,151,176]" pageId="24" pageNumber="275">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="FF018811761A2044F841FF1917814DE1" box="[380,489,151,176]" pageId="24" pageNumber="275" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
, male,
|
||
<collectingCountry id="58AD7623761A2044FB06FF1614164DFE" box="[571,638,152,175]" name="Japan" pageId="24" pageNumber="275">Japan</collectingCountry>
|
||
, collected by G. Lewis, deposited in NHM.
|
||
</paragraph>
|
||
<paragraph id="200536B3761A2044F9FAFF3214D04DA9" blockId="24.[151,1436,151,392]" pageId="24" pageNumber="275">
|
||
<emphasis id="12CEEAA1761A2044F9FAFF3217AE4D84" bold="true" box="[199,454,188,213]" pageId="24" pageNumber="275">Specimens examined.</emphasis>
|
||
<specimenCount id="36BCFD3A761A2044F8F0FF3214734D84" box="[461,539,188,213]" pageId="24" pageNumber="275" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="36BCFD3A761A2044FB1BFF3214E24D84" box="[550,650,188,213]" pageId="24" pageNumber="275" type="female">1 female</specimenCount>
|
||
,
|
||
<collectingCountry id="58AD7623761A2044FBABFF3214B44D84" box="[662,732,188,213]" name="China" pageId="24" pageNumber="275">China</collectingCountry>
|
||
, Hubei Province, Xingshan, Longmenhe,
|
||
<quantity id="E7429B56761A2044FD8CFF3213694D84" box="[1201,1281,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="24" pageNumber="275" unit="m" value="1350.0">1350m</quantity>
|
||
,
|
||
<date id="54041073761A2044FC31FF3213F04D85" box="[1292,1432,188,212]" pageId="24" pageNumber="275" value="1993-07-20">20.VII.1993</date>
|
||
, collected by Yang Xingke, deposited in IZ-CAS.
|
||
</paragraph>
|
||
<paragraph id="200536B3761A2044F9FAFE8A15B84C35" blockId="24.[151,1436,151,392]" pageId="24" pageNumber="275">
|
||
<emphasis id="12CEEAA1761A2044F9FAFE8A17094C4C" bold="true" box="[199,353,260,285]" pageId="24" pageNumber="275">Comparison.</emphasis>
|
||
<taxonomicName id="E7BA4D30761A2044F853FE8A14F74C4D" box="[366,671,260,284]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="setosus" subGenus="Velleius">
|
||
<emphasis id="12CEEAA1761A2044F853FE8A17A34C4D" box="[366,459,260,284]" italics="true" pageId="24" pageNumber="275">Quedius</emphasis>
|
||
(
|
||
<emphasis id="12CEEAA1761A2044F8DCFE8A145D4C4D" box="[481,565,260,284]" italics="true" pageId="24" pageNumber="275">Velleius</emphasis>
|
||
)
|
||
<emphasis id="12CEEAA1761A2044FB73FE8B14F74C4D" box="[590,671,261,284]" italics="true" pageId="24" pageNumber="275">setosus</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="E7BA4D30761A2044FA7AFE8A127E4C4D" box="[839,1046,260,284]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="pectinatus" subGenus="Velleius">
|
||
<emphasis id="12CEEAA1761A2044FA7AFE8A150B4C4A" box="[839,867,260,283]" italics="true" pageId="24" pageNumber="275">Q.</emphasis>
|
||
(
|
||
<emphasis id="12CEEAA1761A2044FA45FE8B15E34C4D" box="[888,907,261,284]" italics="true" pageId="24" pageNumber="275">V.</emphasis>
|
||
)
|
||
<emphasis id="12CEEAA1761A2044FA9EFE8B127E4C4D" box="[931,1046,261,284]" italics="true" pageId="24" pageNumber="275">pectinatus</emphasis>
|
||
</taxonomicName>
|
||
, but its last antennal segment is obviously shorter than preceding three segments combined. It is also similar to
|
||
<taxonomicName id="E7BA4D30761A2044FD72FEA613644C11" box="[1103,1292,296,320]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="dilatatus" subGenus="Velleius">
|
||
<emphasis id="12CEEAA1761A2044FD72FEA612024C6E" box="[1103,1130,296,319]" italics="true" pageId="24" pageNumber="275">Q.</emphasis>
|
||
(
|
||
<emphasis id="12CEEAA1761A2044FDBDFEA712FA4C11" box="[1152,1170,297,320]" italics="true" pageId="24" pageNumber="275">V.</emphasis>
|
||
)
|
||
<emphasis id="12CEEAA1761A2044FD97FEA613644C11" box="[1194,1292,296,320]" italics="true" pageId="24" pageNumber="275">dilatatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E7BA4D30761A2044FC70FEA6175B4C35" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="circumipectus" subGenus="Velleius">
|
||
<emphasis id="12CEEAA1761A2044FC70FEA613004C6E" box="[1357,1384,296,319]" italics="true" pageId="24" pageNumber="275">Q.</emphasis>
|
||
(
|
||
<emphasis id="12CEEAA1761A2044FC43FEA713F84C11" box="[1406,1424,297,320]" italics="true" pageId="24" pageNumber="275">V.</emphasis>
|
||
)
|
||
<emphasis id="12CEEAA1761A2044F9AAFEC3175B4C35" box="[151,307,333,356]" italics="true" pageId="24" pageNumber="275">circumipectus</emphasis>
|
||
</taxonomicName>
|
||
, but with much deeper bifurcated antennal segments IV–X.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="68A06538761A2044F9FAFEE112C74CD6" box="[199,1199,367,392]" pageId="24" pageNumber="275" type="distribution">
|
||
<paragraph id="200536B3761A2044F9FAFEE112C74CD6" blockId="24.[151,1436,151,392]" box="[199,1199,367,392]" pageId="24" pageNumber="275">
|
||
<emphasis id="12CEEAA1761A2044F9FAFEE117084CD9" bold="true" box="[199,352,367,392]" pageId="24" pageNumber="275">Distribution.</emphasis>
|
||
<taxonomicName id="E7BA4D30761A2044F85AFEFE17AC4CD9" box="[359,452,368,392]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="12CEEAA1761A2044F85AFEFE17AC4CD9" box="[359,452,368,392]" italics="true" pageId="24" pageNumber="275">Quedius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="12CEEAA1761A2044F8E9FEFE14404CD6" box="[468,552,368,392]" italics="true" pageId="24" pageNumber="275">Vell eius</emphasis>
|
||
)
|
||
<taxonomicName id="E7BA4D30761A2044FB01FEFF14E54CD9" box="[572,653,369,392]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="setosus">
|
||
<emphasis id="12CEEAA1761A2044FB01FEFF14E54CD9" box="[572,653,369,392]" italics="true" pageId="24" pageNumber="275">setosus</emphasis>
|
||
</taxonomicName>
|
||
is known from
|
||
<collectingCountry id="58AD7623761A2044FA03FEE115E94CD9" box="[830,897,367,392]" name="China" pageId="24" pageNumber="275">China</collectingCountry>
|
||
(Hubei),
|
||
<collectingCountry id="58AD7623761A2044FAD0FEFE12594CD6" box="[1005,1073,368,391]" name="South Korea" pageId="24" pageNumber="275">Korea</collectingCountry>
|
||
and
|
||
<collectingCountry id="58AD7623761A2044FD55FEFE12C34CD6" box="[1128,1195,368,391]" name="Japan" pageId="24" pageNumber="275">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |